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<document id="A851986D04D287906CFA60BFEAF68D41" ID-DOI="10.11646/zootaxa.4019.1.25" ID-GBIF-Dataset="cc992da2-e091-46e5-86ae-5e28d0693feb" ID-ISSN="1175-5326" ID-Zenodo-Dep="242689" ID-ZooBank="C17868C6-847E-4578-B258-32158F87F43C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267192001" checkinUser="plazi" docAuthor="Wong, Eunice &amp; Hutchings, Pat" docDate="2015" docId="03F99517FFED293AFF2C56FFA0C02407" docLanguage="en" docName="zt04019p744.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphictene lizardensis Wong &amp; Hutchings, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="738" masterDocId="FFC0ED6FFFEF293FFFBB5668A2562017" masterDocTitle="New records of Pectinariidae (Polychaeta) from Lizard Island, Great Barrier Reef, Australia and the description of two new species" masterLastPageNumber="744" masterPageNumber="733" pageNumber="735" updateTime="1698610842274" updateUser="plazi">
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<mods:title id="7EBBDC8870F901140B55FD62823349DB">New records of Pectinariidae (Polychaeta) from Lizard Island, Great Barrier Reef, Australia and the description of two new species</mods:title>
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<mods:namePart id="31507545484091D8BF34BA9CE7A48017">Wong, Eunice</mods:namePart>
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<treatment id="03F99517FFED293AFF2C56FFA0C02407" ID-DOI="http://doi.org/10.5281/zenodo.6093555" ID-GBIF-Taxon="119638264" ID-Zenodo-Dep="6093555" LSID="urn:lsid:plazi:treatment:03F99517FFED293AFF2C56FFA0C02407" httpUri="http://treatment.plazi.org/id/03F99517FFED293AFF2C56FFA0C02407" lastPageId="5" lastPageNumber="738" pageId="2" pageNumber="735">
<subSubSection id="C34A778AFFED293DFF2C56FFA35C20C4" pageId="2" pageNumber="735" type="nomenclature">
<paragraph id="8BEF2401FFED293DFF2C56FFA3AF20A5" blockId="2.[151,505,151,211]" box="[151,505,151,178]" pageId="2" pageNumber="735">
<heading id="D0A7936DFFED293DFF2C56FFA3AF20A5" bold="true" box="[151,505,151,178]" fontSize="11" level="1" pageId="2" pageNumber="735" reason="1">
<emphasis id="B924F813FFED293DFF2C56FFA3AF20A5" bold="true" box="[151,505,151,178]" pageId="2" pageNumber="735">
<taxonomicName id="4C505F82FFED293DFF2C56FFA3E620A6" ID-CoL="D28J" box="[151,432,151,177]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="735" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFED293DFF2C56FFA3E620A6" bold="true" box="[151,432,151,177]" italics="true" pageId="2" pageNumber="735">Amphictene lizardensis</emphasis>
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<taxonomicNameLabel id="A2174568FFED293DFE0C56F0A3AF20A5" box="[439,505,152,178]" pageId="2" pageNumber="735" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BEF2401FFED293DFF2C56D2A35C20C4" blockId="2.[151,505,151,211]" box="[151,266,186,211]" pageId="2" pageNumber="735">
(
<figureCitation id="136B3884FFED293DFF2456D2A35720C4" box="[159,257,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1635,1657]" captionStartId-1="3.[151,250,1726,1748]" captionStartId-2="4.[151,250,1673,1695]" captionTargetBox-0="[151,1436,590,1609]" captionTargetBox-1="[151,1436,193,1704]" captionTargetBox-2="[226,1370,178,1657]" captionTargetId-0="figure@2.[151,1436,590,1610]" captionTargetId-1="figure@3.[151,1436,193,1705]" captionTargetId-2="figure@4.[226,1370,178,1658]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Amphictene lizardensis n. sp., stained with methyl green. A. Ventral view of entire animal, paratype, AM W. 47429; B. Dorsal view of entire animal, holotype, AM W. 47432. Scale bars: A B = 0.1 mm." captionText-1="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." captionText-2="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri-0="https://zenodo.org/record/242690/files/figure.png" httpUri-1="https://zenodo.org/record/242691/files/figure.png" httpUri-2="https://zenodo.org/record/242692/files/figure.png" pageId="2" pageNumber="735">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFED293DFF2C576AA33F21D8" pageId="2" pageNumber="735" type="materials_examined">
<paragraph id="8BEF2401FFED293DFF2C576AA33F21D8" blockId="2.[151,1437,258,535]" pageId="2" pageNumber="735">
<emphasis id="B924F813FFED293DFF2C576AA3DD210C" bold="true" box="[151,395,258,283]" pageId="2" pageNumber="735">Material examined.</emphasis>
<typeStatus id="54EB9AA3FFED293DFE27576AA05F210C" box="[412,521,258,283]" pageId="2" pageNumber="735" type="holotype">Holotype</typeStatus>
: AM W.47432, MI QLD 2194, Lizard Island, Watsons Bay,
<geoCoordinate id="EE6442C6FFED293DFAA6576AA7CA210C" box="[1309,1436,258,283]" direction="south" orientation="latitude" pageId="2" pageNumber="735" precision="15" value="-14.657222">14°39'26&quot;S</geoCoordinate>
<geoCoordinate id="EE6442C6FFED293DFF2C574EA34C2128" box="[151,282,294,319]" direction="east" orientation="longitude" pageId="2" pageNumber="735" precision="15" value="145.45084">145°27'3&quot;E</geoCoordinate>
,
<quantity id="4CA889E4FFED293DFE9C574EA33D2129" box="[295,363,294,319]" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="735" unit="m" value="6.5">6.5 m</quantity>
, coll. P. Hutchings &amp; M. Capa,
<date id="FFEE02C1FFED293DFD62574FA1242128" box="[729,882,295,319]" pageId="2" pageNumber="735" value="2010-08-28">28 Aug 2010</date>
, CReefs, gravid with coelomic gametes,
<quantity id="4CA889E4FFED293DFAFE574EA7CB2129" box="[1349,1437,294,319]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="735" unit="mm" value="5.7">5.7 mm</quantity>
long,
<quantity id="4CA889E4FFED293DFF6C5722A3782175" box="[215,302,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="735" unit="mm" value="1.6">1.6 mm</quantity>
wide anteriorly,
<quantity id="4CA889E4FFED293DFE565723A0122175" box="[493,580,331,355]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="735" unit="mm" value="0.7">0.7 mm</quantity>
wide posteriorly.
<typeStatus id="54EB9AA3FFED293DFCAB5723A1D42174" box="[784,898,331,355]" pageId="2" pageNumber="735" type="paratype">Paratypes</typeStatus>
: AM W.47429, MI QLD 2194 (2, 1 gravid with coelomic gametes), same locality as
<typeStatus id="54EB9AA3FFED293DFDF55706A0E52190" box="[590,691,366,391]" pageId="2" pageNumber="735" type="holotype">holotype</typeStatus>
,
<quantity id="4CA889E4FFED293DFD7F5706A1072191" box="[708,849,366,391]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.5" metricValueMin="3.1" pageId="2" pageNumber="735" unit="mm" value="3.8" valueMax="4.5" valueMin="3.1">3.14.5 mm</quantity>
long,
<quantity id="4CA889E4FFED293DFC185706A6662191" box="[931,1072,366,391]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="2" pageNumber="735" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
wide anteriorly,
<quantity id="4CA889E4FFED293DFB455707A70C2191" box="[1278,1370,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="735" unit="mm" value="0.8">0.8 mm</quantity>
wide posteriorly; AM W.47433, MI QLD 2194, same,
<quantity id="4CA889E4FFED293DFD7D57FBA14821BD" box="[710,798,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="735" unit="mm" value="2.8">2.8 mm</quantity>
long,
<quantity id="4CA889E4FFED293DFCDC57FBA1E821BD" box="[871,958,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="735" unit="mm" value="1.2">1.2 mm</quantity>
wide anteriorly,
<quantity id="4CA889E4FFED293DFB3B57FAA68121BD" box="[1152,1239,402,427]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="735" unit="mm" value="0.6">0.6 mm</quantity>
wide posteriorly, mounted for SEM.
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFED293AFF7C57B2A109220F" lastPageId="5" lastPageNumber="738" pageId="2" pageNumber="735" type="description">
<paragraph id="8BEF2401FFED293DFF7C57B2A0242200" blockId="2.[151,1437,258,535]" pageId="2" pageNumber="735">
<emphasis id="B924F813FFED293DFF7C57B2A30F21E4" bold="true" box="[199,345,474,499]" pageId="2" pageNumber="735">Description.</emphasis>
Preserved specimen pale cream in colour, small, conical in shape (
<figureCitation id="136B3884FFED293DFBFB57B2A62821E4" box="[1088,1150,474,499]" captionStart="FIGURE 1" captionStartId="2.[151,250,1635,1657]" captionTargetBox="[151,1436,590,1609]" captionTargetId="figure@2.[151,1436,590,1610]" captionTargetPageId="2" captionText="FIGURE 1. Amphictene lizardensis n. sp., stained with methyl green. A. Ventral view of entire animal, paratype, AM W. 47429; B. Dorsal view of entire animal, holotype, AM W. 47432. Scale bars: A B = 0.1 mm." httpUri="https://zenodo.org/record/242690/files/figure.png" pageId="2" pageNumber="735">Fig. 1</figureCitation>
AB). Tube composed of cemented sand grains and shell fragments.
</paragraph>
<caption id="DF2F7489FFED293DFF2C500BA60D2680" httpUri="https://zenodo.org/record/242690/files/figure.png" pageId="2" pageNumber="735" targetBox="[151,1436,590,1609]" targetPageId="2">
<paragraph id="8BEF2401FFED293DFF2C500BA60D2680" blockId="2.[151,1436,1635,1687]" pageId="2" pageNumber="735">
<emphasis id="B924F813FFED293DFF2C500BA34E266E" bold="true" box="[151,280,1635,1657]" pageId="2" pageNumber="735">FIGURE 1.</emphasis>
<taxonomicName id="4C505F82FFED293DFE9F500BA042266E" box="[292,532,1635,1657]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="735" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFED293DFE9F500BA042266E" box="[292,532,1635,1657]" italics="true" pageId="2" pageNumber="735">Amphictene lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFED293DFD9B500CA00B266E" box="[544,605,1636,1657]" pageId="2" pageNumber="735" rank="species">n. sp.</taxonomicNameLabel>
, stained with methyl green. A. Ventral view of entire animal, paratype, AM W.47429; B. Dorsal view of entire animal, holotype, AM W.47432. Scale bars: AB = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEF2401FFED293DFF7C50B2A30127F8" blockId="2.[151,1436,1754,2031]" pageId="2" pageNumber="735">
Rim of cephalic veil with 11 (1013) long, narrow cirri, tips rounded and slightly expanded. Cephalic veil completely free from operculum forming dorsal semi-circle around the numerous buccal tentacles (
<figureCitation id="136B3884FFED293DFA8B5096A72E2700" box="[1328,1400,1790,1815]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@3.[151,1436,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="2" pageNumber="735">Fig. 2</figureCitation>
A). Buccal tentacles conspicuous, markedly wide and deeply grooved, inverted V shaped (
<figureCitation id="136B3884FFED293DFB26514AA6B3272C" box="[1181,1253,1826,1851]" captionStart="FIGURE 1" captionStartId="2.[151,250,1635,1657]" captionTargetBox="[151,1436,590,1609]" captionTargetId="figure@2.[151,1436,590,1610]" captionTargetPageId="2" captionText="FIGURE 1. Amphictene lizardensis n. sp., stained with methyl green. A. Ventral view of entire animal, paratype, AM W. 47429; B. Dorsal view of entire animal, holotype, AM W. 47432. Scale bars: A B = 0.1 mm." httpUri="https://zenodo.org/record/242690/files/figure.png" pageId="2" pageNumber="735">Figs 1</figureCitation>
A, 2A). Raised opercular margin well developed, divided into 15 (1319) triangular lappets. Operculum with 14 (1314) pairs of long paleae, yellow-gold, curved dorsally, thick at base and flatten towards distal end, tips extended (
<figureCitation id="136B3884FFED293DFA885102A72C2794" box="[1331,1402,1898,1923]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@3.[151,1436,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="2" pageNumber="735">Fig. 2</figureCitation>
B), filiform when viewed under SEM (
<figureCitation id="136B3884FFED293DFDA651E6A00827B0" box="[541,606,1934,1959]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="2" pageNumber="735">Fig. 3</figureCitation>
A). A stalked spherical structure was noted between the two sets of paleae, resembling a balloon (
<figureCitation id="136B3884FFED293DFE2451DAA3BD27DC" box="[415,491,1970,1995]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@3.[151,1436,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="2" pageNumber="735">Figs 2</figureCitation>
B, 3A). Length of the structure approximately half of immediately neighbouring (shortest) paleae.
</paragraph>
<caption id="DF2F7489FFEC293CFF2C50D6A0472727" httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="3" pageNumber="736" targetBox="[151,1436,193,1704]" targetPageId="3">
<paragraph id="8BEF2401FFEC293CFF2C50D6A0472727" blockId="3.[151,1436,1726,1840]" pageId="3" pageNumber="736">
<emphasis id="B924F813FFEC293CFF2C50D6A34426C3" bold="true" box="[151,274,1726,1748]" pageId="3" pageNumber="736">FIGURE 2.</emphasis>
<taxonomicName id="4C505F82FFEC293CFEA250D6A05526C3" box="[281,515,1726,1748]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="736" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEC293CFEA250D6A05526C3" box="[281,515,1726,1748]" italics="true" pageId="3" pageNumber="736">Amphictene lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEC293CFDB250D7A01726C3" box="[521,577,1727,1748]" pageId="3" pageNumber="736" rank="species">n. sp.</taxonomicNameLabel>
, stained with methyl green, holotype, AM W.47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: AD = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEF2401FFEC293CFF7C51E9A64227F5" blockId="3.[151,1436,1921,2018]" pageId="3" pageNumber="736">
Tentacular cirri arise on anterior margin of segment 2. Segment 2 without anterodorsal lobe. Two pairs of comb-like branchiae on segments 3 and 4, situated laterally and consisting of loose flat lamellae (
<figureCitation id="136B3884FFEC293CFB4451CCA71F27AA" box="[1279,1353,1956,1981]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@3.[151,1436,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="3" pageNumber="736">Figs 2</figureCitation>
C, 3B). Anterior pair situated more ventrally and almost twice as large as posterior pair.
</paragraph>
<caption id="DF2F7489FFEB293BFF2C50E1A3B427C6" httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="4" pageNumber="737" targetBox="[226,1370,178,1657]" targetPageId="4">
<paragraph id="8BEF2401FFEB293BFF2C50E1A3B427C6" blockId="4.[151,1436,1673,2001]" pageId="4" pageNumber="737">
<emphasis id="B924F813FFEB293BFF2C50E1A3452688" bold="true" box="[151,275,1673,1695]" pageId="4" pageNumber="737">FIGURE 3.</emphasis>
SEM images of
<taxonomicName id="4C505F82FFEB293BFE0550E1A0FC2688" box="[446,682,1673,1695]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="737" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEB293BFE0550E1A0FC2688" box="[446,682,1673,1695]" italics="true" pageId="4" pageNumber="737">Amphictene lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEB293BFD0B50E2A0BE2688" box="[688,744,1674,1695]" pageId="4" pageNumber="737" rank="species">n. sp.</taxonomicNameLabel>
, paratype, AM W.47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of
<taxonomicName id="4C505F82FFEB293BFDB350CFA0D626AA" box="[520,640,1703,1725]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="737" phylum="Annelida" rank="genus">
<emphasis id="B924F813FFEB293BFDB350CFA0D626AA" box="[520,640,1703,1725]" italics="true" pageId="4" pageNumber="737">Amphictene</emphasis>
</taxonomicName>
species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 36 longitudinal rows, each row with 69 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: AC, FG = 0.1 mm, B (insert), DE = 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BEF2401FFEA293AFF7C56FFA0E620C2" blockId="5.[151,1437,151,1040]" pageId="5" pageNumber="738">Chaetigers 1 and 2 with broad anteroventral lobe, with that of chaetiger 1 more prominent. Anterior margin of lobes smooth. Nephridial papillae not observed.</paragraph>
<paragraph id="8BEF2401FFEA293AFF7C56B7A762219F" blockId="5.[151,1437,151,1040]" pageId="5" pageNumber="738">
Chaetigers 13 (segments 57) with notopodia and notochaetae only. Chaetigers 416 biramous with notopodia, neuropodia, notochaetae and neurochaetae (
<figureCitation id="136B3884FFEA293AFCAD576CA10A210B" box="[790,860,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
C). Chaetiger 17 with notopodia and notochaetae only. All notochaetae finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wing (
<figureCitation id="136B3884FFEA293AFF245724A2B72173" box="[159,225,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
D). Neuropodia wedge-shaped, erect tori with numerous neurochaetae. Neurochaetal uncini with major teeth arranged in 36 longitudinal rows, each row with 69 teeth, with size of teeth declining basally (
<figureCitation id="136B3884FFEA293AFB6B5707A745219F" box="[1232,1299,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
E).
</paragraph>
<paragraph id="8BEF2401FFEA293AFF7C57FCA109220F" blockId="5.[151,1437,151,1040]" pageId="5" pageNumber="738">
Posterior 5 segments fused to form a flattened plate or scaphe, broader than long, or as broad as long (
<figureCitation id="136B3884FFEA293AFA8157FCA72A21BB" box="[1338,1404,404,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
F). Scaphe with anal flap and dorsal papilla, scaphal margins crenulated and lobed (
<figureCitation id="136B3884FFEA293AFBA257DFA60D21C7" box="[1049,1115,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
FG). Small circular ciliated patches scattered across scaphal lobes, visible only under SEM (
<figureCitation id="136B3884FFEA293AFCC357B4A1EB21E3" box="[888,957,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1673,1695]" captionTargetBox="[226,1370,178,1657]" captionTargetId="figure@4.[226,1370,178,1658]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Amphictene lizardensis n. sp., paratype, AM W. 47433. A. Antero-dorsal view, opercular rim with triangular lappets, characteristic of Amphictene species, paleae thick, distally curved and extended to form filiform tips; B. Ventral view of anterior end of specimen, buccal tentacles conspicuous and markedly wide, two pairs of branchiae inserted laterally on segments 3 and 4, consisting of loose flat lamellae, chaetae commence from segment 5; C. Ventro-lateral view of specimen, neuropodia with neurochaetae commencing from chaetiger 4; D. Notochaetae of chaetiger 4 finely hirsute, some with smooth tip capillaries and some with broad distal pectinate wings; E. Neurochaetae of thoracic chaetiger, uncini arranged in 3 6 longitudinal rows, each row with 6 9 teeth; F. Dorsal view of posterior end, scaphe as broad as long, scaphal hooks broad based with pointed tips, most pairs are lost in this specimen; G. Lateral view of posterior end, scaphal margin crenulated forming lobes. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, nec = neurochaetae, noc = notochaetae, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hook, ss = unknown spherical structure. Scale bars: A C, F G = 0.1 mm, B (insert), D E = 0.01 mm." httpUri="https://zenodo.org/record/242692/files/figure.png" pageId="5" pageNumber="738">Fig. 3</figureCitation>
G). Scaphal hooks present, 4 pairs, broad, golden or yellow-brown, tips curved and pointed (
<figureCitation id="136B3884FFEA293AFD735797A146220F" box="[712,784,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@3.[151,1436,193,1705]" captionTargetPageId="3" captionText="FIGURE 2. Amphictene lizardensis n. sp., stained with methyl green, holotype, AM W. 47432. A. Ventral view of anterior end; B. Dorsal view of anterior end; C. Lateral view of anterior end; D. Dorsal view of posterior end. Abbreviations: br = branchiae, bt = buccal tentacles, cv = cephalic veil, or = opercular rim, p = paleae, s = scaphe, sh = scaphal hooks, ss = unknown spherical structure. Scale bars: A D = 0.1 mm." httpUri="https://zenodo.org/record/242691/files/figure.png" pageId="5" pageNumber="738">Figs 2</figureCitation>
D, 3F).
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFEA293AFF7C544CA1642368" pageId="5" pageNumber="738" type="discussion">
<paragraph id="8BEF2401FFEA293AFF7C544CA1642368" blockId="5.[151,1437,151,1040]" pageId="5" pageNumber="738">
<emphasis id="B924F813FFEA293AFF7C544CA36D222A" bold="true" box="[199,315,548,573]" pageId="5" pageNumber="738">Remarks.</emphasis>
This new species
<taxonomicName id="4C505F82FFEA293AFDB0544CA158222B" box="[523,782,548,572]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEA293AFDB0544CA158222B" box="[523,782,548,572]" italics="true" pageId="5" pageNumber="738">Amphictene lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEA293AFCAD544DA105222B" box="[790,851,549,572]" pageId="5" pageNumber="738" rank="species">n. sp.</taxonomicNameLabel>
is characterized by 4 pairs of robust or broad based scaphal hooks with curved point tips and the arrangement of uncini teeth in 36 rows.
<bibRefCitation id="EFC159F0FFEA293AFBCE542FA7CA2277" author="Hutchings" box="[1141,1436,583,608]" pageId="5" pageNumber="738" refString="Hutchings, P. &amp; Peart, P. (2002) A review of the genera of Pectinariidae (Polychaeta) together with a description of the Australian Fauna. Records of the Australian Museum, 54, 99 - 127. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.54.2002.1356" type="journal article" year="2002">Hutchings &amp; Peart (2002)</bibRefCitation>
provided a summary of the diagnostic feature of all known species of
<taxonomicName id="4C505F82FFEA293AFC125404A67B2293" box="[937,1069,620,644]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="genus">
<emphasis id="B924F813FFEA293AFC125404A67B2293" box="[937,1069,620,644]" italics="true" pageId="5" pageNumber="738">Amphictene</emphasis>
</taxonomicName>
(see Table 1,
<bibRefCitation id="EFC159F0FFEA293AFB705404A28722BF" author="Hutchings" pageId="5" pageNumber="738" refString="Hutchings, P. &amp; Peart, P. (2002) A review of the genera of Pectinariidae (Polychaeta) together with a description of the Australian Fauna. Records of the Australian Museum, 54, 99 - 127. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.54.2002.1356" type="journal article" year="2002">Hutchings &amp; Peart 2002</bibRefCitation>
) and a new species from the Gulf of
<collectingCountry id="F3476491FFEA293AFD3954E7A08122BF" box="[642,727,655,680]" name="Mexico" pageId="5" pageNumber="738">Mexico</collectingCountry>
has since been described (García-Garza &amp; de
<bibRefCitation id="EFC159F0FFEA293AFB4B54E7A28222DB" author="Leon-Gonzalez" pageId="5" pageNumber="738" refString="Garcia-Garza, M. E. &amp; de Leon-Gonzalez, J. A. (2014) A new species of Amphictene (Annelida, Pectinariidae) from the Gulf of Mexico, with a redescription of Amphictene guatemalensis (Nilsson, 1928). Zookeys, 367, 1 - 9. http: // dx. doi. org / 10.3897 / zookeys. 367.6038" type="journal article" year="2014">León-González 2014</bibRefCitation>
).
<taxonomicName id="4C505F82FFEA293AFF5054DCA3A522DB" box="[235,499,692,716]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEA293AFF5054DCA3A522DB" box="[235,499,692,716]" italics="true" pageId="5" pageNumber="738">Amphictene lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEA293AFE4554DDA06922DB" box="[510,575,693,716]" pageId="5" pageNumber="738" rank="species">n. sp.</taxonomicNameLabel>
most closely resembles
<taxonomicName id="4C505F82FFEA293AFCDF54DCA6B522DA" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[868,1251,692,717]" class="Polychaeta" family="Terebellidae" genus="Amphitrite" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="auricoma">
<emphasis id="B924F813FFEA293AFCDF54DCA1A522DB" box="[868,1011,692,716]" italics="true" pageId="5" pageNumber="738">A. auricoma</emphasis>
(
<bibRefCitation id="EFC159F0FFEA293AFBB354DCA68C22DA" author="Muller" box="[1032,1242,692,717]" pageId="5" pageNumber="738" refString="Muller, O. F. (1776) Zoologicae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum characteres, nomina et synonyma imprimis popularium. typis Hallageriis, Havniae (Copenhagen), xxxii, 282 pp, plates published in 1777." type="book" year="1776">O.F. Müller, 1776</bibRefCitation>
)
</taxonomicName>
described from
<collectingCountry id="F3476491FFEA293AFF2C54BFA2A922E7" box="[151,255,727,752]" name="Denmark" pageId="5" pageNumber="738">Denmark</collectingCountry>
in terms of number of cirri on cephalic veil and on the opercular rim. However,
<taxonomicName id="4C505F82FFEA293AFB6154B0A73D22F8" box="[1242,1387,728,751]" class="Polychaeta" family="Terebellidae" genus="Amphitrite" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="auricoma">
<emphasis id="B924F813FFEA293AFB6154B0A73D22F8" box="[1242,1387,728,751]" italics="true" pageId="5" pageNumber="738">A. auricoma</emphasis>
</taxonomicName>
has significantly more scaphal hooks (818 pairs) than
<taxonomicName id="4C505F82FFEA293AFD685494A13C2303" box="[723,874,764,788]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEA293AFD685494A13C2303" box="[723,874,764,788]" italics="true" pageId="5" pageNumber="738">A. lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEA293AFCCA5495A1FA2303" box="[881,940,765,788]" pageId="5" pageNumber="738" rank="species">n. sp.</taxonomicNameLabel>
(4 pairs). The same character can be used to distinguish
<taxonomicName id="4C505F82FFEA293AFEA15548A3E7232F" box="[282,433,800,824]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEA293AFEA15548A3E7232F" box="[282,433,800,824]" italics="true" pageId="5" pageNumber="738">A. lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEA293AFE015548A3A12320" box="[442,503,800,823]" pageId="5" pageNumber="738" rank="species">n. sp.</taxonomicNameLabel>
from the Australian species
<taxonomicName id="4C505F82FFEA293AFC875548A1FE232F" box="[828,936,800,824]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="favona">
<emphasis id="B924F813FFEA293AFC875548A1FE232F" box="[828,936,800,824]" italics="true" pageId="5" pageNumber="738">A. favona</emphasis>
</taxonomicName>
with 16 pairs of scaphal hooks.
<taxonomicName id="4C505F82FFEA293AFAA35548A2BA234B" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="uniloba">
<emphasis id="B924F813FFEA293AFAA35548A2BA234B" italics="true" pageId="5" pageNumber="738">Amphictene uniloba</emphasis>
</taxonomicName>
, the other Australian species, has 410 pairs of scaphal hooks, but these are fine, in contrast to the broad based shape with curved pointed tips in
<taxonomicName id="4C505F82FFEA293AFDEC5500A0B82397" box="[599,750,872,896]" class="Polychaeta" family="Pectinariidae" genus="Amphictene" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="738" phylum="Annelida" rank="species" species="lizardensis" status="sp. nov.">
<emphasis id="B924F813FFEA293AFDEC5500A0B82397" box="[599,750,872,896]" italics="true" pageId="5" pageNumber="738">A. lizardensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2174568FFEA293AFD4D5500A1642368" box="[758,818,872,895]" pageId="5" pageNumber="738" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFEA293AFF7C55E4A0E723DF" pageId="5" pageNumber="738" type="etymology">
<paragraph id="8BEF2401FFEA293AFF7C55E4A69923B2" blockId="5.[151,1437,151,1040]" box="[199,1231,908,933]" pageId="5" pageNumber="738">
<emphasis id="B924F813FFEA293AFF7C55E4A31823B2" bold="true" box="[199,334,908,933]" pageId="5" pageNumber="738">Etymology.</emphasis>
The species is named after Lizard Island where the species was described from.
</paragraph>
<paragraph id="8BEF2401FFEA293AFF7C55C7A0E723DF" blockId="5.[151,1437,151,1040]" box="[199,689,943,968]" pageId="5" pageNumber="738">
<emphasis id="B924F813FFEA293AFF7C55C7A37C23DF" bold="true" box="[199,298,943,968]" pageId="5" pageNumber="738">Habitat.</emphasis>
Shallow subtidal sandy substrates.
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFEA293AFF7C55BCA60B23FA" box="[199,1117,980,1005]" pageId="5" pageNumber="738" type="materials_examined">
<paragraph id="8BEF2401FFEA293AFF7C55BCA60B23FA" blockId="5.[151,1437,151,1040]" box="[199,1117,980,1005]" pageId="5" pageNumber="738">
<emphasis id="B924F813FFEA293AFF7C55BCA33323FA" bold="true" box="[199,357,980,1005]" pageId="5" pageNumber="738">
<typeStatus id="54EB9AA3FFEA293AFF7C55BCA35423FA" box="[199,258,980,1005]" pageId="5" pageNumber="738">Type</typeStatus>
locality.
</emphasis>
Queensland: Lizard Island, Watsons Bay,
<geoCoordinate id="EE6442C6FFEA293AFCF355BCA19F23FA" box="[840,969,980,1005]" direction="south" orientation="latitude" pageId="5" pageNumber="738" precision="15" value="-14.657222">14°39'26&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6442C6FFEA293AFC6E55BCA60F23FA" box="[981,1113,980,1005]" direction="east" orientation="longitude" pageId="5" pageNumber="738" precision="15" value="145.45084">145°27'3&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A778AFFEA293AFF7C559FA0C02407" box="[199,662,1015,1040]" pageId="5" pageNumber="738" type="distribution">
<paragraph id="8BEF2401FFEA293AFF7C559FA0C02407" blockId="5.[151,1437,151,1040]" box="[199,662,1015,1040]" pageId="5" pageNumber="738">
<emphasis id="B924F813FFEA293AFF7C559FA3362407" bold="true" box="[199,352,1015,1040]" pageId="5" pageNumber="738">Distribution.</emphasis>
Lizard Island, Queensland.
</paragraph>
</subSubSection>
</treatment>
</document>