270 lines
34 KiB
XML
270 lines
34 KiB
XML
<document id="30583A0B40881472607F9E15665600F4" ID-CLB-Dataset="32119" ID-DOI="10.5852/ejt.2017.293" ID-GBIF-Dataset="3099a544-54cd-48e5-8aac-8e1ca24c0e8b" ID-ISSN="2118-9773" ID-Zenodo-Dep="889857" ID-ZooBank="urn:lsid:zoobank.org:pub:608B2AAB-39A4-494C-9CE0-AA84916008BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505237457348" checkinUser="plazi" docAuthor="Anh D. Nguyen, Zoltán Korsós, Kuem-Hee Jang & Ui-Wook Hwang" docDate="2017" docId="287287F2FFD03329DF0C1955FE508750" docLanguage="en" docName="EJT-293_Nguyen.pdf" docOrigin="European Journal of Taxonomy 293" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oxidus gigas" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="D44BFF8AFFD73323DD631D42FF8F8422" masterDocTitle="A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="8" updateTime="1698468380887" updateUser="ExternalLinkService">
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<mods:title id="741176580FAA42981F9A41E9A73976B7">A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="8B6D9D68D5B571C6FC944CBD419143F7">Anh D. Nguyen</mods:namePart>
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<mods:namePart id="13268725870A810798002FC5C5863E34">Zoltán Korsós</mods:namePart>
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<mods:namePart id="CD69B1DC2BD749FFA79C0EE071D8BA7A">Kuem-Hee Jang</mods:namePart>
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<mods:namePart id="F7C9EAA4D3105040F80CCBCF56295A77">Ui-Wook Hwang</mods:namePart>
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<mods:title id="C6AE48E8A83650F0ED66F063C2DFBBB3">European Journal of Taxonomy</mods:title>
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<mods:date id="6C54F822A63AC8DC4DE03FDC95F2CD89">2017</mods:date>
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<treatment id="287287F2FFD03329DF0C1955FE508750" ID-DOI="http://doi.org/10.5281/zenodo.5672738" ID-GBIF-Taxon="134034201" ID-Zenodo-Dep="5672738" LSID="urn:lsid:plazi:treatment:287287F2FFD03329DF0C1955FE508750" httpUri="http://treatment.plazi.org/id/287287F2FFD03329DF0C1955FE508750" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
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<subSubSection id="E8C1656FFFD03324DF0C1955FC4B8011" box="[623,964,1047,1075]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="A06436E4FFD03324DF0C1955FC4B8011" blockId="7.[623,964,1047,1109]" box="[623,964,1047,1075]" pageId="7" pageNumber="8">
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<heading id="FB2C8188FFD03324DF0C1955FC4B8011" box="[623,964,1047,1075]" centered="true" fontSize="11" level="4" pageId="7" pageNumber="8" reason="4">
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<taxonomicName id="67DB4D67FFD03324DF0C1955FC4B8011" ID-CoL="4BBXR" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[623,964,1047,1075]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigas">
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<emphasis id="92AFEAF6FFD03324DF0C1955FC838010" bold="true" box="[623,780,1047,1074]" italics="true" pageId="7" pageNumber="8">Oxidus gigas</emphasis>
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(
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<bibRefCitation id="C44A4B15FFD03324DE78195AFC338011" author="Attems C." box="[795,956,1048,1075]" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie" pageId="7" pageNumber="8" part="5" refString="Attems C. 1953. Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie 5 (3), Museum national d'Histoire naturelle, Paris." title="Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939)" type="journal volume" year="1953">Attems, 1953</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD03324DF841979FCC28077" box="[743,845,1083,1109]" pageId="7" pageNumber="8" type="description">
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||
<paragraph id="A06436E4FFD03324DF841979FCC28077" blockId="7.[623,964,1047,1109]" box="[743,845,1083,1109]" pageId="7" pageNumber="8">
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<figureCitation id="38E02A61FFD03324DF841979FCC28077" box="[743,845,1083,1109]" captionStart-0="Fig. 3" captionStart-1="Fig. 4" captionStartId-0="8.[191,234,1965,1991]" captionStartId-1="9.[189,232,1965,1991]" captionTargetBox-0="[310,1280,812,1925]" captionTargetBox-1="[187,1397,1247,1920]" captionTargetId-0="figure@8.[311,1279,812,1924]" captionTargetId-1="figure@9.[187,1397,1247,1920]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 3. Oxidus gigas (Attems, 1953) from Duc Xuan Commune, Ha Giang Prov., Vietnam. A. Entire body, length ca 34 mm. B – C. Segments 8 – 9 – 10. B. ♂. C. ♀. Scale bars = 1 mm. (photo by Anh Nguyen)" captionText-1="Fig. 4. Right gonopod of Oxidus gigas (Attems, 1953), from sample IEBR-Myr 113. A. Lateral view. B. Ventral view. C. Mesal view. The picture has been Fipped horizontally. Scale bars = 1 um." httpUri-0="https://zenodo.org/record/889863/files/figure.png" httpUri-1="https://zenodo.org/record/889865/files/figure.png" pageId="7" pageNumber="8">Figs 3–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD03324DDDE193EFBB9813D" pageId="7" pageNumber="8" type="reference_group">
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<paragraph id="A06436E4FFD03324DDDE193EFCE380B5" blockId="7.[189,876,1148,1175]" box="[189,876,1148,1175]" pageId="7" pageNumber="8">
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<taxonomicName id="67DB4D67FFD03324DDDE193EFC8280B5" authority="Attems, 1953: 165" authorityName="Attems" authorityPageNumber="165" authorityYear="1953" box="[189,781,1148,1175]" class="Diplopoda" family="Paradoxosomatidae" genus="Kalorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="gracilis" subSpecies="gigas">
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<emphasis id="92AFEAF6FFD03324DDDE193EFDA580B4" box="[189,554,1148,1174]" italics="true" pageId="7" pageNumber="8">Kalorthomorpha gracilis gigas</emphasis>
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<bibRefCitation id="C44A4B15FFD03324DF52193FFD5B80B5" author="Attems C." box="[561,724,1148,1175]" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie" pageId="7" pageNumber="8" part="5" refString="Attems C. 1953. Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie 5 (3), Museum national d'Histoire naturelle, Paris." title="Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939)" type="journal volume" year="1953">Attems, 1953</bibRefCitation>
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: 165
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</taxonomicName>
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, fg. 44.
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</paragraph>
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<paragraph id="A06436E4FFD03324DDDE19FCFB1B80FA" blockId="7.[189,1172,1214,1311]" box="[189,1172,1214,1240]" pageId="7" pageNumber="8">
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<taxonomicName id="67DB4D67FFD03324DDDE19FCFE3780FA" ID-CoL="4BBXR" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[189,440,1214,1240]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="gracilis" subSpecies="gigas">
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<emphasis id="92AFEAF6FFD03324DDDE19FCFE3780FA" box="[189,440,1214,1240]" italics="true" pageId="7" pageNumber="8">Oxidus gracilis gigas</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="C44A4B15FFD03324DCB619FCFDFD80FA" author="Jeekel C. A. W." box="[469,626,1214,1240]" journalOrPublisher="Studies on the Fauna of Suriname and other Guyanas" pageId="7" pageNumber="8" pagination="1 - 157" part="4" refString="Jeekel C. A. W. 1963 a. Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas 4 (11): 1 - 157." title="Diplopoda of Guiana" type="journal article" year="1963" yearSuffix="a">Jeekel 1963a</bibRefCitation>
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: 20;
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<bibRefCitation id="C44A4B15FFD03324DFC919FCFCB580FA" author="Jeekel C. A. W." box="[682,826,1214,1240]" journalOrPublisher="PhD Thesis, University of Amsterdam, The Netherlands" pageId="7" pageNumber="8" refString="Jeekel C. A. W. 1968. On the Classifcation and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). PhD Thesis, University of Amsterdam, The Netherlands." title="On the Classifcation and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida)" type="book" year="1968">Jeekel 1968</bibRefCitation>
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: 60. —
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<bibRefCitation id="C44A4B15FFD03324DEF619FCFBD980FA" author="Golovatch S. I." box="[917,1110,1214,1240]" editor="Sokolov V. E." journalOrPublisher="Nauka " Publishing, Moscow" pageId="7" pageNumber="8" pagination="178 - 186" refString="Golovatch S. I. 1983. Millipedes (Diplopoda) of the fauna of Vietnam. In: Sokolov V. E. (ed.) Fauna and Ecology of the Animals of Vietnam: 178 - 186. " Nauka " Publishing, Moscow." title="Millipedes (Diplopoda) of the fauna of Vietnam" type="book chapter" volumeTitle="Fauna and Ecology of the Animals of Vietnam" year="1983">Golovatch 1983</bibRefCitation>
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: 181.
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</paragraph>
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<paragraph id="A06436E4FFD03324DDDE19A3FDF080DE" blockId="7.[189,1172,1214,1311]" box="[189,639,1249,1276]" pageId="7" pageNumber="8">
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<taxonomicName id="67DB4D67FFD03324DDDE19A3FEFE80D9" ID-CoL="4BBXS" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[189,369,1249,1275]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="92AFEAF6FFD03324DDDE19A3FEFE80D9" box="[189,369,1249,1275]" italics="true" pageId="7" pageNumber="8">Oxidus gracilis</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="C44A4B15FFD03324DCED19A3FDC080D9" author="Golovatch S. I." box="[398,591,1249,1275]" journalOrPublisher="Acta Zoologica Hungarica" pageId="7" pageNumber="8" pagination="53 - 77" part="30" refString="Golovatch S. I. 1984. Contribution to the millipede fauna of Vietnam (Diplopoda), II. Acta Zoologica Hungarica 30 (1 - 2): 53 - 77." title="Contribution to the millipede fauna of Vietnam (Diplopoda), II" type="journal article" year="1984">Golovatch 1984</bibRefCitation>
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: 54.
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</paragraph>
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<paragraph id="A06436E4FFD03324DDDE1846FBB9813D" blockId="7.[189,1172,1214,1311]" box="[189,1078,1284,1311]" pageId="7" pageNumber="8">
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<taxonomicName id="67DB4D67FFD03324DDDE1846FED8813C" ID-CoL="4BBXR" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[189,343,1284,1310]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gigas">
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<emphasis id="92AFEAF6FFD03324DDDE1846FED8813C" box="[189,343,1284,1310]" italics="true" pageId="7" pageNumber="8">Oxidus gigas</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="C44A4B15FFD03324DC171846FDD3813C" author="Enghoff H. & Golovatch S. & Nguyen D. A." box="[372,604,1284,1310]" journalOrPublisher="Arthropoda Selecta" pageId="7" pageNumber="8" pagination="29 - 43" part="13" refString="Enghoff H., Golovatch S. I. & Nguyen D. A. 2004. A review of the millipede fauna of Vietnam (Diplopoda). Arthropoda Selecta 13 (1 / 2): 29 - 43." title="A review of the millipede fauna of Vietnam (Diplopoda)" type="journal article" year="2004">
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Enghoff
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<emphasis id="92AFEAF6FFD03324DCB81847FD99813C" box="[475,534,1284,1310]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
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2004
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</bibRefCitation>
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: 39. —
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<bibRefCitation id="C44A4B15FFD03324DFDB1847FC66813C" author="Nguyen A. D. & Sierwald P." box="[696,1001,1284,1311]" journalOrPublisher="CheckList" pageId="7" pageNumber="8" pagination="1132 - 1353" part="9" refString="Nguyen A. D. & Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). CheckList 9: 1132 - 1353." title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)" type="journal article" year="2013">Nguyen & Sierwald 2013</bibRefCitation>
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: 1295.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="E8C1656FFFD03324DDDE1804FEBB81DA" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="A06436E4FFD03324DDDE1804FEB98142" blockId="7.[189,1399,1350,1528]" box="[189,310,1350,1376]" pageId="7" pageNumber="8">
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||
<heading id="FB2C8188FFD03324DDDE1804FEB98142" bold="true" box="[189,310,1350,1376]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="6">
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||
<emphasis id="92AFEAF6FFD03324DDDE1804FEB98142" bold="true" box="[189,310,1350,1376]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A06436E4FFD03324DDDE1836FEBB81DA" blockId="7.[189,1399,1350,1528]" pageId="7" pageNumber="8">
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This species is particularly similar to
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<taxonomicName id="67DB4D67FFD03324DF0A1836FC7A81AC" authority="(C. L. Koch, 1847)" authorityName="C.L. Koch" authorityYear="1847" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[617,1013,1396,1422]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="92AFEAF6FFD03324DF0A1836FC9381AC" box="[617,796,1396,1422]" italics="true" pageId="7" pageNumber="8">Oxidus gracilis</emphasis>
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(
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<bibRefCitation id="C44A4B15FFD03324DE491836FC6281AC" author="Koch C. L." box="[810,1005,1396,1422]" editor="Panzer G. W. F." journalOrPublisher="Bandchen, Regensburg" pageId="7" pageNumber="8" pagination="1 - 196" refString="Koch C. L. 1847. System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden. In: Panzer G. W. F. & Herrich-Schaffer A. (eds) Kritische Revision der Insectenfauna Deutschlands, III: 1 - 196. Bandchen, Regensburg." title="System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden" type="book chapter" volumeTitle="Kritische Revision der Insectenfauna Deutschlands" year="1847">C.L. Koch, 1847</bibRefCitation>
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)
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</taxonomicName>
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in body appearance and gonopod conformation, but differs in larger size and in details of gonopod structure: process
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<emphasis id="92AFEAF6FFD03324D9F418D5FB298193" box="[1175,1190,1431,1457]" italics="true" pageId="7" pageNumber="8">h</emphasis>
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narrow and long, pointed at the end and not serrated at distolateral margin; bases of process
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<emphasis id="92AFEAF6FFD03324D95118F9FBCE81F7" box="[1074,1089,1467,1493]" italics="true" pageId="7" pageNumber="8">h</emphasis>
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and spine
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<emphasis id="92AFEAF6FFD03324D9A218FEFB4381F7" box="[1217,1228,1468,1493]" italics="true" pageId="7" pageNumber="8">z</emphasis>
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less distinctly separated.
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD03324DDDE1B5DFDF3833A" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="A06436E4FFD03324DDDE1B5DFE25821B" blockId="7.[189,426,1567,1593]" box="[189,426,1567,1593]" pageId="7" pageNumber="8">
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||
<heading id="FB2C8188FFD03324DDDE1B5DFE25821B" bold="true" box="[189,426,1567,1593]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="2">
|
||
<emphasis id="92AFEAF6FFD03324DDDE1B5DFE25821B" bold="true" box="[189,426,1567,1593]" pageId="7" pageNumber="8">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFD03324DDDE1B0CFDF3833A" blockId="7.[189,1399,1614,1816]" pageId="7" pageNumber="8">VIETNAM: 1 ♂, 1 ♀, Lao Cai Prov., Sa Pa, Ham Rong Mts., limestone mountain, 1600 m, 16 Jul. 2006, Anh D. Nguyen, IEBR-113; 1 ♂, 6 ♀♀ (IEBR-Myr 516), 7 ♀♀ (IEBR-Myr 517), Ha Giang Prov., Bac Quang Distr., Duc Xuan Commune, Na Po village, limestone forest, under logs, 14−15 Apr. 2013, leg. Anh D. Nguyen; 1 ♂, 3 ♀♀, Vinh Phuc Prov., Tam Dao National Park, forest, 900−1000 m, Mar. 2012, leg. Hoang Long, IEBR-Myr 518; 2 ♂♂, 1 ♀, Lao Cai Prov., Co xan, in and around a cave, 1971, leg. Gy. Topál & I. Matskási, IEBR-H133.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8C1656FFFD03329DDDE1A7DFB878576" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="A06436E4FFD03324DDDE1A7DFEE4837B" blockId="7.[189,363,1855,1881]" box="[189,363,1855,1881]" pageId="7" pageNumber="8">
|
||
<heading id="FB2C8188FFD03324DDDE1A7DFEE4837B" bold="true" box="[189,363,1855,1881]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="2">
|
||
<emphasis id="92AFEAF6FFD03324DDDE1A7DFEE4837B" bold="true" box="[189,363,1855,1881]" pageId="7" pageNumber="8">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFD03324DDDE1A2CFB088389" blockId="7.[189,1398,1902,1964]" pageId="7" pageNumber="8">SIZE. Body length 31−34.5 mm (♂♂), 29.9−30.2 mm (♀); width of midbody pro- and metazonae 1.9−2.2 mm (♂), 2.2 mm (♀) and 2.8−3.2 mm (♂), 3.1− 3.2 mm (♀), respectively.</paragraph>
|
||
<paragraph id="A06436E4FFDF332BDDDE1C49FD37854E" blockId="8.[189,1399,267,364]" pageId="8" pageNumber="9">COLORATION. Generally castaneous brown, but anterior half of metaterga and posterior margin of prozonae darkish brown. Posterior half of metaterga, anterior margin of prozonae and pleura castaneous brown; paraterga, sterna and legs brownish yellow.</paragraph>
|
||
<paragraph id="A06436E4FFDF332BDDDE1CD6FCB085D1" blockId="8.[189,1399,403,500]" pageId="8" pageNumber="9">HEAD. Slightly narrower than collum; labrum sparsely setose; epicranial suture distinct, dividing frons into two equal parts. Antennae long and slender, not claviform, reaching segment 3 if stretched posteriorly. Antennomere 1<7<<2=3=4=5=6 in length.</paragraph>
|
||
<paragraph id="A06436E4FFDF332BDDDE1F59FACE8659" blockId="8.[189,1399,539,636]" pageId="8" pageNumber="9">COLLUM. Subequal to, or slightly narrower than collum, trapeziform; surface shining and smooth, without rugosity; setae broken, but traces of two rows: 3+3 close to anterior margin and 1+1 in middle. Paraterga large, ear-shaped with broadly rounded laterocaudal corners; lateral side with a setiferous incision.</paragraph>
|
||
<paragraph id="A06436E4FFDF332ADDDE1FE1FBFF85AD" blockId="8.[189,1398,675,737]" lastBlockId="9.[189,1399,267,399]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">In width, segment 4≤3<2<5–17, thereafter gradually tapering towards telson. Prozonae and metaterga shining, smooth, without rugosity. Metaterga with two rows of setae: 2+2 close to anterior margin and 2+2 close to posterior margin. Transverse sulcus starting on metatergum 4, well developed, reaching base of paraterga on metaterga 5−19, beaded at bottom on some caudal segments. Stricture between pro- and metazonae clearly distinct, broad and striolate at bottom. Pleura shining, smooth; pleurosternal carinae completely absent or poorly developed only on pre-gonopodal segments.</paragraph>
|
||
<caption id="F4A4666CFFDF332BDDDC1AEFFAF783C9" httpUri="https://zenodo.org/record/889863/files/figure.png" pageId="8" pageNumber="9" targetBox="[310,1280,812,1925]" targetPageId="8">
|
||
<paragraph id="A06436E4FFDF332BDDDC1AEFFAF783C9" blockId="8.[191,1400,1965,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="92AFEAF6FFDF332BDDDC1AEFFE8183E5" bold="true" box="[191,270,1965,1991]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
|
||
<taxonomicName id="67DB4D67FFDF332BDC7B1AEFFDFE83EA" authority="(Attems, 1953)" authorityName="Attems" authorityYear="1953" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[280,625,1965,1992]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="92AFEAF6FFDF332BDC7B1AEFFE3B83E5" box="[280,436,1965,1991]" italics="true" pageId="8" pageNumber="9">Oxidus gigas</emphasis>
|
||
(Attems, 1953)
|
||
</taxonomicName>
|
||
from Duc Xuan Commune, Ha Giang Prov., Vietnam.
|
||
<emphasis id="92AFEAF6FFDF332BD8691AEFFA9083E5" bold="true" box="[1290,1311,1965,1991]" pageId="8" pageNumber="9">A</emphasis>
|
||
. Entire body, length ca 34 mm.
|
||
<emphasis id="92AFEAF6FFDF332BDCB01A93FD8583C8" bold="true" box="[467,522,2000,2027]" pageId="8" pageNumber="9">B–C</emphasis>
|
||
. Segments 8–9–10.
|
||
<emphasis id="92AFEAF6FFDF332BDF951A93FC8583C9" bold="true" box="[758,778,2001,2027]" pageId="8" pageNumber="9">B</emphasis>
|
||
. ♂.
|
||
<emphasis id="92AFEAF6FFDF332BDE591A92FCC083C8" bold="true" box="[826,847,2000,2026]" pageId="8" pageNumber="9">C</emphasis>
|
||
. ♀. Scale bars = 1 mm. (photo by Anh Nguyen)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A06436E4FFDE332ADDDE1CF5FC6D8619" blockId="9.[189,1399,438,571]" pageId="9" pageNumber="10">PARATERGA. Well developed, lying subequal to or lower than metatergal surface. Calluses small, but obvious; lateral side with 2 setiferous incisions at ⅓ and ⅔ of its length.Anterior corner broadly rounded, but posterior corner acute, produced into a pointed projection on segments 15−17. Ozopores located behind second lateral incision of paraterga 5, 7, 9, 10, 12, 13, 15–19.</paragraph>
|
||
<paragraph id="A06436E4FFDE332ADDDE1F20FADE86BD" blockId="9.[189,1399,609,671]" pageId="9" pageNumber="10">EPIPROCT. Long, but broadly truncated, dorsoventrally Fattened, with two minute apical tubercles. Tip with four spinnerets. Hypoproct trapeziform, with two well separated, distolateral, setiferous knobs.</paragraph>
|
||
<paragraph id="A06436E4FFDE332ADDDE1F84FABE86C3" blockId="9.[189,1329,710,737]" box="[189,1329,710,737]" pageId="9" pageNumber="10">STERNA. Modestly setose, without modifcations, but with a minute cone caudally near each coxa.</paragraph>
|
||
<paragraph id="A06436E4FFDE332ADDDE1E4AFC178767" blockId="9.[189,1399,775,837]" pageId="9" pageNumber="10">LEG. Long and slender, about 1.6−1.7 (♂), 1.4−1.5 (♀) times as long as midbody height. Tarsal brushes absent. Prefemora not swollen. Femora without modifcations.</paragraph>
|
||
<paragraph id="A06436E4FFDE332ADDDE1E2EFE68805E" blockId="9.[189,1399,876,1148]" pageId="9" pageNumber="10">GONOPOD. Relatively complicated. Coxite cylindrical, long, subequal to femorite in length; distoventral part sparsely setose. Prefemorite densely setose, separated laterally from femorite by a distinct oblique sulcus. Femorite weakly twisted, grooved mesally, more or less slightly enlarged distally, without modifcations and demarcated from postfemoral region by a lateral sulcus. Postfemoral region shorter than femorite, with a lateral sub-pentagonal lamina l; spine z pointed; process h long and pointed, not serrated laterally; bases of process h and spine z not separated. Solenophore strongly spiral; distomesal lobule well developed and distinctly separated from solenophore tip. Solenomere Fagelliform, completely sheathed by solenophore.</paragraph>
|
||
<caption id="F4A4666CFFDE332ADDDE1AEFFA8E83C9" httpUri="https://zenodo.org/record/889865/files/figure.png" pageId="9" pageNumber="10" targetBox="[187,1397,1247,1920]" targetPageId="9">
|
||
<paragraph id="A06436E4FFDE332ADDDE1AEFFA8E83C9" blockId="9.[189,1399,1965,2027]" pageId="9" pageNumber="10">
|
||
<emphasis id="92AFEAF6FFDE332ADDDE1AEFFE8383E5" bold="true" box="[189,268,1965,1991]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
|
||
Right gonopod of
|
||
<taxonomicName id="67DB4D67FFDE332ADC8C1AEFFCC683EA" authority="(Attems, 1953)" authorityName="Attems" authorityYear="1953" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[495,841,1965,1992]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="92AFEAF6FFDE332ADC8C1AEFFD0483E5" box="[495,651,1965,1991]" italics="true" pageId="9" pageNumber="10">Oxidus gigas</emphasis>
|
||
(Attems, 1953)
|
||
</taxonomicName>
|
||
, from sample IEBR-Myr 113.
|
||
<emphasis id="92AFEAF6FFDE332AD9D41AEFFB4383E5" bold="true" box="[1207,1228,1965,1991]" pageId="9" pageNumber="10">A</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="92AFEAF6FFDE332ADDDE1A93FF5E83C9" bold="true" box="[189,209,2001,2027]" pageId="9" pageNumber="10">B</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="92AFEAF6FFDE332ADCE31A92FE1A83C8" bold="true" box="[384,405,2000,2026]" pageId="9" pageNumber="10">C</emphasis>
|
||
. Mesal view. The picture has been Fipped horizontally. Scale bars = 1 um.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A06436E4FFDD3329DDDE1C49FF728507" blockId="10.[189,253,267,293]" box="[189,253,267,293]" pageId="10" pageNumber="11">
|
||
<heading id="FB2C8188FFDD3329DDDE1C49FF728507" allCaps="true" bold="true" box="[189,253,267,293]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
|
||
<emphasis id="92AFEAF6FFDD3329DDDE1C49FF728507" bold="true" box="[189,253,267,293]" pageId="10" pageNumber="11">DNA</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFDD3329DDDE1C78FB878576" blockId="10.[189,1032,313,340]" box="[189,1032,313,340]" pageId="10" pageNumber="11">
|
||
COI and 16S barcode data (partial) are deposited in GenBank (
|
||
<tableCitation id="ED59035FFFDD3329DEFC1C7BFC788576" box="[927,1015,313,340]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Details of specimens analysed for DNA, including GenBank numbers." pageId="10" pageNumber="11">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8C1656FFFDD3329DDDE1C39FC3485E1" box="[189,955,379,452]" pageId="10" pageNumber="11" type="biology_ecology">
|
||
<paragraph id="A06436E4FFDD3329DDDE1C39FEA685B7" blockId="10.[189,297,379,405]" box="[189,297,379,405]" pageId="10" pageNumber="11">
|
||
<heading id="FB2C8188FFDD3329DDDE1C39FEA685B7" bold="true" box="[189,297,379,405]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="2">
|
||
<emphasis id="92AFEAF6FFDD3329DDDE1C39FEA685B7" bold="true" box="[189,297,379,405]" pageId="10" pageNumber="11">Habitats</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFDD3329DDDE1CEBFC3485E1" blockId="10.[189,955,425,452]" box="[189,955,425,452]" pageId="10" pageNumber="11">All material was found under leaf-litter, logs and decaying wood.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8C1656FFFDD3329DDDE1CA9FAF88616" box="[189,1399,491,564]" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="A06436E4FFDD3329DDDE1CA9FED98627" blockId="10.[189,342,491,517]" box="[189,342,491,517]" pageId="10" pageNumber="11">
|
||
<heading id="FB2C8188FFDD3329DDDE1CA9FED98627" bold="true" box="[189,342,491,517]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="2">
|
||
<emphasis id="92AFEAF6FFDD3329DDDE1CA9FED98627" bold="true" box="[189,342,491,517]" pageId="10" pageNumber="11">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFDD3329DDDE1F5BFAF88616" blockId="10.[189,1399,537,564]" box="[189,1399,537,564]" pageId="10" pageNumber="11">The species has only been recorded from northern Vietnam (Lao Cai, Ha Giang and Vinh Phuc Provinces).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8C1656FFFDD3329DDDE1F19FE508750" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="A06436E4FFDD3329DDDE1F19FEA08657" blockId="10.[189,303,603,629]" box="[189,303,603,629]" pageId="10" pageNumber="11">
|
||
<heading id="FB2C8188FFDD3329DDDE1F19FEA08657" bold="true" box="[189,303,603,629]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="2">
|
||
<emphasis id="92AFEAF6FFDD3329DDDE1F19FEA08657" bold="true" box="[189,303,603,629]" pageId="10" pageNumber="11">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFDD3329DDDE1FC8FD75872F" blockId="10.[189,1399,649,781]" pageId="10" pageNumber="11">
|
||
<bibRefCitation id="C44A4B15FFDD3329DDDE1FC8FEE58686" author="Attems C." box="[189,362,649,676]" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie" pageId="10" pageNumber="11" part="5" refString="Attems C. 1953. Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, Serie A - Zoologie 5 (3), Museum national d'Histoire naturelle, Paris." title="Myriopoden von Indochina: Expedition von C. Dawydoff (1938 - 1939)" type="journal volume" year="1953">Attems (1953)</bibRefCitation>
|
||
proposed a new subspecies
|
||
<taxonomicName id="67DB4D67FFDD3329DFA31FCBFBBD8681" authorityName="Attems" authorityYear="1953" box="[704,1074,649,675]" class="Diplopoda" family="Paradoxosomatidae" genus="Kalorthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="gracilis" subSpecies="gigas">
|
||
<emphasis id="92AFEAF6FFDD3329DFA31FCBFBBD8681" box="[704,1074,649,675]" italics="true" pageId="10" pageNumber="11">Kalorthomorpha gracilis gigas</emphasis>
|
||
</taxonomicName>
|
||
with a short note.
|
||
<bibRefCitation id="C44A4B15FFDD3329D8741FCBFEDF86E5" author="Enghoff H. & Golovatch S. & Nguyen D. A." journalOrPublisher="Arthropoda Selecta" pageId="10" pageNumber="11" pagination="29 - 43" part="13" refString="Enghoff H., Golovatch S. I. & Nguyen D. A. 2004. A review of the millipede fauna of Vietnam (Diplopoda). Arthropoda Selecta 13 (1 / 2): 29 - 43." title="A review of the millipede fauna of Vietnam (Diplopoda)" type="journal article" year="2004">
|
||
Enghoff
|
||
<emphasis id="92AFEAF6FFDD3329DDDE1FEFFF7686E4" box="[189,249,684,710]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
raised this subspecies to full rank as
|
||
<taxonomicName id="67DB4D67FFDD3329DE711FEEFC4786E4" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[786,968,684,710]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="92AFEAF6FFDD3329DE711FEEFC4786E4" box="[786,968,684,710]" italics="true" pageId="10" pageNumber="11">Oxidus gracilis</emphasis>
|
||
</taxonomicName>
|
||
, and also showed minor differences in size and gonopod process h. The species is fairly similar to
|
||
<taxonomicName id="67DB4D67FFDD3329DEC81F92FBA486C8" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[939,1067,720,746]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="92AFEAF6FFDD3329DEC81F92FBA486C8" box="[939,1067,720,746]" italics="true" pageId="10" pageNumber="11">O. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, but the molecular data has provided strong evidence to separate the species.
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFDD3329DDDE1E76FE508750" blockId="10.[189,1398,820,882]" pageId="10" pageNumber="11">
|
||
<bibRefCitation id="C44A4B15FFDD3329DDDE1E76FE05876C" author="Golovatch S. I." box="[189,394,820,846]" journalOrPublisher="Acta Zoologica Hungarica" pageId="10" pageNumber="11" pagination="53 - 77" part="30" refString="Golovatch S. I. 1984. Contribution to the millipede fauna of Vietnam (Diplopoda), II. Acta Zoologica Hungarica 30 (1 - 2): 53 - 77." title="Contribution to the millipede fauna of Vietnam (Diplopoda), II" type="journal article" year="1984">Golovatch (1984)</bibRefCitation>
|
||
misidentifed
|
||
<taxonomicName id="67DB4D67FFDD3329DF4E1E77FD27876C" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[557,680,820,847]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="92AFEAF6FFDD3329DF4E1E77FD27876C" box="[557,680,820,847]" italics="true" pageId="10" pageNumber="11">O. gracilis</emphasis>
|
||
</taxonomicName>
|
||
in Vietnam (sample IEBR-H133), which is currently corrected as
|
||
<taxonomicName id="67DB4D67FFDD3329DDBE1E1AFECE8753" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[221,321,856,882]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="92AFEAF6FFDD3329DDBE1E1AFECE8753" box="[221,321,856,882]" italics="true" pageId="10" pageNumber="11">O. gigas</emphasis>
|
||
</taxonomicName>
|
||
in this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |