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<document id="438313A8B43CBA24A5BA60BE3630E353" ID-CLB-Dataset="3586" ID-DOI="10.5852/ejt.2021.777.1561" ID-GBIF-Dataset="e8ade5ef-ebef-4519-85fe-50919b4b9fe1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5637112" ID-ZooBank="C86302D6-49F7-41A6-9BAC-94FDFED0D27B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635793154903" checkinUser="felipe" docAuthor="Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor &amp; Raspotnig, Günther" docDate="2021" docId="534F336D7706BD1BFD89FB90B945539D" docLanguage="en" docName="ejt-777_1-67.pdf" docOrigin="European Journal of Taxonomy 777" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nemastoma bidentatum Roewer 1914" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="AF764B157701BD1CFFBCFF85BC51553B" masterDocTitle="Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence" masterLastPageNumber="67" masterPageNumber="1" pageNumber="8" updateTime="1699467332017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="207472C696BEBFB66F1368BEE613EFE4">Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence</mods:title>
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<mods:affiliation id="0A472666D6415E2933A5CAAFA84EA115">Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška 160, 2000 Maribor, Slovenia.</mods:affiliation>
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<mods:namePart id="7A452321E02BD4ED5AC668AABE409961">Kozel, Peter</mods:namePart>
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<mods:affiliation id="8572DAA59192FC149808463F54D3EC5D">Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška 160, 2000 Maribor, Slovenia. &amp; Karst Research Institute IZRK ZRC SAZU, Titov trg 2, 6230 Postojna, Slovenia. &amp; Chair on Karst Education, University of Nova Gorica, Vipava, Slovenia.</mods:affiliation>
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<mods:namePart id="6F7AEB93187D64F4784045F0AB3F352C">Schaider, Miriam Gudrun</mods:namePart>
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<mods:namePart id="DCA88F51AD836311FA43FBE5D085F219">Komposch, Christian</mods:namePart>
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<mods:affiliation id="1F12120B545F8FEA6850F7C49EB3A751">Institut für Tierökologie und Naturraumplanung, Institute for Animal Ecology and Landscape Planning, Bergmanngasse 22, 8010 Graz, Austria.</mods:affiliation>
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<mods:affiliation id="97DA02D0D9ADBC11FA5F6A0F311E496E">Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška 160, 2000 Maribor, Slovenia. &amp; Faculty of Medicine, University of Maribor, Taborska ulica 8, 2000 Maribor, Slovenia. &amp; Faculty of Chemistry and Chemical Engineering, University of Maribor, Smetanova ulica 17, 2000 Maribor, Slovenia. &amp; Gottfried Schatz Research Center, Division of Cell Biology, Histology and Embryology, Medical University of Graz, Neue Stiftingtalstrasse 6, 8010 Graz, Austria.</mods:affiliation>
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<mods:namePart id="F77D943A9950F4D19960EAC2494A8247">Podlesnik, Jan</mods:namePart>
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<mods:affiliation id="E4E330D01998370E4A4DB0E96E144DA9">Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška 160, 2000 Maribor, Slovenia.</mods:affiliation>
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<mods:namePart id="41759BA1D9618B8CB1C589CDBCCDE9C7">Paušič, Igor</mods:namePart>
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<mods:affiliation id="6F91E575001B0D8B4D3F684F1D135BFA">Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška 160, 2000 Maribor, Slovenia.</mods:affiliation>
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<mods:namePart id="19DBCA0A56991BA33F397E855B4C77BB">Raspotnig, Günther</mods:namePart>
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<mods:affiliation id="EBBD1048211B7D24AEF323782E02690D">Institute of Biology, Karl-Franzens University, Universitätsplatz 2, 8010 Graz, Austria.</mods:affiliation>
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<mods:title id="D06D100673C82E2372E69AC616B85F42">European Journal of Taxonomy</mods:title>
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<mods:date id="18BBEA0E7247065FAD38F7690094BA1D">2021</mods:date>
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<taxonomicName id="1CE6F9F87706BD1BFD89FB90BFAE5114" authority="Roewer, 1914" authorityName="Roewer" authorityYear="1914" box="[565,1023,1044,1071]" class="Florideophyceae" family="Nemastomataceae" genus="Nemastoma" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="8" phylum="Rhodophyta" rank="species" species="bidentatum">
<emphasis id="E9925E697706BD1BFD89FB90BF055114" bold="true" box="[565,852,1044,1071]" italics="true" pageId="7" pageNumber="8">Nemastoma bidentatum</emphasis>
<bibRefCitation id="BF77FF8A7706BD1BFCE0FB90BFAE5114" author="Roewer C. F." box="[860,1023,1045,1071]" pageId="7" pageNumber="8" pagination="99 - 169" refId="ref59529" refString="Roewer C. F. 1914. Die Familien der Ischyropsalidae und Nemastomatidae der Opiliones-Palpatores. Archives fur Naturgeschichte 80 A (3): 99 - 169." type="journal article" year="1914">Roewer, 1914</bibRefCitation>
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<treatmentCitation id="5A47A46A7706BD1BFF01FBD2BE93514B" author="Roewer C. F." box="[189,706,1110,1136]" page="141" pageId="7" pageNumber="8" year="1914">
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<emphasis id="E9925E697706BD1BFF01FBD2BD84514B" box="[189,469,1110,1136]" italics="true" pageId="7" pageNumber="8">Nemastoma bidentatum</emphasis>
<bibRefCitation id="BF77FF8A7706BD1BFE62FBD3BE93514B" author="Roewer C. F." box="[478,706,1110,1136]" pageId="7" pageNumber="8" pagination="99 - 169" refId="ref59529" refString="Roewer C. F. 1914. Die Familien der Ischyropsalidae und Nemastomatidae der Opiliones-Palpatores. Archives fur Naturgeschichte 80 A (3): 99 - 169." type="journal article" year="1914">Roewer, 1914: 141</bibRefCitation>
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, figs 20a–d (original description) [partim;
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series also includes
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<emphasis id="E9925E697706BD1BFEEFFBFCBDE351A9" box="[339,434,1145,1170]" italics="true" pageId="7" pageNumber="8">N. triste</emphasis>
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].
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<taxonomicName id="1CE6F9F87706BD1BFF01FB3FBD8551EF" authorityName="Roewer" authorityYear="1914" box="[189,468,1210,1236]" class="Florideophyceae" family="Nemastomataceae" genus="Nemastoma" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="8" phylum="Rhodophyta" rank="species" species="bidentatum">
<emphasis id="E9925E697706BD1BFF01FB3FBD8551EF" box="[189,468,1210,1236]" italics="true" pageId="7" pageNumber="8">Nemastoma bidentatum</emphasis>
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–
<treatmentCitation id="5A47A46A7706BD1BFE49FB3FBE8551EF" author="Roewer C. F." box="[501,724,1209,1236]" page="658" pageId="7" pageNumber="8" year="1923">
<bibRefCitation id="BF77FF8A7706BD1BFE49FB3FBE8551EF" author="Roewer C. F." box="[501,724,1209,1236]" pageId="7" pageNumber="8" refId="ref59584" refString="Roewer C. F. 1923. Die Weberknechte der Erde. Gustav Fischer, Jena." type="book" year="1923">Roewer 1923: 658</bibRefCitation>
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, figs 817a–d [partim;
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series also includes
<taxonomicName id="1CE6F9F87706BD1BFAB9FB3FB93451E8" baseAuthorityName="C. L. Koch" baseAuthorityYear="1835" box="[1285,1381,1210,1235]" class="Florideophyceae" family="Nemastomataceae" genus="Nemastoma" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="8" phylum="Rhodophyta" rank="species" species="triste">
<emphasis id="E9925E697706BD1BFAB9FB3FB93451E8" box="[1285,1381,1210,1235]" italics="true" pageId="7" pageNumber="8">N. triste</emphasis>
</taxonomicName>
]. —
<treatmentCitation id="5A47A46A7706BD1BFEADFB59BEFA51CC" author="Gruber J. &amp; Martens J." box="[273,683,1244,1271]" page="141" pageId="7" pageNumber="8" year="1968">
<bibRefCitation id="BF77FF8A7706BD1BFEADFB59BEFA51CC" author="Gruber J. &amp; Martens J." box="[273,683,1244,1271]" pageId="7" pageNumber="8" pagination="137 - 172" refId="ref57425" refString="Gruber J. &amp; Martens J. 1968. Morphologie, Systematik und Okologie der Gattung Nemastoma C. L. Koch (s. str.) (Opiliones, Nemastomatidae). Senckenbergiana Biologica 49 (2): 137 - 172." type="journal article" year="1968">Gruber &amp; Martens 1968: 141–142</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="5A47A46A7706BD1BFC25FB58B8EA51CD" author="Martens J." box="[921,1211,1244,1271]" page="106" pageId="7" pageNumber="8" year="1978">
<bibRefCitation id="BF77FF8A7706BD1BFC25FB58B8EA51CD" author="Martens J." box="[921,1211,1244,1271]" pageId="7" pageNumber="8" pagination="1 - 464" refId="ref58493" refString="Martens J. 1978. Spinnentiere, Arachnida: Weberknechte, Opiliones. In: Senglaub F., Hannemann H. &amp; Schumann H. (eds) Die Tierwelt Deutschlands 64: 1 - 464. Fischer Verlag, Jena." type="journal article" year="1978">Martens 1978: 106–107</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="5A47A46A7706BD1BFB52FB59BD065022" author="Schonhofer A. L." page="36" pageId="7" pageNumber="8" year="2013">
<bibRefCitation id="BF77FF8A7706BD1BFB52FB59BD065022" author="Schonhofer A. L." pageId="7" pageNumber="8" pagination="1 - 68" refId="ref59736" refString="Schonhofer A. L. 2013. A taxonomic catalogue of the Dyspnoi Hansen and Sorensen, 1904 (Arachnida: Opiliones). Zootaxa 3679 (1): 1 - 68. https: // doi. org / 10.11646 / zootaxa. 3679.1.1" type="journal article" year="2013">Schönhofer 2013: 36</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="93FCD1F07706BD1BFF01FAC5B82950F5" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="DB59827B7706BD1BFF01FAC5BF8A5060" blockId="7.[189,987,1344,1371]" box="[189,987,1344,1371]" pageId="7" pageNumber="8">
<emphasis id="E9925E697706BD1BFF01FAC5BD675061" bold="true" box="[189,310,1344,1370]" pageId="7" pageNumber="8">Diagnosis</emphasis>
(according to
<bibRefCitation id="BF77FF8A7706BD1BFE5CFAC5BEAC5061" author="Gruber J. &amp; Martens J." box="[480,765,1344,1371]" pageId="7" pageNumber="8" pagination="137 - 172" refId="ref57425" refString="Gruber J. &amp; Martens J. 1968. Morphologie, Systematik und Okologie der Gattung Nemastoma C. L. Koch (s. str.) (Opiliones, Nemastomatidae). Senckenbergiana Biologica 49 (2): 137 - 172." type="journal article" year="1968">Gruber &amp; Martens 1968</bibRefCitation>
, slightly modified)
</paragraph>
<paragraph id="DB59827B7706BD1BFF01FAEBB82950F5" blockId="7.[189,1399,1390,1487]" pageId="7" pageNumber="8">
Species of the
<emphasis id="E9925E697706BD1BFED4FAEABE5E50B3" box="[360,527,1390,1416]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="1CE6F9F87706BD1BFED4FAEABDA250B3" authorityName="C. L. Koch" authorityYear="1836" box="[360,499,1391,1416]" class="Florideophyceae" family="Nemastomataceae" genus="Nemastoma" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="8" phylum="Rhodophyta" rank="genus">Nemastoma</taxonomicName>
b.
</emphasis>
complex with male Pa-Fe having either at least two conspicuous spines or long, rigid setae instead of some spines, or 6–7 irregular tubercles, or 5–11 denticles, and Pa-Fe min:max width 1:2.5–3.5, Rec sem with 1 or none tubular and 12–18 balloon-like vesicles.
</paragraph>
</subSubSection>
<subSubSection id="93FCD1F07706BD1BFF01FA70B945539D" pageId="7" pageNumber="8" type="discussion">
<paragraph id="DB59827B7706BD1BFF01FA70BD7E5334" blockId="7.[189,303,1525,1551]" box="[189,303,1525,1551]" pageId="7" pageNumber="8">
<heading id="801135177706BD1BFF01FA70BD7E5334" bold="true" box="[189,303,1525,1551]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="E9925E697706BD1BFF01FA70BD7E5334" bold="true" box="[189,303,1525,1551]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DB59827B7706BD1BFF01F9A1B945539D" blockId="7.[189,1399,1571,1702]" pageId="7" pageNumber="8">
To date, the species includes six subspecies:
<emphasis id="E9925E697706BD1BFD6EF9A1BF5A5306" box="[722,779,1571,1597]" italics="true" pageId="7" pageNumber="8">N. b.</emphasis>
<taxonomicName id="1CE6F9F87706BD1BFCA8F9A6B8195305" authority="Roewer, 1914" authorityName="Roewer" authorityYear="1914" box="[788,1096,1571,1598]" class="Florideophyceae" family="Nemastomataceae" genus="Nemastoma" kingdom="Plantae" order="Gigartinales" pageId="7" pageNumber="8" phylum="Rhodophyta" rank="species" species="bidentatum">
<emphasis id="E9925E697706BD1BFCA8F9A6BFC95306" box="[788,920,1571,1597]" italics="true" pageId="7" pageNumber="8">bidentatum</emphasis>
<bibRefCitation id="BF77FF8A7706BD1BFC1DF9A1B8195305" author="Roewer C. F." box="[929,1096,1572,1598]" pageId="7" pageNumber="8" pagination="99 - 169" refId="ref59529" refString="Roewer C. F. 1914. Die Familien der Ischyropsalidae und Nemastomatidae der Opiliones-Palpatores. Archives fur Naturgeschichte 80 A (3): 99 - 169." type="journal article" year="1914">Roewer, 1914</bibRefCitation>
</taxonomicName>
,
<emphasis id="E9925E697706BD1BFBEAF9A1B8AA5306" box="[1110,1275,1571,1597]" italics="true" pageId="7" pageNumber="8">N. b. sparsum</emphasis>
<bibRefCitation id="BF77FF8A7706BD1BFAB8F9A6BD36535B" author="Gruber J. &amp; Martens J." pageId="7" pageNumber="8" pagination="137 - 172" refId="ref57425" refString="Gruber J. &amp; Martens J. 1968. Morphologie, Systematik und Okologie der Gattung Nemastoma C. L. Koch (s. str.) (Opiliones, Nemastomatidae). Senckenbergiana Biologica 49 (2): 137 - 172." type="journal article" year="1968">Gruber &amp; Martens, 1968</bibRefCitation>
, and the newly discovered
<emphasis id="E9925E697706BD1BFD24F9C2BF7B535B" box="[664,810,1606,1632]" italics="true" pageId="7" pageNumber="8">N. b. gruberi</emphasis>
Novak, Slana Novak, Kozel &amp; Raspotnig
<taxonomicNameLabel id="F2A1E3127706BD1BFAB1F9C2B923535A" box="[1293,1394,1607,1633]" pageId="7" pageNumber="8" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
,
<emphasis id="E9925E697706BD1BFF01F9EFBD3653B8" box="[189,359,1641,1667]" italics="true" pageId="7" pageNumber="8">N. b. martensi</emphasis>
Novak, Slana Novak &amp; Raspotnig
<taxonomicNameLabel id="F2A1E3127706BD1BFCACF9EFBF2B53BF" box="[784,890,1642,1668]" pageId="7" pageNumber="8" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
,
<emphasis id="E9925E697706BD1BFC3BF9EFB87F53B8" box="[903,1070,1641,1667]" italics="true" pageId="7" pageNumber="8">N. b. schmidti</emphasis>
Novak, Raspotnig &amp; Slana Novak
<taxonomicNameLabel id="F2A1E3127706BD1BFEAFF909BD28539D" box="[275,377,1676,1702]" pageId="7" pageNumber="8" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
and
<emphasis id="E9925E697706BD1BFE0DF908BED3539D" box="[433,642,1676,1702]" italics="true" pageId="7" pageNumber="8">N. b. sneznikensis</emphasis>
Novak, Komposch, Slana Novak &amp; Raspotnig
<taxonomicNameLabel id="F2A1E3127706BD1BFB12F909B945539D" box="[1198,1300,1676,1702]" pageId="7" pageNumber="8" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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