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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.94.95751" ID-Pensoft-Pub="1314-4049-94-51" ID-Pensoft-UUID="1E443200E6585C66B1D669737794361F" ModsDocID="1314-4049-94-51" checkinTime="1670952471273" checkinUser="pensoft" docAuthor="Lu, Li, Karunarathna, Samantha C., Dai, Dong-qin, Jayawardena, Ruvishika S., Suwannarach, Nakarin &amp; Tibpromma, Saowaluck" docDate="2022" docId="6C59D4C7E1AF5732BDD602F00DCC1623" docLanguage="en" docName="MycoKeys 94: 51-71" docOrigin="MycoKeys 94" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/mycokeys.94.95751" docTitle="Nigrograna asexualis L. Lu &amp; Tibpromma 2022, sp. nov." docType="treatment" docVersion="1" id="1E443200E6585C66B1D669737794361F" lastPageNumber="51" masterDocId="1E443200E6585C66B1D669737794361F" masterDocTitle="Three new species of Nigrograna (Dothideomycetes, Pleosporales) associated with Arabica coffee from Yunnan Province, China" masterLastPageNumber="71" masterPageNumber="51" pageNumber="51" updateTime="1670952471273" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of Nigrograna (Dothideomycetes, Pleosporales) associated with Arabica coffee from Yunnan Province, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lu, Li</mods:namePart>
<mods:affiliation>Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karunarathna, Samantha C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7080-0781</mods:nameIdentifier>
<mods:affiliation>Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dai, Dong-qin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8935-8807</mods:nameIdentifier>
<mods:affiliation>Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jayawardena, Ruvishika S.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suwannarach, Nakarin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2653-1913</mods:nameIdentifier>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">suwan.462@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tibpromma, Saowaluck</mods:namePart>
<mods:affiliation>Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
<mods:nameIdentifier type="email">saowaluckfai@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title>MycoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>94</mods:number>
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<mods:start>51</mods:start>
<mods:end>71</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.94.95751</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.94.95751</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-94-51</mods:identifier>
<mods:identifier type="Pensoft-UUID">1E443200E6585C66B1D669737794361F</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="51" type="results">
<treatment LSID="urn:lsid:plazi:treatment:6C59D4C7E1AF5732BDD602F00DCC1623" httpUri="http://treatment.plazi.org/id/6C59D4C7E1AF5732BDD602F00DCC1623" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="6C59D4C7-E1AF-5732-BDD6-02F00DCC1623" authority="L. Lu &amp; Tibpromma" authorityName="L. Lu &amp; Tibpromma" authorityYear="2022" class="Dothideomycetes" family="Biatriosporaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna asexualis" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="species" species="asexualis" status="sp. nov.">Nigrograna asexualis L. Lu &amp; Tibpromma</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nigrograna asexualis (ZHKU 22 - 0123, holotype) a, b conidiomata on the host substrate c, d vertical sections of a conidioma e peridium f, g conidiophores with phialides h conidia i culture on PDA from above and reverse. Scale bars: 100 μm (c); 50 μm (d); 15 μm (e); 30 μm (f); 20 μm (g); 10 μm (h)." figureDoi="10.3897/mycokeys.94.95751.figure4" httpUri="https://binary.pensoft.net/fig/781091" pageId="0" pageNumber="51">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="etymology">
<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="51">
The species epithet '
<taxonomicName authorityName="L. Lu &amp; Tibpromma" authorityYear="2022" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna asexualis" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna asexualis</emphasis>
</taxonomicName>
' refers to the asexual morph.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="holotype">
<paragraph pageId="0" pageNumber="51">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="51">ZHKU 22-0123.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Saprobic</emphasis>
on decaying branch of
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" lsidName="Coffea arabica." order="Gentianales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="species" species="arabica.">
<emphasis italics="true" pageId="0" pageNumber="51">Coffea arabica.</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">Sexual morph</emphasis>
: Undetermined.
<emphasis bold="true" pageId="0" pageNumber="51">Asexual morph</emphasis>
: Coelomycetous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Pycnidia</emphasis>
100-230
<normalizedToken originalValue="µm">µm</normalizedToken>
high, 120-180
<normalizedToken originalValue="µm">µm</normalizedToken>
wide (x̄ = 156
<normalizedToken originalValue="×">x</normalizedToken>
144
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 10), globose to subglobose, or pyriform, immersed, solitary, unilocular, dark brown, papillate ostiole, appearing as black spots on host surface.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Pycnidial wall</emphasis>
11-16
<normalizedToken originalValue="µm">µm</normalizedToken>
wide (x̄ = 14
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 15), brown (#937463), the wall with pseudoparenchymatous cells.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Conidiophores</emphasis>
arising from the pycnidial wall, up to 46
<normalizedToken originalValue="µm">µm</normalizedToken>
long and 3-4.4
<normalizedToken originalValue="µm">µm</normalizedToken>
wide (x̄ = 3.4
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 25), filiform, septate, hyaline, simple to sparsely branched, with pegs along one or two sides and solitary phialides terminally.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Phialides</emphasis>
3-6
<normalizedToken originalValue="×">x</normalizedToken>
1-2
<normalizedToken originalValue="µm">µm</normalizedToken>
(x̄ = 4.5
<normalizedToken originalValue="×">x</normalizedToken>
1.5
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 15), variable in shape, phialidic, discrete, ampulliform-lageniform-subcylindrical.
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Conidia</emphasis>
5-6.5
<normalizedToken originalValue="×">x</normalizedToken>
3-4
<normalizedToken originalValue="µm">µm</normalizedToken>
(x̄ = 5.5
<normalizedToken originalValue="×">x</normalizedToken>
3.7
<normalizedToken originalValue="µm">µm</normalizedToken>
, n = 30), ellipsoidal, unicellular, aseptate with 1-2 granules, subhyaline, smooth-walled.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Culture characteristics.</paragraph>
<paragraph pageId="0" pageNumber="51">Conidium germinated on PDA within 24 h. Colonies growing on PDA reaching 5 cm diam. after two months at room temperature (22-26 °C). Colony dense, circular, surface sparsely hairy, radially striate, with a fimbriate edge, yellowish (#eabf83) to pale brown (#e1af33) at the center and dark brown (#6e5031) at the margin, reverse dark brown (#6e5031).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="materials_examined">
<paragraph pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="51">
<materialsCitation accessionNumber="OP450965, OP450971, OP432241, OP450979, OP432245, OP450966, OP450972, OP432242, OP450980, OP432246" collectingDate="2021-09-16" country="China" elevation="1016" latitude="22.600555" location="Pu'er City" longLatPrecision="21" longitude="101.01639" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<location LSID="urn:lsid:plazi:treatment:6C59D4C7E1AF5732BDD602F00DCC1623:715A2A6D87F06CDBCCCBA3C58A5FA55E" country="China" latitude="22.600555" longLatPrecision="21" longitude="101.01639" name="Pu'er City" stateProvince="Yunnan Province">
<normalizedToken originalValue="Puer">Pu'er</normalizedToken>
City
</location>
,
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCountry name="China">China</collectingCountry>
, on a decaying branch of
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" lsidName="Coffea arabica" order="Gentianales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis italics="true" pageId="0" pageNumber="51">Coffea arabica</emphasis>
</taxonomicName>
, (
<geoCoordinate degrees="22" direction="north" minutes="36" orientation="latitude" precision="15" seconds="2" value="22.600555">22°36'2&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="0" orientation="longitude" precision="15" seconds="59" value="101.01639">101°0'59&quot;E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.01643" unit="m" value="1016.43">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.01643" unit="m" value="1016.43">1016.43 m</elevation>
</quantity>
),
<collectingDate value="2021-09-16">16 September 2021</collectingDate>
, LiLu, Puer 1-14 (ZHKU 22-0123,
<typeStatus>holotype</typeStatus>
), ZHKUCC 22-0214 = ZHKUCC 22-0215. GenBank number; ITS:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450965">OP450965</accessionNumber>
, LSU:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450971">OP450971</accessionNumber>
,
<emphasis italics="true" pageId="0" pageNumber="51">rpb</emphasis>
2:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP432241">OP432241</accessionNumber>
, SSU:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450979">OP450979</accessionNumber>
,
<emphasis italics="true" pageId="0" pageNumber="51">tef</emphasis>
1-α:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP432245">OP432245</accessionNumber>
(ZHKUCC 22-0214, ex-type); ITS:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450966">OP450966</accessionNumber>
, LSU:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450972">OP450972</accessionNumber>
,
<emphasis italics="true" pageId="0" pageNumber="51">rpb</emphasis>
2:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP432242">OP432242</accessionNumber>
, SSU:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP450980">OP450980</accessionNumber>
,
<emphasis italics="true" pageId="0" pageNumber="51">tef</emphasis>
1-α:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OP432246">OP432246</accessionNumber>
(ZHKUCC 22-0215)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="notes">
<paragraph pageId="0" pageNumber="51">Notes.</paragraph>
<paragraph pageId="0" pageNumber="51">
In multi-gene phylogeny,
<taxonomicName authorityName="L. Lu &amp; Tibpromma" authorityYear="2022" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna asexualis" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna asexualis</emphasis>
</taxonomicName>
formed a separate (68% ML, 0.97 BIPP) and distinct clade within
<taxonomicName authorityName="J.de Gruyter, G.J.M.Verkley &amp; P.W.Crous" authorityYear="2012" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The maximum-likelihood phylogram of Nigrograna based on a combined SSU, LSU, ITS, rpb 2 and tef 1 - α sequence dataset with Cyclothyriella rubronotata CBS 141486 and CBS 419.85 as the outgroup taxa (Dayarathne et al. 2020). The maximum-likelihood bootstrap values (ML ≥ 60 %, left) and Bayesian Inference Posterior Probability values (BIPP ≥ 0.90, right) are shown above the nodes. Strains derived from the current study are in blue, while type strains are in bold." figureDoi="10.3897/mycokeys.94.95751.figure1" httpUri="https://binary.pensoft.net/fig/781088" pageId="0" pageNumber="51">1</figureCitation>
). Morphologically,
<taxonomicName lsidName="N. asexualis" pageId="0" pageNumber="51" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">N. asexualis</emphasis>
</taxonomicName>
conforms to the morphological characteristics of
<taxonomicName authorityName="J.de Gruyter, G.J.M.Verkley &amp; P.W.Crous" authorityYear="2012" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna</emphasis>
</taxonomicName>
by having hyaline or subhyaline, long and branched conidiophores, solitary phialides, and aseptate, ellipsoidal or cylindrical conidia (
<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2016.09.002" author="Jaklitsch, WM" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="51" pagination="35 - 64" refId="B19" refString="Jaklitsch, WM, Voglmayr, H, 2016. Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae. Studies in Mycology 85 (1): 35 - 64, DOI: https://doi.org/10.1016/j.simyco.2016.09.002" title="Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae." url="https://doi.org/10.1016/j.simyco.2016.09.002" volume="85" year="2016">Jaklitsch and Voglmayr 2016</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5943/mycosphere/11/1/1" author="Dayarathne, MC" journalOrPublisher="Mycosphere" pageId="0" pageNumber="51" pagination="1 - 188" refId="B7" refString="Dayarathne, MC, Jones, E, Maharachchikumbura, S, Devadatha, B, Sarma, V, Khongphinitbunjong, K, Chomnunti, P, Hyde, KD, 2020. Morpho-molecular characterization of microfungi associated with marine based habitats. Mycosphere 11: 1 - 188, DOI: https://doi.org/10.5943/mycosphere/11/1/1" title="Morpho-molecular characterization of microfungi associated with marine based habitats." url="https://doi.org/10.5943/mycosphere/11/1/1" volume="11" year="2020">Dayarathne et al. 2020</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0235855" author="Wanasinghe, DN" journalOrPublisher="Journal of Bacteriology" pageId="0" pageNumber="51" refId="B49" refString="Wanasinghe, DN, Wijayawardene, NN, Xu, J, Cheewangkoon, R, Mortimer, PE, 2020. Taxonomic novelties in Magnolia-associated pleosporalean fungi in the Kunming Botanical Gardens (Yunnan, China). PLoS ONE 15(7): e0235855. https://doi.org/10.1371/journal.pone.0235855" title="Taxonomic novelties in Magnolia-associated pleosporalean fungi in the Kunming Botanical Gardens (Yunnan, China). PLoS ONE 15 (7): e 0235855." url="https://doi.org/10.1371/journal.pone.0235855" year="2020">Wanasinghe et al. 2020</bibRefCitation>
). Blast results of the sequences show that ITS is similar to
<taxonomicName lsidName="N. fuscidula" pageId="0" pageNumber="51" rank="species" species="fuscidula">
<emphasis italics="true" pageId="0" pageNumber="51">N. fuscidula</emphasis>
</taxonomicName>
with 89% (MH856004), and SSU is similar to
<taxonomicName lsidName="N. mycophila" pageId="0" pageNumber="51" rank="species" species="mycophila">
<emphasis italics="true" pageId="0" pageNumber="51">N. mycophila</emphasis>
</taxonomicName>
with 99.8% (KX650510).
<taxonomicName authorityName="L. Lu &amp; Tibpromma" authorityYear="2022" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna asexualis" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna asexualis</emphasis>
</taxonomicName>
is different from
<taxonomicName lsidName="N. fuscidula" pageId="0" pageNumber="51" rank="species" species="fuscidula">
<emphasis italics="true" pageId="0" pageNumber="51">N. fuscidula</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="N. mycophila" pageId="0" pageNumber="51" rank="species" species="mycophila">
<emphasis italics="true" pageId="0" pageNumber="51">N. mycophila</emphasis>
</taxonomicName>
by its ellipsoidal conidia, but the similarities of these three species are hyaline, 1-celled, smooth-walled conidia forming on philipides (
<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2016.09.002" author="Jaklitsch, WM" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="51" pagination="35 - 64" refId="B19" refString="Jaklitsch, WM, Voglmayr, H, 2016. Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae. Studies in Mycology 85 (1): 35 - 64, DOI: https://doi.org/10.1016/j.simyco.2016.09.002" title="Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae." url="https://doi.org/10.1016/j.simyco.2016.09.002" volume="85" year="2016">Jaklitsch and Voglmayr 2016</bibRefCitation>
). The LSU and
<emphasis italics="true" pageId="0" pageNumber="51">rpb</emphasis>
2 sequences of our strain blast results are similar to
<taxonomicName lsidName="N. obliqua" pageId="0" pageNumber="51" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="51">N. obliqua</emphasis>
</taxonomicName>
, and the similarities are 98.9% (KX650560) and 87% (KX650579) respectively, but
<taxonomicName lsidName="N. obliqua" pageId="0" pageNumber="51" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="51">N. obliqua</emphasis>
</taxonomicName>
lacks the asexual morph (
<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2016.09.002" author="Jaklitsch, WM" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="51" pagination="35 - 64" refId="B19" refString="Jaklitsch, WM, Voglmayr, H, 2016. Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae. Studies in Mycology 85 (1): 35 - 64, DOI: https://doi.org/10.1016/j.simyco.2016.09.002" title="Hidden diversity in Thyridaria and a new circumscription of the Thyridariaceae." url="https://doi.org/10.1016/j.simyco.2016.09.002" volume="85" year="2016">Jaklitsch and Voglmayr 2016</bibRefCitation>
). The
<emphasis italics="true" pageId="0" pageNumber="51">tef</emphasis>
1-α sequence of our strain is 95.8% (MF939615) similar to
<taxonomicName lsidName="N. locuta-pollinis" pageId="0" pageNumber="51" rank="species" species="locuta-pollinis">
<emphasis italics="true" pageId="0" pageNumber="51">N. locuta-pollinis</emphasis>
</taxonomicName>
, which was isolated from hive-stored pollen of
<taxonomicName class="Magnoliopsida" family="Brassicaceae" genus="Brassica" kingdom="Plantae" lsidName="Brassica campestris" order="Brassicales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="species" species="campestris">
<emphasis italics="true" pageId="0" pageNumber="51">Brassica campestris</emphasis>
</taxonomicName>
L. that lacks morphology (
<bibRefCitation DOI="https://doi.org/10.5943/mycosphere/9/6/3" author="Zhao, YZ" journalOrPublisher="Mycosphere" pageId="0" pageNumber="51" pagination="1089 - 1116" refId="B57" refString="Zhao, YZ, Zhang, ZF, Cai, L, Peng, JW, Liu, F, 2018. Four new filamentous fungal species from newly-collected and hive stored bee pollen. Mycosphere 9: 1089 - 1116, DOI: https://doi.org/10.5943/mycosphere/9/6/3" title="Four new filamentous fungal species from newly-collected and hive stored bee pollen." url="https://doi.org/10.5943/mycosphere/9/6/3" volume="9" year="2018">Zhao et al. 2018</bibRefCitation>
). Therefore, we introduce
<taxonomicName lsidName="N. asexualis" pageId="0" pageNumber="51" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">N. asexualis</emphasis>
</taxonomicName>
as a distinct new species from coffee in China.
</paragraph>
<caption doi="10.3897/mycokeys.94.95751.figure4" httpUri="https://binary.pensoft.net/fig/781091" pageId="0" pageNumber="51" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 4.</emphasis>
<taxonomicName authorityName="L. Lu &amp; Tibpromma" authorityYear="2022" class="Dothideomycetes" family="Nigrogranaceae" genus="Nigrograna" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Nigrograna asexualis" order="Pleosporales" pageId="0" pageNumber="51" phylum="Ascomycota" rank="species" species="asexualis">
<emphasis italics="true" pageId="0" pageNumber="51">Nigrograna asexualis</emphasis>
</taxonomicName>
(ZHKU 22-0123, holotype)
<emphasis bold="true" pageId="0" pageNumber="51">a, b</emphasis>
conidiomata on the host substrate
<emphasis bold="true" pageId="0" pageNumber="51">c, d</emphasis>
vertical sections of a conidioma
<emphasis bold="true" pageId="0" pageNumber="51">e</emphasis>
peridium
<emphasis bold="true" pageId="0" pageNumber="51">f, g</emphasis>
conidiophores with phialides
<emphasis bold="true" pageId="0" pageNumber="51">h</emphasis>
conidia
<emphasis bold="true" pageId="0" pageNumber="51">i</emphasis>
culture on PDA from above and reverse. Scale bars: 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">c</emphasis>
); 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">d</emphasis>
); 15
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">e</emphasis>
); 30
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">f</emphasis>
); 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">g</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="51">h</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>