253 lines
32 KiB
XML
253 lines
32 KiB
XML
<document id="FC9A717879F37C4CFABB77499F5496C5" ID-DOI="10.2478/aemnp-2018-0013" ID-GBIF-Dataset="7b94dd56-d3a7-4462-8da2-72ceb277835f" ID-ISSN="1804-6487" ID-Zenodo-Dep="3699163" ID-ZooBank="BF2CDC12-C555-4187-BFAB-274C9C674A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583496890333" checkinUser="tatiana" docAuthor="Löbl, Ivan" docDate="2018" docId="03F94B1B2179FF8A645FE28A01AD85BC" docLanguage="en" docName="ActaEntMusNatPra.58.1.151-165.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Baeocera inoptata Löbl 2018, sp. nov." docType="treatment" docVersion="12" lastPageNumber="152" masterDocId="FFC033632178FF8B656AE114055B8309" masterDocTitle="A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="165" masterPageNumber="151" pageNumber="152" updateTime="1698725294610" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="C8DEF1FAAB6EE0A7A3F5FCE7A9B346CE">A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
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<mods:name id="C1B45D657881E20754569DB1B967B3DD" type="personal">
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<mods:roleTerm id="869282FB377664A8C2AC20D84D7B925D">Author</mods:roleTerm>
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<mods:namePart id="3F3D505737E8E80A9B850515501B01E9">Löbl, Ivan</mods:namePart>
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<mods:affiliation id="4969CC1712CDA09B1FA7670860BA41E8">Muséum d’histoire naturelle, Route de Malagnou 1, CH- 1208 Geneva, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="9FD34D3E3AF2215D9788A131CCDB99CF" type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
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<mods:typeOfResource id="EE49611AD91C55C2173E62CAD7F2F090">text</mods:typeOfResource>
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<mods:title id="513CA0289C29668CEBD4024C517F32A5">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="91BF879EE7485E9D7BAD946D1CEA71E9">
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<mods:date id="A67A71D5B3D53ACAD0A78C6C198A0713">2018</mods:date>
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<mods:number id="1DFB6851FB6E4182C9985F1ECABFD287">2018-05-30</mods:number>
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<mods:number id="D81F0156DF95A83BDF34C83285C0C188">58</mods:number>
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<mods:start id="78842BCA58EF1DA59E7C08516D242432">151</mods:start>
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<mods:classification id="514B5AD691F9DCAA1D6229DF3845666E">journal article</mods:classification>
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<mods:identifier id="FE179764D79A57E2395364FF52F23343" type="DOI">10.2478/aemnp-2018-0013</mods:identifier>
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<treatment id="03F94B1B2179FF8A645FE28A01AD85BC" ID-DOI="http://doi.org/10.5281/zenodo.3706167" ID-GBIF-Taxon="162467412" ID-Zenodo-Dep="3706167" LSID="urn:lsid:plazi:treatment:03F94B1B2179FF8A645FE28A01AD85BC" httpUri="http://treatment.plazi.org/id/03F94B1B2179FF8A645FE28A01AD85BC" lastPageNumber="152" pageId="1" pageNumber="152">
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<subSubSection id="C34AA9862179FF8A645FE28A070180BD" box="[309,602,925,949]" pageId="1" pageNumber="152" type="nomenclature">
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<paragraph id="8BEFFA0D2179FF8A645FE28A070180BD" blockId="1.[309,602,925,974]" box="[309,602,925,949]" pageId="1" pageNumber="152">
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<heading id="D0A74D612179FF8A645FE28A070180BD" box="[309,602,925,949]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="152" reason="3">
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<emphasis id="B924261F2179FF8A645FE28A070180BD" bold="true" box="[309,602,925,949]" pageId="1" pageNumber="152">
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<taxonomicName id="4C50818E2179FF8A645FE28A04A680BD" ID-CoL="KCD4" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[309,509,925,949]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="inoptata" status="sp. nov.">
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<emphasis id="B924261F2179FF8A645FE28A04A680BD" bold="true" box="[309,509,925,949]" italics="true" pageId="1" pageNumber="152">Baeocera inoptata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2179B642179FF8A676EE289070180BD" box="[516,602,925,948]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34AA9862179FF8A64F1E2AF04AF80C7" box="[411,500,955,974]" pageId="1" pageNumber="152" type="description">
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<paragraph id="8BEFFA0D2179FF8A64F1E2AF04AF80C7" blockId="1.[309,602,925,974]" box="[411,500,955,974]" pageId="1" pageNumber="152">
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(
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<figureCitation id="136BE6882179FF8A64CBE2AF04B680C7" box="[417,493,955,974]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="1" pageNumber="152">Figs 1–5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34AA9862179FF8A65FDE2F104238724" pageId="1" pageNumber="152" type="materials_examined">
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<paragraph id="8BEFFA0D2179FF8A65FDE2F104238724" blockId="1.[151,760,997,1203]" pageId="1" pageNumber="152">
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<emphasis id="B924261F2179FF8A65FDE2F1044880F1" bold="true" box="[151,275,997,1016]" pageId="1" pageNumber="152">
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<typeStatus id="54EB44AF2179FF8A65FDE2F1059F80F1" box="[151,196,997,1016]" pageId="1" pageNumber="152">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F347BA9D2179FF8A6471E2F1042A80F1" box="[283,369,997,1016]" name="Indonesia" pageId="1" pageNumber="152">Indonesia</collectingCountry>
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,
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<collectingRegion id="499434EF2179FF8A6416E2F1075D80F1" box="[380,518,997,1016]" country="Indonesia" name="Sulawesi Selatan" pageId="1" pageNumber="152">South Sulawesi</collectingRegion>
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, Gowa District,
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<quantity id="4CA857E82179FF8A67F0E2F1079380F1" box="[666,712,997,1016]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="152" unit="km" value="6.0">6 km</quantity>
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E of Malino, Gunung Bawakaraeng [Mt.] Area, near Lembanna Base camp, ca.
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<geoCoordinate id="EE649CCA2179FF8A65DFE50E04518724" box="[181,266,1050,1069]" degrees="05" direction="south" minutes="15.4" orientation="latitude" pageId="1" pageNumber="152" precision="92" value="-5.2566667">
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05°15.4
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<emphasis id="B924261F2179FF8A6592E50E05A68724" box="[248,253,1050,1069]" italics="true" pageId="1" pageNumber="152">′</emphasis>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="EE649CCA2179FF8A6479E50E042F8724" box="[275,372,1050,1069]" degrees="119" direction="east" minutes="54.5" orientation="longitude" pageId="1" pageNumber="152" precision="92" value="119.90833">
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119°54.5
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<emphasis id="B924261F2179FF8A640AE50E043E8724" box="[352,357,1050,1069]" italics="true" pageId="1" pageNumber="152">′</emphasis>
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E
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</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34AA9862179FF8A65FDE521071887BA" pageId="1" pageNumber="152" type="materials_examined">
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<paragraph id="8BEFFA0D2179FF8A65FDE521071887BA" blockId="1.[151,760,997,1203]" pageId="1" pageNumber="152">
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<emphasis id="B924261F2179FF8A65FDE52104418741" bold="true" box="[151,282,1077,1096]" pageId="1" pageNumber="152">Type material.</emphasis>
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<materialsCitation id="3B38F0502179FF8A6477E52107FB8791" collectingDate="2015-02-11" collectingDateMax="2015-02-13" collectingDateMin="2015-02-11" collectionCode="NMPC" collectorName="J. Hajek & J. Sumpich" country="Indonesia" elevation="1520" latitude="-5.2566667" location="Gowa Distr" longLatPrecision="130" longitude="119.90833" municipality="Gowa Distr" pageId="1" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="S SULAWESI" typeStatus="holotype">
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<typeStatus id="54EB44AF2179FF8A6477E521042A874F" box="[285,369,1077,1096]" pageId="1" pageNumber="152" type="holotype">HOLOTYPE</typeStatus>
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: (
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<collectionCode id="ED4162C82179FF8A64F9E521048F8741" box="[403,468,1077,1096]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="152">NMPC</collectionCode>
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),
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||
<collectingCountry id="F347BA9D2179FF8A648BE521070C8741" box="[481,599,1077,1096]" name="Indonesia" pageId="1" pageNumber="152">INDONESIA</collectingCountry>
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||
,
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<collectingRegion id="499434EF2179FF8A6734E52107E0874F" box="[606,699,1077,1096]" pageId="1" pageNumber="152">S SULAWESI</collectingRegion>
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:
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||
<collectingMunicipality id="6B8B60772179FF8A67AFE5210598876A" pageId="1" pageNumber="152">Gowa Distr</collectingMunicipality>
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. /
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<locationDeviation id="9A5DE32F2179FF8A65BEE54407D08774" pageId="1" pageNumber="152">
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<quantity id="4CA857E82179FF8A65BEE54405A5876A" box="[212,254,1104,1123]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="152" unit="km" value="6.0">6 km</quantity>
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||
E of Malino, Gn. Bawakaraeng Area / border of gardens and mixed forest / (dominant Pinus) nr Lembanna Base camp
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||
</locationDeviation>
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||
/
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||
<geoCoordinate id="EE649CCA2179FF8A67F5E57E07A88774" box="[671,755,1130,1149]" degrees="05" direction="south" minutes="15.4" orientation="latitude" pageId="1" pageNumber="152" precision="92" value="-5.2566667">
|
||
05°15.4
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||
<emphasis id="B924261F2179FF8A6788E57E07BC8774" box="[738,743,1130,1149]" italics="true" pageId="1" pageNumber="152">′</emphasis>
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||
S
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||
</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE649CCA2179FF8A65FDE59105AC8791" box="[151,247,1157,1176]" degrees="119" direction="east" minutes="54.5" orientation="longitude" pageId="1" pageNumber="152" precision="92" value="119.90833">
|
||
119°54.5
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||
<emphasis id="B924261F2179FF8A658EE59105B28791" box="[228,233,1157,1176]" italics="true" pageId="1" pageNumber="152">′</emphasis>
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E
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</geoCoordinate>
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||
,
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||
<quantity id="4CA857E82179FF8A6594E59104658791" box="[254,318,1157,1176]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="152" unit="m" value="1520.0">
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<elevation id="007D1D3E2179FF8A6594E59104658791" box="[254,318,1157,1176]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="152" unit="m" value="1520.0">1520 m</elevation>
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</quantity>
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||
/
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<collectorName id="26A59FDB2179FF8A6421E59104A78791" box="[331,508,1157,1176]" pageId="1" pageNumber="152">J.Hájek & J.Šumpich</collectorName>
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leg.,
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<date id="FFEEDCCD2179FF8A6743E59107C78791" box="[553,668,1157,1176]" pageId="1" pageNumber="152" value="2015-02-11" valueMax="2015-02-13" valueMin="2015-02-11">
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<collectingDate id="EFAA25252179FF8A6743E59107C78791" box="[553,668,1157,1176]" pageId="1" pageNumber="152" value="2015-02-11" valueMax="2015-02-13" valueMin="2015-02-11">11-13.ii.2015</collectingDate>
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</date>
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||
.
|
||
</materialsCitation>
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||
<materialsCitation id="3B38F0502179FF8A67C9E591071887BA" collectingDate="2015-02-11" collectingDateMax="2015-02-13" collectingDateMin="2015-02-11" collectionCode="MHNG" collectorName="J. Hajek & J. Sumpich" country="Indonesia" elevation="1520" latitude="-5.2566667" location="Gowa Distr" longLatPrecision="130" longitude="119.90833" municipality="Gowa Distr" pageId="1" pageNumber="152" specimenCount="5" specimenCount-female="5" stateProvince="S SULAWESI" typeStatus="paratype">
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||
<typeStatus id="54EB44AF2179FF8A67C9E59107A9879F" box="[675,754,1157,1176]" pageId="1" pageNumber="152" type="paratype">PARATYPE</typeStatus>
|
||
:
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||
<specimenCount id="9D5631842179FF8A65FDE5B405EC87BA" box="[151,183,1180,1203]" pageId="1" pageNumber="152" type="female">1 ♀</specimenCount>
|
||
(
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||
<collectionCode id="ED4162C82179FF8A65A8E5B4045387BA" box="[194,264,1184,1203]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="152" type="Museum">MHNG</collectionCode>
|
||
), with the same data as the holotype.
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||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA9862179FF8A65FDE5DF0688803C" pageId="1" pageNumber="152" type="description">
|
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<paragraph id="8BEFFA0D2179FF8A65FDE5DF0072829C" blockId="1.[151,760,1227,2082]" lastBlockId="1.[828,1437,190,1717]" pageId="1" pageNumber="152">
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<emphasis id="B924261F2179FF8A65FDE5DF044187EB" bold="true" box="[151,282,1227,1250]" pageId="1" pageNumber="152">Description.</emphasis>
|
||
Length
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||
<quantity id="4CA857E82179FF8A6406E5DF049387EB" box="[364,456,1227,1250]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="1" pageNumber="152" unit="mm" value="2.03">2.03 mm</quantity>
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||
, width
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||
<quantity id="4CA857E82179FF8A6764E5DF073187EB" box="[526,618,1227,1250]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="152" unit="mm" value="1.42">1.42 mm</quantity>
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||
. Body convex dorsally; black, elytral apices and apical abdominal segments blackish-brown, mouth-parts ochreous, femora dark reddish-brown near bases and apices, blackish in middle, tibiae dark reddish-brown, tarsi ochreous, lighter than tibiae, antennomeres I to III ochreous, following antennomeres darkened, brown. Eyes large. Length/width ratios of antennomeres as: III 20/10: IV 20/8: V 32/8: VI 35/15: VII 55/20: VIII 43/16: IX 57/20: X 55/ 23 (
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<figureCitation id="136BE6882179FF8A6724E4DF07D686EB" box="[590,653,1483,1506]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="1" pageNumber="152">Fig. 1</figureCitation>
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||
; antennae of the
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||
<typeStatus id="54EB44AF2179FF8A65BDE4FF0469850B" box="[215,306,1515,1538]" pageId="1" pageNumber="152" type="holotype">holotype</typeStatus>
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||
broken off,
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||
<typeStatus id="54EB44AF2179FF8A64DAE4FF0751850B" box="[432,522,1515,1538]" pageId="1" pageNumber="152" type="paratype">paratype</typeStatus>
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||
with one antenna only, lacking apical antennomere). Lateral contours of pronotum and elytra separately rounded. Pronotum with lateral margin carinae concealed near angles, discal punctation very fine. Tip of scutellum exposed. Elytra weakly narrowed apically, middle section of lateral margins straight, lateral margin carinae exposed in dorsal view, sutural striae curved at base and extending laterally to form basal striae joined with lateral striae, adsutural areas flat. Elytral punctation on inner halves of disc somewhat coarser than pronotal punctation and finer than on outer discal halves. Hind wings fully developed. Hypomera very finely punctate. Mesoventrite with very shallow mesal stria, lacking large marginal punctures. Mesepimera large, slightly narrowed mesally, about 5 times as long as wide and 6 times as long as intervals to mesocoxae. Median part of metaventrite convex, smooth in middle, with several coarse punctures near metacoxal process. Lateral parts of metaventrite appearing impunctate. Metacoxal process flat, submesocoxal areas
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<quantity id="4CA857E82179FF8A6160E1AA013C83DC" box="[1034,1127,190,213]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="152" unit="mm" value="0.03">0.03 mm</quantity>
|
||
, with marginal punctures coarse, not elongate, extended to inner margins of mesepimera. Metanepisterna flat, about
|
||
<quantity id="4CA857E82179FF8A6127E1EA01F2821C" box="[1101,1193,254,277]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="152" unit="mm" value="0.13">0.13 mm</quantity>
|
||
wide, with suture deep, slightly sinuate. Abdominal ventrite I not microsculptured, with coarse, not elongate basal punctures, lacking basal rugosity; following ventrites distinctly punctate; ventrite VI with punctulate microsculpture. Tibiae straight.
|
||
</paragraph>
|
||
<paragraph id="8BEFFA0D2179FF8A6630E08A00C681DC" blockId="1.[828,1437,190,1717]" pageId="1" pageNumber="152">
|
||
<emphasis id="B924261F2179FF8A6630E08A014F82BC" bold="true" box="[858,1044,414,437]" italics="true" pageId="1" pageNumber="152">Male characters.</emphasis>
|
||
Protarsomeres I–III distinctly widened, narrower than apices of protibiae. Mesotarsomeres I and II slightly widened. Aedeagus (
|
||
<figureCitation id="136BE6882179FF8A61A6E0CA007682FC" box="[1228,1325,478,501]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="1" pageNumber="152">Figs 2–4</figureCitation>
|
||
)
|
||
<quantity id="4CA857E82179FF8A6056E0CA00C782FC" box="[1340,1436,478,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="152" unit="mm" value="0.63">0.63 mm</quantity>
|
||
long, strongly sclerotized, asymmetrical. Apical process of median lobe short, robust, overlapped by basal bulb. Internal sac with complex sclerites and two apical tufts of spine-like structures. Right parameres simple, bent in middle, in apical half almost evenly wide seen in ventral view, sinuate and widened in apical section in lateral view; left paramere lacking apophyse, wide, expanded ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BEFFA0D2179FF8A6630E3CA0688803C" blockId="1.[828,1437,190,1717]" pageId="1" pageNumber="152">
|
||
<emphasis id="B924261F2179FF8A6630E3CA016A81FC" bold="true" box="[858,1073,734,757]" italics="true" pageId="1" pageNumber="152">Female characters.</emphasis>
|
||
Protarsomeres not widened. Gonocoxite almost straight, evenly narrow, with subapical pores (
|
||
<figureCitation id="136BE6882179FF8A66EEE20A069E803C" box="[900,965,798,821]" captionStart="Figs 1–19. 1–5" captionStartId="3.[151,188,1957,1976]" captionTargetBox="[163,1426,411,1929]" captionTargetId="figure@3.[151,1439,398,1942]" captionTargetPageId="3" captionText="Figs 1–19. 1–5 – Baeocera inoptata sp. nov.: 1 – antennomeres 3 to 10; 2–4 – aedeagus in ventral, lateral and dorsal views (laterally impressed basal buld is an artefact), scale 0.2 mm; 5 – gonocoxite. 6–12 – Scaphisoma ancora sp. nov.: 6, 7 – aedeagus in dorsal and lateral views; 8 – male protibia and protarsus in lateral view; 9 – male mesotibia in lateral view; 10 – male ventrite 6; 11 – male tergite 9; 12 – gonocoxite. 13–19 – S. hulai sp. nov.: 13 – male tergite 9; 14 – male protarsus in dorsal view; 15 – male mesotarsus; 16, 17 – aedeagus in ventral and lateral views; 18 – male ventrite 6; 19 – gonocoxite. Scale bars: 0.1 mm (Figs 1, 5, 12, 19); 0.2 mm (Figs 2–4, 6–9, 14–17); 0.3 mm (10–11, 13, 18)." figureDoi="http://doi.org/10.5281/zenodo.3699165" httpUri="https://zenodo.org/record/3699165/files/figure.png" pageId="1" pageNumber="152">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA9862179FF8A6656E22A01F0855C" pageId="1" pageNumber="152" type="diagnosis">
|
||
<paragraph id="8BEFFA0D2179FF8A6656E22A01F0855C" blockId="1.[828,1437,190,1717]" pageId="1" pageNumber="152">
|
||
<emphasis id="B924261F2179FF8A6656E22A0175805C" bold="true" box="[828,1070,830,853]" pageId="1" pageNumber="152">Differential diagnosis.</emphasis>
|
||
Only a single species of
|
||
<taxonomicName id="4C50818E2179FF8A605BE22B00CE805F" authorityName="Erichson" authorityYear="1845" box="[1329,1429,831,854]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924261F2179FF8A605BE22B00CE805F" box="[1329,1429,831,854]" italics="true" pageId="1" pageNumber="152">Baeocera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C50818E2179FF8A6656E24B0104807C" authority="Lobl, 1983" authorityName="Lobl" authorityYear="1983" box="[828,1119,862,886]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="derougemonti">
|
||
<emphasis id="B924261F2179FF8A6656E24B06BC807C" box="[828,999,862,886]" italics="true" pageId="1" pageNumber="152">B. derougemonti</emphasis>
|
||
<bibRefCitation id="EFC187FC2179FF8A6681E24A0104807C" author="LOBL I." box="[1003,1119,862,885]" firstAuthor="Lobl" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="1" pageNumber="152" pagination="285 - 293" part="56" refId="ref12872" refString="LOBL I. 1983: Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 56: 285 - 293." title="Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien" type="journal article" year="1983">Löbl, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
, has been described from
|
||
<collectingRegion id="499434EF2179FF8A600FE24A0634809C" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="152">Sulawesi</collectingRegion>
|
||
. This species is a member of the
|
||
<emphasis id="B924261F2179FF8A61D1E26B0051809C" box="[1211,1290,894,918]" italics="true" pageId="1" pageNumber="152">B. lenta</emphasis>
|
||
species group defined by symmetrical aedeagi with a long ejaculatory duct extruded in resting position. The new species, a member of the
|
||
<emphasis id="B924261F2179FF8A6609E2CB06B680FF" box="[867,1005,991,1014]" italics="true" pageId="1" pageNumber="152">B. monstrosa</emphasis>
|
||
species group, defined by asymmetrical, voluminous aedeagi with complex internal sacs, with the ejaculatory duct not extruded in repos (see
|
||
<bibRefCitation id="EFC187FC2179FF8A6198E50A0038873C" author="LOBL I." box="[1266,1379,1054,1077]" firstAuthor="LOBL" journalOrPublisher="Revue Suisse de Zoologie" pageId="1" pageNumber="152" pagination="937 - 1006" part="78" refId="ref12828" refString="LOBL I. 1971: Scaphidiidae von Ceylon (Coleoptera). Revue Suisse de Zoologie 78: 937 - 1006." title="Scaphidiidae von Ceylon (Coleoptera)" type="journal article" year="1971">LÖBL 1971</bibRefCitation>
|
||
). The sole known Indonesian member of the
|
||
<emphasis id="B924261F2179FF8A61BAE52B0002875F" box="[1232,1369,1087,1110]" italics="true" pageId="1" pageNumber="152">B. monstrosa</emphasis>
|
||
group is the Sumatran
|
||
<emphasis id="B924261F2179FF8A6684E54B0127877C" box="[1006,1148,1118,1142]" italics="true" pageId="1" pageNumber="152">B. robertiana</emphasis>
|
||
Löbl, 1990, a species that differs drastically from
|
||
<taxonomicName id="4C50818E2179FF8A615AE56B01F8879F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1072,1187,1151,1174]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="inoptata" status="sp. nov.">
|
||
<emphasis id="B924261F2179FF8A615AE56B01F8879F" box="[1072,1187,1151,1174]" italics="true" pageId="1" pageNumber="152">B. inoptata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B642179FF8A61C3E56A01A1879C" box="[1193,1274,1150,1173]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the shape of the parameres: it has the right paramere strongly expanded and lamellar, abruptly narrowed and denticular near the tip. Also notable is the internal sac in
|
||
<emphasis id="B924261F2179FF8A61F9E5CB004787FC" box="[1171,1308,1246,1270]" italics="true" pageId="1" pageNumber="152">B. robertiana</emphasis>
|
||
lacking apical tufts of spine-like structures. Both species may be easily distinguished by their antennae: the antennomere IV in
|
||
<emphasis id="B924261F2179FF8A60EFE40B06F1865C" italics="true" pageId="1" pageNumber="152">B. robertiana</emphasis>
|
||
is about 1.5 times as long as antennomere III and similar to antennomere VI, while antennomeres III and IV are equally long in
|
||
<taxonomicName id="4C50818E2179FF8A6696E46B0136869F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1020,1133,1407,1430]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="inoptata" status="sp. nov.">
|
||
<emphasis id="B924261F2179FF8A6696E46B0136869F" box="[1020,1133,1407,1430]" italics="true" pageId="1" pageNumber="152">B. inoptata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B642179FF8A6118E46A019A869C" box="[1138,1217,1406,1429]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with III much shorter than antennomere VI. The median lobe and the parameres of the new species are comparatively similar with those of the Philippine
|
||
<emphasis id="B924261F2179FF8A66BBE4CB016086FC" box="[977,1083,1502,1526]" italics="true" pageId="1" pageNumber="152">B. alticola</emphasis>
|
||
<bibRefCitation id="EFC187FC2179FF8A612BE4CA01ED86FC" author="LOBL I." box="[1089,1206,1502,1525]" firstAuthor="Lobl" journalOrPublisher="Vernate" pageId="1" pageNumber="152" pagination="309 - 317" part="31" refId="ref12955" refString="LOBL I. 2012 a: Irianscapha dimorpha, an unusual new Scaphisomatini from Western New Guinea, and two new Indonesian species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae). Vernate 31: 309 - 317." title="Irianscapha dimorpha, an unusual new Scaphisomatini from Western New Guinea, and two new Indonesian species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae)" type="journal article" year="2012">Löbl, 2012</bibRefCitation>
|
||
. However, the internal sac bearing robust, elongate sclerites in the latter species is distinctive (see
|
||
<bibRefCitation id="EFC187FC2179FF8A66B0E70A010C853C" author="LOBL I." box="[986,1111,1566,1589]" firstAuthor="LOBL" journalOrPublisher="Revue Suisse de Zoologie" pageId="1" pageNumber="152" pagination="351 - 383" part="119" refId="ref12996" refString="LOBL I. 2012 b: On the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of the Philippines, III: the genus Baeocera Erichson. Revue Suisse de Zoologie 119: 351 - 383." title="On the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of the Philippines, III: the genus Baeocera Erichson" type="journal article" year="2012" yearSuffix="b">LÖBL 2012b</bibRefCitation>
|
||
). The “
|
||
<taxonomicName id="4C50818E2179FF8A61F5E70B0059853F" authorityName="Erichson" authorityYear="1845" box="[1183,1282,1567,1590]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924261F2179FF8A61F5E70B0059853F" box="[1183,1282,1567,1590]" italics="true" pageId="1" pageNumber="152">Baeocera</emphasis>
|
||
</taxonomicName>
|
||
sp1” in
|
||
<bibRefCitation id="EFC187FC2179FF8A603EE70A06DA855C" author="OGAWA R." bookContentInfo="180 pp." firstAuthor="OGAWA" journalOrPublisher="Graduate School of Agricultural Science, Kobe University" pageId="1" pageNumber="152" refId="ref13104" refString="OGAWA R. 2015: Phylogeography of the subfamily Scaphidiinae (Coleoptera, Staphylinidae) in Sulawesi, with its systematic revision. Graduate School of Agricultural Science, Kobe University, 180 pp. [PhD thesis, unpublished]." title="Phylogeography of the subfamily Scaphidiinae (Coleoptera, Staphylinidae) in Sulawesi, with its systematic revision" type="book" year="2015">OGAWA (2015)</bibRefCitation>
|
||
is likely
|
||
<taxonomicName id="4C50818E2179FF8A668BE72B010E855F" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[993,1109,1599,1622]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="inoptata" status="sp. nov.">
|
||
<emphasis id="B924261F2179FF8A668BE72B010E855F" box="[993,1109,1599,1622]" italics="true" pageId="1" pageNumber="152">B. inoptata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2179B642179FF8A6131E72A01F0855C" box="[1115,1195,1598,1621]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA9862179FF8A6656E74A01EF859C" pageId="1" pageNumber="152" type="etymology">
|
||
<paragraph id="8BEFFA0D2179FF8A6656E74A01EF859C" blockId="1.[828,1437,190,1717]" pageId="1" pageNumber="152">
|
||
<emphasis id="B924261F2179FF8A6656E74A06E0857C" bold="true" box="[828,955,1630,1653]" pageId="1" pageNumber="152">Etymology.</emphasis>
|
||
The species epithet is a Latin adjective
|
||
<emphasis id="B924261F2179FF8A603AE74B0600859F" italics="true" pageId="1" pageNumber="152">inoptatus</emphasis>
|
||
, -
|
||
<emphasis id="B924261F2179FF8A661BE76B0625859F" box="[881,894,1663,1686]" italics="true" pageId="1" pageNumber="152">a</emphasis>
|
||
, -
|
||
<emphasis id="B924261F2179FF8A66FFE76B06ED859F" box="[917,950,1663,1686]" italics="true" pageId="1" pageNumber="152">um</emphasis>
|
||
, meaning ‘not wanted’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34AA9862179FF8A6656E78A01AD85BC" box="[828,1270,1694,1717]" pageId="1" pageNumber="152" type="distribution">
|
||
<paragraph id="8BEFFA0D2179FF8A6656E78A01AD85BC" blockId="1.[828,1437,190,1717]" box="[828,1270,1694,1717]" pageId="1" pageNumber="152">
|
||
<emphasis id="B924261F2179FF8A6656E78A069585BC" bold="true" box="[828,974,1694,1717]" pageId="1" pageNumber="152">Distribution.</emphasis>
|
||
<collectingCountry id="F347BA9D2179FF8A66BEE78A016585BC" box="[980,1086,1694,1717]" name="Indonesia" pageId="1" pageNumber="152">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="499434EF2179FF8A6123E78A01A985BC" box="[1097,1266,1694,1717]" country="Indonesia" name="Sulawesi Selatan" pageId="1" pageNumber="152">South Sulawesi</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |