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<document id="BF1B7F86919F737ADC7C3A63AA9AF0DF" ID-DOI="10.1163/18759866-20191416" ID-GBIF-Dataset="6388d839-a72d-4598-a141-053897bfc900" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF96B14F53679F26F6B7FC24" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.3:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="3" docTitle="Fastosarion mcdonaldi Stanisic 1993" docType="treatment" docVersion="9" lastPageNumber="430" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="430" updateTime="1731578815876" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="36DE49B7E0DD020A49C029A0886B7320">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="6650CC42A834C5D8F7AB955B9EEDD418">Hyman, Isabel T.</mods:namePart>
<mods:affiliation id="A1BFC51E3C8F383D3BFA16F78FCBC6E8">Australian Museum, 1 William St, Sydney NSW 2010, Australia Isabel. Hyman @ austmus. gov. au</mods:affiliation>
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<mods:namePart id="AC3C970475E84FED70F9C36113DAE360">Köhler, Frank</mods:namePart>
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<treatment id="03F32A40FF96B14F53679F26F6B7FC24" ID-DOI="http://doi.org/10.5281/zenodo.8366913" ID-GBIF-Taxon="213408543" ID-Zenodo-Dep="8366913" LSID="urn:lsid:plazi:treatment:03F32A40FF96B14F53679F26F6B7FC24" httpUri="http://treatment.plazi.org/id/03F32A40FF96B14F53679F26F6B7FC24" lastPageNumber="430" pageId="79" pageNumber="430">
<subSubSection id="C340C8DDFF96B14F53679F26F671FCB4" box="[93,518,821,848]" pageId="79" pageNumber="430" type="nomenclature">
<paragraph id="8BE59B56FF96B14F53679F26F671FCB4" blockId="79.[93,518,821,848]" box="[93,518,821,848]" pageId="79" pageNumber="430">
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<taxonomicName id="4C5AE0D5FF96B14F53679F26F671FCB4" ID-CoL="7RBRN" ID-ENA="2658036" authority="Stanisic, 1993" authorityName="Stanisic" authorityYear="1993" box="[93,518,821,848]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF96B14F53679F26F671FCB4" bold="true" box="[93,518,821,848]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F53679F26F51FFCAB" bold="true" box="[93,360,821,848]" italics="true" pageId="79" pageNumber="430">Fastosarion mcdonaldi</emphasis>
Stanisic, 1993
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF96B14F53589F4AF5EBFC8F" box="[98,412,858,884]" pageId="79" pageNumber="430" type="description">
<paragraph id="8BE59B56FF96B14F53589F4AF5EBFC8F" blockId="79.[98,609,858,1063]" box="[98,412,858,884]" pageId="79" pageNumber="430">
<figureCitation id="136187D3FF96B14F53589F4AF4C5FC8F" box="[98,178,858,884]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="79" pageNumber="430">Figs. 18</figureCitation>
,
<figureCitation id="136187D3FF96B14F53869F4AF490FC8F" box="[188,231,858,884]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="79" pageNumber="430">19H</figureCitation>
,
<figureCitation id="136187D3FF96B14F53CB9F4AF568FC8F" box="[241,287,858,884]" captionStart="FIGURE 20" captionStartId="69.[98,171,1087,1108]" captionTargetBox="[98,1157,188,1050]" captionTargetId="figure-74@69.[98,1157,188,1050]" captionTargetPageId="69" captionText="FIGURE 20 Fastosarion live images (Clade B). A-D.Fastosarion alyssa, A:AM C.512537, Mt Etna Caves; B:AM C.512480, Mt Ossa; C:AM C.512579, The Hummock. D: NMV F230748,Brandy Creek (Type locality of F.peterbrocki; A.Moussalli, NMV). E. Fastosarion ephelis (J. Stanisic).F.Fastosarion hannianus (QM image).G. Fastosarion mcdonaldi, Mt Dalrymple (QM image). H. Fastosarion rowani, AM C.512541,Dan Dan NP." figureDoi="http://doi.org/10.5281/zenodo.8343101" httpUri="https://zenodo.org/record/8343101/files/figure.png" pageId="79" pageNumber="430">20G</figureCitation>
,
<figureCitation id="136187D3FF96B14F52129F4AF50FFC8F" box="[296,376,858,884]" captionStart="FIGURE 22" captionStartId="71.[98,171,1710,1731]" captionTargetBox="[98,887,184,1677]" captionTargetId="figure-39@71.[98,887,184,1677]" captionTargetPageId="71" captionText="FIGURE 22 Spermatophores of Clade B species. A-B. F.alyssa, MO77158.C-D.F.deensis, MO85834.E-F. F.hannianus, MO72277.G-H.F.mcdonaldi, MO50434.I-J. F.minerva, MO78358." figureDoi="http://doi.org/10.5281/zenodo.8343105" httpUri="https://zenodo.org/record/8343105/files/figure.png" pageId="79" pageNumber="430">22GH</figureCitation>
,
<figureCitation id="136187D3FF96B14F52B89F4AF5EBFC8F" box="[386,412,858,884]" captionStart="FIGURE 28" captionStartId="80.[129,202,1223,1244]" captionTargetBox="[129,1167,188,1192]" captionTargetId="figure-8@80.[129,1187,188,1192]" captionTargetPageId="80" captionText="FIGURE 28 Genitalia of F.mcdonaldi, MO504354,Eungella NP. A. Reproductive system.Arrow shows the junction of the penis and epiphallus. B. Spermatophore.C.Penis interior. Scale bars 1 mm (BC), 2 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.8343117" httpUri="https://zenodo.org/record/8343117/files/figure.png" pageId="79" pageNumber="430">28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF96B14F53BB9F6EF4DDFBDC" pageId="79" pageNumber="430" type="reference_group">
<paragraph id="8BE59B56FF96B14F53BB9F6EF4DDFBDC" blockId="79.[98,609,858,1063]" pageId="79" pageNumber="430">
<taxonomicName id="4C5AE0D5FF96B14F53BB9F6EF62AFC63" ID-CoL="7R9JY" authority="Stanisic, 1993: 28" authorityName="Stanisic" authorityPageNumber="28" authorityYear="1993" box="[129,605,893,920]" class="Gastropoda" family="Helicarionidae" genus="Eungarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF96B14F53BB9F6EF50EFC6C" box="[129,377,893,919]" italics="true" pageId="79" pageNumber="430">Eungarion mcdonaldi</emphasis>
Stanisic, 1993: 28
</taxonomicName>
, figs. 1ab,
<figureCitation id="136187D3FF96B14F53DA9FB2F499FC47" box="[224,238,930,956]" captionStart="FIGURE 2" captionStartId="31.[98,171,1692,1713]" captionTargetBox="[98,1125,170,1660]" captionTargetId="figure-6@31.[98,1125,169,1660]" captionText="FIGURE 2 Majority-rule consensus tree based on Bayesian analysis of the concatenated data set of fragments of the mitochondrial genes 16S and COI. Ambiguous alignment sites in 16S removed using MAFFT. Numbers on branches indicate posterior probabilities. Scale bar indicating 40% of modelled sequence divergence. Shells shown to scale." figureDoi="http://doi.org/10.5281/zenodo.8343065" httpUri="https://zenodo.org/record/8343065/files/figure.png" pageId="79" pageNumber="430">2</figureCitation>
, 3af, 4ac;
<bibRefCitation id="EFCBE6A7FF96B14F52B19FB2F4CBFC1B" author="Hyman, I. T. &amp; Ponder, W. F." pageId="79" pageNumber="430" pagination="1 - 148" refId="ref41527" refString="Hyman, I. T. &amp; Ponder, W. F. (2010) A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family. Zootaxa, 2462, 1 - 148." type="journal article" year="2010">Hyman &amp; Ponder, 2010: 56</bibRefCitation>
, figs. 6LM, 13K, 14EF, 18C, 19C, 21C;
<bibRefCitation id="EFCBE6A7FF96B14F535E9FF9F5F8FBF8" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[100,399,1001,1028]" pageId="79" pageNumber="430" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 316317</bibRefCitation>
, 333;
<bibRefCitation id="EFCBE6A7FF96B14F52F29FF9F62DFBF8" author="Stanisic, J." box="[456,602,1001,1027]" pageId="79" pageNumber="430" pagination="43 - 51" refId="ref43100" refString="Stanisic, J. (2018) Description of a new species of semi-slug, Fastosarioncomerfordae sp. nov., from EungellaNationalPark, mid-easternQueensland (Gastropoda: Eupulmonata: Helicarionidae). Mem. Queensl. Mus. - Nature, 61, 43 - 51." type="journal article" year="2018">Stanisic, 2018</bibRefCitation>
: fig.
<specimenCount id="9D5C50DFFF96B14F53B3981DF4DDFBDC" box="[137,170,1037,1063]" pageId="79" pageNumber="430" type="generic">4F.</specimenCount>
</paragraph>
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<subSubSection id="C340C8DDFF96B14F53669849F6B7FC24" pageId="79" pageNumber="430" type="materials_examined">
<paragraph id="8BE59B56FF96B14F53669849F547FB88" blockId="79.[92,609,1113,1319]" box="[92,304,1113,1139]" pageId="79" pageNumber="430">Material examined</paragraph>
<paragraph id="8BE59B56FF96B14F5358986EF4D0FB24" blockId="79.[92,609,1113,1319]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F5358986EF4D0FB6C" box="[98,167,1150,1175]" italics="true" pageId="79" pageNumber="430">Types:</emphasis>
<materialsCitation id="3B32910BFF96B14F538E986DF4D0FB24" ID-GBIF-Occurrence="4413347305" collectionCode="QM" elevation="1000" location="Dalrymple Heights" municipality="Eungella National Park" pageId="79" pageNumber="430" specimenCode="MO43309" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E125F4FF96B14F538E986DF568FB6C" box="[180,287,1149,1175]" pageId="79" pageNumber="430" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED4B0393FF96B14F5215986DF52DFB6C" box="[303,346,1149,1175]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F525D986DF5A4FB6C" box="[359,467,1149,1175]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO43309</specimenCode>
(
<location id="8E85CD8DFF96B14F52D2986DF4CDFB40" LSID="urn:lsid:plazi:treatment:03F32A40FF96B14F53679F26F6B7FC24:8E85CD8DFF96B14F52D2986DF4CDFB40" municipality="Eungella National Park" name="Dalrymple Heights" pageId="79" pageNumber="430">Dalrymple Heights</location>
,
<collectingMunicipality id="6B81012CFF96B14F53FD98B1F5B8FB40" box="[199,463,1185,1211]" pageId="79" pageNumber="430">Eungella National Park</collectingMunicipality>
,
<quantity id="4CA236B3FF96B14F52E198B1F65AFB40" box="[475,557,1185,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="79" pageNumber="430" unit="m" value="1000.0">
<elevation id="00777C65FF96B14F52E198B1F65AFB40" box="[475,557,1185,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="79" pageNumber="430" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, ME QLD).
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF96B14F53BB98F9F5E0FAF8" blockId="79.[92,609,1113,1319]" box="[129,407,1257,1283]" pageId="79" pageNumber="430">
<materialsCitation id="3B32910BFF96B14F53BB98F9F5E4FAF8" ID-GBIF-Occurrence="4413347321" box="[129,403,1257,1283]" collectionCode="QM" pageId="79" pageNumber="430" specimenCode="MO43310" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E125F4FF96B14F53BB98F9F487FAF8" box="[129,240,1257,1283]" pageId="79" pageNumber="430" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED4B0393FF96B14F53C398F9F553FAF8" box="[249,292,1257,1283]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF96B14F521398F9F5E4FAF8" box="[297,403,1257,1283]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="79" pageNumber="430" type="Museum">MO43310</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE59B56FF96B14F53BB991DF5BCFADC" blockId="79.[92,609,1113,1319]" box="[129,459,1292,1319]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F53BB991DF53BFADD" box="[129,332,1292,1318]" italics="true" pageId="79" pageNumber="430">Non-type material:</emphasis>
See
<tableCitation id="C6D8AEEDFF96B14F5244991CF5B0FADC" box="[382,455,1292,1319]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="79" pageNumber="430" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
.
</paragraph>
<paragraph id="8BE59B56FF96B14F53669948F4BEFA89" blockId="79.[92,609,1368,1754]" box="[92,201,1368,1394]" pageId="79" pageNumber="430">Diagnosis</paragraph>
<paragraph id="8BE59B56FF96B14F5358996CF66FF969" blockId="79.[92,609,1368,1754]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F5358996CF526FA6D" box="[98,337,1404,1430]" italics="true" pageId="79" pageNumber="430">External morphology</emphasis>
: Shell (fig. 19H) small (11.311.6), orange amber, 3.23.5 whorls, flattened, thin, base uncalcified, membraneous. Shell sculpture very faint on protoconch, obsolete on teleoconch. Body (fig. 20G)
<quantity id="4CA236B3FF96B14F51259A1CF4E6F9B1" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="79" pageNumber="430" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
, reddish brown. Shell lappets and mantle lobes large with pigmented pustules, lappets joined on both sides by a narrow collar.
</paragraph>
<paragraph id="8BE59B56FF96B14F53BB9A8CF7CBFE1F" blockId="79.[92,609,1368,1754]" lastBlockId="79.[646,1157,171,484]" pageId="79" pageNumber="430">
<emphasis id="B92E4744FF96B14F53BB9A8CF532F94E" box="[129,325,1692,1718]" italics="true" pageId="79" pageNumber="430">Genital anatomy:</emphasis>
Genitalia (figs. 22GH, 28) with medium length vagina, very swollen; bursa copulatrix moderately short, duct distinct, bursa elongate oval. Penis long, swollen proximally, internally with 24 longitudinal pilasters, internal wall pustulose; 60% of penis contained in penial tunica. Epiphallus approx. equal length to penis, enters penis through small rounded verge; epiphallic caecum of moderate length; flagellum short, slender. Spermatophore simple.
</paragraph>
<paragraph id="8BE59B56FF96B14F51BA9E06F695FDCB" blockId="79.[640,1159,534,991]" box="[640,738,534,560]" pageId="79" pageNumber="430">Remarks</paragraph>
<paragraph id="8BE59B56FF96B14F51BC9E2BF6B7FC24" blockId="79.[640,1159,534,991]" pageId="79" pageNumber="430">
<taxonomicName id="4C5AE0D5FF96B14F51BC9E2BF7FDFDAF" authorityName="Stanisic" authorityYear="1993" box="[646,906,570,596]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF96B14F51BC9E2BF7FDFDAF" box="[646,906,570,596]" italics="true" pageId="79" pageNumber="430">Fastosarion mcdonaldi</emphasis>
</taxonomicName>
is restricted to the Clarke Ranges in Mideastern Qld, usually found above
<quantity id="4CA236B3FF96B14F50249E92F710FD66" box="[798,871,642,669]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="79" pageNumber="430" unit="m" value="900.0">900 m</quantity>
altitude (fig. 18). Several other semislugs are found in this region, including
<taxonomicName id="4C5AE0D5FF96B14F51DE9EDBF71CFD1F" box="[740,875,715,740]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="aquavitae">
<emphasis id="B92E4744FF96B14F51DE9EDBF71CFD1F" box="[740,875,715,740]" italics="true" pageId="79" pageNumber="430">F. aquavitae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AE0D5FF96B14F50419EDBF068FD1F" box="[891,1055,714,740]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="comerfordae">
<emphasis id="B92E4744FF96B14F50419EDBF068FD1F" box="[891,1055,714,740]" italics="true" pageId="79" pageNumber="430">F. comerfordae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AE0D5FF96B14F57149EDBF6E9FCF3" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="ephelis">
<emphasis id="B92E4744FF96B14F57149EDBF6E9FCF3" italics="true" pageId="79" pageNumber="430">F. ephelis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF96B14F51DB9EFFF748FCF3" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[737,831,750,776]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="tuljun">
<emphasis id="B92E4744FF96B14F51DB9EFFF748FCF3" box="[737,831,750,776]" italics="true" pageId="79" pageNumber="430">F. tuljun</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5AE0D5FF96B14F506B9EFFF027FCF3" authorityName="Stanisic" authorityYear="1993" box="[849,1104,750,776]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF96B14F506B9EFFF027FCF3" box="[849,1104,750,776]" italics="true" pageId="79" pageNumber="430">Fastosarion mcdonaldi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C5AE0D5FF96B14F50AF9F03F068FCD7" box="[917,1055,787,812]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="aquavitae">
<emphasis id="B92E4744FF96B14F50AF9F03F068FCD7" box="[917,1055,787,812]" italics="true" pageId="79" pageNumber="430">F. aquavitae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AE0D5FF96B14F570B9F03F695FCAB" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="comerfordae">
<emphasis id="B92E4744FF96B14F570B9F03F695FCAB" italics="true" pageId="79" pageNumber="430">F. comerfordae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF96B14F50259F26F7F2FCAB" box="[799,901,822,848]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="ephelis">
<emphasis id="B92E4744FF96B14F50259F26F7F2FCAB" box="[799,901,822,848]" italics="true" pageId="79" pageNumber="430">F. ephelis</emphasis>
</taxonomicName>
by its more flattened, reduced shell and smaller size, and from
<taxonomicName id="4C5AE0D5FF96B14F57499F4AF6B0FC6C" authorityName="Hyman &amp; Köhler" authorityYear="2019" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="79" pageNumber="430" phylum="Mollusca" rank="species" species="tuljun">
<emphasis id="B92E4744FF96B14F57499F4AF6B0FC6C" italics="true" pageId="79" pageNumber="430">F. tuljun</emphasis>
</taxonomicName>
by its larger size, pustulose shell lappets and more reduced shell with a membraneous base.
</paragraph>
</subSubSection>
</treatment>
</document>