treatments-xml/data/03/EB/D7/03EBD72BFFF9FFCAF2C277B50B7E4839.xml
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<document id="EA37C72C6C221337E14E8348A5CB52DE" ID-DOI="10.5281/zenodo.10793369" ID-ISSN="1942-1354" ID-Zenodo-Dep="10793369" ID-ZooBank="7DD2EF0F-7DC9-4A85-83DA-10F4EE8CA8CB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709733663575" checkinUser="felipe" docAuthor="Santos-Silva, Antonio &amp; Botero, Juan Pablo" docDate="2024" docId="03EBD72BFFF9FFCAF2C277B50B7E4839" docLanguage="en" docName="InsectaMundi.2024.1028.1-17.pdf" docOrigin="Insecta Mundi 2024 (28)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Leptostylus cristulatus Bates 1872" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFD2AF53FFF2FFC7F2027117084D4E46" masterDocTitle="The unbelievable fauna of American Lamiinae (Coleoptera: Cerambycidae): description of two new species, transference, and notes in Acanthocinini, and a new record in Calliini" masterLastPageNumber="17" masterPageNumber="1" pageNumber="10" updateTime="1709812233359" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="D50D9585DE292119984DA953F17621E8">The unbelievable fauna of American Lamiinae (Coleoptera: Cerambycidae): description of two new species, transference, and notes in Acanthocinini, and a new record in Calliini</mods:title>
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<mods:namePart id="2765C5378288B951E5BEE6AB832DFC2B">Santos-Silva, Antonio</mods:namePart>
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<mods:namePart id="5B432F06E6AFCF75E03774996E0BA8A4">Botero, Juan Pablo</mods:namePart>
<mods:affiliation id="AFA73A0E2F3897708A508B728D6662B2">Sistemática Molecular, Laboratorio de Entomología, Pontificia Universidad Javeriana. Bogotá</mods:affiliation>
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<treatment id="03EBD72BFFF9FFCAF2C277B50B7E4839" LSID="urn:lsid:plazi:treatment:03EBD72BFFF9FFCAF2C277B50B7E4839" httpUri="http://treatment.plazi.org/id/03EBD72BFFF9FFCAF2C277B50B7E4839" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10">
<subSubSection id="C35835B6FFF9FFCCF2C277B50A3D48F9" box="[192,624,1698,1727]" pageId="11" pageNumber="10" type="nomenclature">
<paragraph id="8BFD663DFFF9FFCCF2C277B50A3D48F9" blockId="11.[192,624,1698,1761]" box="[192,624,1698,1727]" pageId="11" pageNumber="10">
<heading id="D0B5D151FFF9FFCCF2C277B50A3D48F9" box="[192,624,1698,1727]" fontSize="12" level="2" pageId="11" pageNumber="10" reason="2">
<taxonomicName id="4C421DBEFFF9FFCCF2C277B50A3D48F9" ID-CoL="3TWWF" authority="Bates, 1872" authorityName="Bates" authorityYear="1872" box="[192,624,1698,1727]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFF9FFCCF2C277B50A3D48F9" bold="true" box="[192,624,1698,1727]" pageId="11" pageNumber="10">
<emphasis id="B936BA2FFFF9FFCCF2C277B5099748F9" bold="true" box="[192,474,1698,1727]" italics="true" pageId="11" pageNumber="10">Leptostylus cristulatus</emphasis>
<bibRefCitation id="EFD31BCCFFF9FFCCF3E277B50A3D48F9" author="Bates HW" box="[480,624,1698,1727]" pageId="11" pageNumber="10" pagination="163 - 238" refId="ref9942" refString="Bates HW. 1872. On the longicorn Coleoptera of Chontales, Nicaragua. The Transactions of the Entomological Society of London 1872: 163 - 238." type="journal article" year="1872">Bates, 1872</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35835B6FFF9FFCCF2C277D1090548A7" box="[192,328,1734,1761]" pageId="11" pageNumber="10" type="description">
<paragraph id="8BFD663DFFF9FFCCF2C277D1090548A7" blockId="11.[192,624,1698,1761]" box="[192,328,1734,1761]" pageId="11" pageNumber="10">
(
<figureCitation id="13797AB8FFF9FFCCF2CA77D1090D48A7" box="[200,320,1734,1761]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="11" pageNumber="10">Fig. 2025</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35835B6FFF9FFCCF2E277E60A2E494F" box="[224,611,1777,1801]" pageId="11" pageNumber="10" type="reference_group">
<paragraph id="8BFD663DFFF9FFCCF2E277E60A2E494F" blockId="11.[224,611,1777,1801]" box="[224,611,1777,1801]" pageId="11" pageNumber="10">
<treatmentCitationGroup id="AB524113FFF9FFCCF2E277E60A2E494F" box="[224,611,1777,1801]" pageId="11" pageNumber="10">
<treatmentCitation id="0AE3402CFFF9FFCCF2E277E60A12494F" author="Bates HW" box="[224,607,1777,1801]" page="213" pageId="11" pageNumber="10" year="1872">
<taxonomicName id="4C421DBEFFF9FFCCF2E277E60A12494F" ID-CoL="3TWWF" authority="Bates 1872: 213" authorityName="Bates" authorityPageNumber="213" authorityYear="1872" box="[224,607,1777,1801]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFF9FFCCF2E277E609F6494F" box="[224,443,1777,1801]" italics="true" pageId="11" pageNumber="10">Leptostylus cristulatus</emphasis>
<bibRefCitation id="EFD31BCCFFF9FFCCF3C377E60A12494F" author="Bates HW" box="[449,607,1777,1801]" pageId="11" pageNumber="10" pagination="163 - 238" refId="ref9942" refString="Bates HW. 1872. On the longicorn Coleoptera of Chontales, Nicaragua. The Transactions of the Entomological Society of London 1872: 163 - 238." type="journal article" year="1872">Bates 1872: 213</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35835B6FFF9FFCAF2C2760E0BCF4875" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10" type="discussion">
<paragraph id="8BFD663DFFF9FFCAF2C2760E0AB44CD5" blockId="11.[192,1441,1816,1910]" lastBlockId="13.[191,1442,230,895]" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10">
<emphasis id="B936BA2FFFF9FFCCF2C2760E09624975" bold="true" box="[192,303,1817,1843]" pageId="11" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="4C421DBEFFF9FFCCF337760F0A654974" authorityName="Bates" authorityYear="1872" box="[309,552,1816,1842]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFF9FFCCF337760F0A654974" box="[309,552,1816,1842]" italics="true" pageId="11" pageNumber="10">Leptostylus cristulatus</emphasis>
</taxonomicName>
was described based on a single specimen from
<collectingCountry id="F35526ADFFF9FFCCF649760F0C8C4974" box="[1099,1217,1816,1842]" name="Nicaragua" pageId="11" pageNumber="10">Nicaragua</collectingCountry>
. According to
<bibRefCitation id="EFD31BCCFFF9FFCCF767760E09474912" author="Bates HW" pageId="11" pageNumber="10" pagination="163 - 238" refId="ref9942" refString="Bates HW. 1872. On the longicorn Coleoptera of Chontales, Nicaragua. The Transactions of the Entomological Society of London 1872: 163 - 238." type="journal article" year="1872">Bates (1872)</bibRefCitation>
(translated): “Oval, brown, with a grayish-white discoidal macula on each side behind the middle of the elytra; thorax with two large tubercles near the anterior margin, and two other tubercles on each side [prothoracic tubercles]; elytra acutely rounded apically, with centrobasal crest bearing four tubercles, and with three rows of tubercles on the disc; antennae with segments 49 yellowish-testaceous basally.” However, the photograph of the
<typeStatus id="54F9D89FFFFFFFCAF2C2703E096F4F05" box="[192,290,297,323]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
(
<figureCitation id="13797AB8FFFFFFCAF333703E09CC4F02" box="[305,385,297,324]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 25</figureCitation>
) shows that there is an elongated whitish pubescent macula close to the suture on the anterior third, and two small, narrow, and elongated whitish pubescent spots on the dorsal surface near the superior region of the wide and oblique whitish pubescent band on the posterior half. Furthermore, the base of antennomere IV is as on III, and it is antennomeres VIX that have a whitish pubescent ring anteriorly; in fact, the pubescent ring does not start on the base in some of these antennomeres.
<bibRefCitation id="EFD31BCCFFFFFFCAF14770A70B824F8C" author="Bates HW" box="[837,975,431,458]" pageId="13" pageNumber="12" pagination="249 - 436" refId="ref9970" refString="Bates HW. 1885. Supplement. p. 249 - 436. In: Godman FD, Salvin O (eds.). Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis; London; xii + 525 p." type="book chapter" year="1885">Bates (1885)</bibRefCitation>
recorded
<taxonomicName id="4C421DBEFFFFFFCAF63D70A70C814F8F" authorityName="Bates" authorityYear="1872" box="[1087,1228,431,457]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFFFFCAF63D70A70C814F8F" box="[1087,1228,431,457]" italics="true" pageId="13" pageNumber="12">L. cristulatus</emphasis>
</taxonomicName>
from
<collectingCountry id="F35526ADFFFFFFCAF71270A70D214F8C" box="[1296,1388,432,458]" name="Panama" pageId="13" pageNumber="12">Panama</collectingCountry>
, and we do not know how many specimens from this country Bates examined. However, we believe that the specimen illustrated in
<bibRefCitation id="EFD31BCCFFFFFFCAF35070E509B74C4B" author="Bezark LG" box="[338,506,498,525]" pageId="13" pageNumber="12" refId="ref10017" refString="Bezark LG. 2023 a. A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available at http: // bezbycids. com / byciddb / wdefault. asp? w = n / (Last accessed 30 June 2023.)" type="url" year="2023">Bezark (2023a)</bibRefCitation>
(
<figureCitation id="13797AB8FFFFFFCAF00570E50A184C4B" box="[519,597,498,525]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 24</figureCitation>
) is, at least, one of these specimens. This specimen does not have the whitish pubescent macula on the anterior third of the elytra and the two small whitish pubescent spots were replaced by a wide and transverse whitish pubescent band that reaches the suture, which was not mentioned by
<bibRefCitation id="EFD31BCCFFFFFFCAF71273210DD14C17" author="Bates HW" box="[1296,1436,566,593]" pageId="13" pageNumber="12" pagination="249 - 436" refId="ref9970" refString="Bates HW. 1885. Supplement. p. 249 - 436. In: Godman FD, Salvin O (eds.). Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis; London; xii + 525 p." type="book chapter" year="1885">Bates (1885)</bibRefCitation>
. Furthermore, the dark and arched area starting on apex of the posterior whitish pubescent band is more distinct, forming a semicircle when considering both elytra.
</paragraph>
<caption id="DF3D36B5FFFEFFCBF2C2763E0A4C49C0" ID-DOI="http://doi.org/10.5281/zenodo.10793380" ID-Zenodo-Dep="10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="12" pageNumber="11" startId="12.[192,279,1833,1859]" targetBox="[192,1440,229,1802]" targetPageId="12" targetType="figure">
<paragraph id="8BFD663DFFFEFFCBF2C2763E0A4C49C0" blockId="12.[192,1440,1832,1927]" pageId="12" pageNumber="11">
<emphasis id="B936BA2FFFFEFFCBF2C2763E09214905" bold="true" box="[192,364,1833,1859]" pageId="12" pageNumber="11">Figures 2025.</emphasis>
<taxonomicName id="4C421DBEFFFEFFCBF374763E0ABE4905" authority="Bates, 1872" authorityName="Bates" authorityYear="1872" box="[374,755,1833,1859]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFEFFCBF374763E0A274905" box="[374,618,1833,1859]" italics="true" pageId="12" pageNumber="11">Leptostylus cristulatus</emphasis>
<bibRefCitation id="EFD31BCCFFFEFFCBF072763E0ABE4905" author="Bates HW" box="[624,755,1833,1859]" pageId="12" pageNumber="11" pagination="163 - 238" refId="ref9942" refString="Bates HW. 1872. On the longicorn Coleoptera of Chontales, Nicaragua. The Transactions of the Entomological Society of London 1872: 163 - 238." type="journal article" year="1872">Bates, 1872</bibRefCitation>
</taxonomicName>
.
<emphasis id="B936BA2FFFFEFFCBF0FC763F0B034904" bold="true" box="[766,846,1832,1858]" pageId="12" pageNumber="11">2023)</emphasis>
Female from Belize;
<emphasis id="B936BA2FFFFEFFCBF643763E0C164905" bold="true" box="[1089,1115,1833,1859]" pageId="12" pageNumber="11">20</emphasis>
) Dorsal habitus;
<emphasis id="B936BA2FFFFEFFCBF720763F0D0A4904" bold="true" box="[1314,1351,1832,1858]" pageId="12" pageNumber="11">21)</emphasis>
Ventral habitus;
<emphasis id="B936BA2FFFFEFFCBF31D765D09094922" bold="true" box="[287,324,1866,1892]" pageId="12" pageNumber="11">22)</emphasis>
Lateral habitus;
<emphasis id="B936BA2FFFFEFFCBF3FD765D0A684922" bold="true" box="[511,549,1866,1892]" pageId="12" pageNumber="11">23)</emphasis>
Head, frontal view.
<emphasis id="B936BA2FFFFEFFCBF105765D0B614922" bold="true" box="[775,812,1866,1892]" pageId="12" pageNumber="11">24)</emphasis>
Holotype, by Jesus Santiago Moure.
<emphasis id="B936BA2FFFFEFFCBF6CA765D0CA34922" bold="true" box="[1224,1262,1866,1892]" pageId="12" pageNumber="11">25)</emphasis>
Specimen from Panama, by Larry G. Bezark.
</paragraph>
</caption>
<paragraph id="8BFD663DFFFFFFCAF302738D0CFB4D7D" blockId="13.[191,1442,230,895]" pageId="13" pageNumber="12">
Although we examined a single specimen of
<taxonomicName id="4C421DBEFFFFFFCAF0F2738C0B304CF2" authorityName="Bates" authorityYear="1872" box="[752,893,666,692]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFFFFCAF0F2738C0B304CF2" box="[752,893,666,692]" italics="true" pageId="13" pageNumber="12">L. cristulatus</emphasis>
</taxonomicName>
, we did examine photographs of specimens from
<collectingCountry id="F35526ADFFFFFFCAF2C273AB095E4C90" box="[192,275,700,726]" name="Mexico" pageId="13" pageNumber="12">Mexico</collectingCountry>
and
<collectingCountry id="F35526ADFFFFFFCAF34C73AB09F34C90" box="[334,446,700,726]" name="Honduras" pageId="13" pageNumber="12">Honduras</collectingCountry>
identified as
<taxonomicName id="4C421DBEFFFFFFCAF05873AA0AA64C90" authorityName="Bates" authorityYear="1872" box="[602,747,700,726]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFFFFCAF05873AA0AA64C90" box="[602,747,700,726]" italics="true" pageId="13" pageNumber="12">L. cristulatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFD31BCCFFFFFFCAF12A73AB0B8F4C90" author="Bezark LG" box="[808,962,700,726]" pageId="13" pageNumber="12" refId="ref10017" refString="Bezark LG. 2023 a. A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available at http: // bezbycids. com / byciddb / wdefault. asp? w = n / (Last accessed 30 June 2023.)" type="url" year="2023">Bezark 2023a</bibRefCitation>
) as well as photographs of
<specimenCount id="9D44ADB4FFFFFFCAF6F973AB0DED4C90" box="[1275,1440,700,726]" count="5" pageId="13" pageNumber="12" type="generic">five specimens</specimenCount>
from
<collectingCountry id="F35526ADFFFFFFCAF30273C909344CBE" box="[256,377,734,760]" name="Costa Rica" pageId="13" pageNumber="12">Costa Rica</collectingCountry>
(as we are not authorized to illustrate these specimens from
<collectingCountry id="F35526ADFFFFFFCAF63373C90CE64CBE" box="[1073,1195,734,760]" name="Costa Rica" pageId="13" pageNumber="12">Costa Rica</collectingCountry>
and since they are of low resolution, we prefer not to indicate the website). However, these specimens from
<collectingCountry id="F35526ADFFFFFFCAF68773E80CB34D5F" box="[1157,1278,767,793]" name="Costa Rica" pageId="13" pageNumber="12">Costa Rica</collectingCountry>
are practically identical to the
<typeStatus id="54F9D89FFFFFFFCAF36C7236099D4D7D" box="[366,464,801,827]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
in shape and general appearance of the elytral pubescent maculae.
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF302725509AB4D39" blockId="13.[191,1442,230,895]" pageId="13" pageNumber="12">
Based on the specimen examined and photographs (including the
<typeStatus id="54F9D89FFFFFFFCAF1ED72550C184D1A" box="[1007,1109,834,860]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
), we can report the following variations in
<taxonomicName id="4C421DBEFFFFFFCAF350727209AD4D38" authorityName="Bates" authorityYear="1872" box="[338,480,868,894]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFFFFCAF350727209AD4D38" box="[338,480,868,894]" italics="true" pageId="13" pageNumber="12">L. cristulatus</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C272860BE74DA8" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">
1. Antennomeres VIX with wide and dense yellowish-white pubescent ring (
<figureCitation id="13797AB8FFFFFFCAF65A72870CE44DED" box="[1112,1193,912,939]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 25</figureCitation>
) or absent on VI and VIII (
<figureCitation id="13797AB8FFFFFFCAF34272A509DD4D8A" box="[320,400,946,972]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 20</figureCitation>
). The absence of the yellowish-white pubescent ring on the antennomeres VI and VIII was also observed through photograph in a specimen from
<collectingCountry id="F35526ADFFFFFFCAF13672C30BEB4DA8" box="[820,934,980,1006]" name="Honduras" pageId="13" pageNumber="12">Honduras</collectingCountry>
;
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C272E20C9E4A49" blockId="13.[192,1442,912,1476]" box="[192,1235,1013,1040]" pageId="13" pageNumber="12">2. Anterolateral tubercles on the pronotum from subconical to distinctly rounded apically;</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C275000D504AD0" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">
<materialsCitation id="3B2A6C60FFFFFFCAF2C275000D544AD0" country="Belize" location="Elytral" pageId="13" pageNumber="12" specimenCount="3" typeStatus="holotype">
3.
<location id="8E9D30E6FFFFFFCAF302750009074A77" LSID="urn:lsid:plazi:treatment:03EBD72BFFF9FFCAF2C277B50B7E4839:8E9D30E6FFFFFFCAF302750009074A77" box="[256,330,1047,1073]" country="Belize" name="Elytral" pageId="13" pageNumber="12">Elytral</location>
length from 1.6 (specimen from
<collectingCountry id="F35526ADFFFFFFCAF0CB75000B434A77" box="[713,782,1047,1073]" name="Belize" pageId="13" pageNumber="12">Belize</collectingCountry>
) to 1.7 (
<typeStatus id="54F9D89FFFFFFFCAF16D75000B984A77" box="[879,981,1047,1073]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
, measured through the photograph) the humeral width. Although the specimen from
<collectingCountry id="F35526ADFFFFFFCAF108752F0B004A14" box="[778,845,1080,1106]" name="Belize" pageId="13" pageNumber="12">Belize</collectingCountry>
is the widest specimen the other features agree very well with the
<typeStatus id="54F9D89FFFFFFFCAF39C754D0A174A32" box="[414,602,1114,1140]" isOtherTypeStatus="true" pageId="13" pageNumber="12">other specimens</typeStatus>
. Furthermore, the measurement of some
<typeStatus id="54F9D89FFFFFFFCAF630754D0CA64A32" box="[1074,1259,1114,1140]" isOtherTypeStatus="true" pageId="13" pageNumber="12">other specimens</typeStatus>
through photographs showed intermediate widths (1.75). Therefore, we consider as an intraspecific variation
</materialsCitation>
;
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C2758A0D1A4AF1" blockId="13.[192,1442,912,1476]" box="[192,1367,1181,1208]" pageId="13" pageNumber="12">
4. Whitish pubescent macula on the anterior third of the elytra present (
<figureCitation id="13797AB8FFFFFFCAF605758A0C1D4AFE" box="[1031,1104,1181,1208]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig 25</figureCitation>
) or absent (
<figureCitation id="13797AB8FFFFFFCAF6D1758A0D064AF1" box="[1235,1355,1181,1207]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 20, 24</figureCitation>
);
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C275A80B984ABD" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">
5. Elytra with whitish pubescence before the oblique whitish pubescent band as two small isolated spots (
<figureCitation id="13797AB8FFFFFFCAF77675A809504ABD" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 25</figureCitation>
) or as a wide transverse band reaching the suture (
<figureCitation id="13797AB8FFFFFFCAF15375F70B854ABD" box="[849,968,1248,1275]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 20, 24</figureCitation>
);
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C274150DC44B78" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">
6. Dark band close to the inferior margin of the first whitish pubescent band or spots on the posterior half of the elytra small and elongate (
<figureCitation id="13797AB8FFFFFFCAF04F74330AD04B78" box="[589,669,1315,1342]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 25</figureCitation>
), absent (
<figureCitation id="13797AB8FFFFFFCAF10574330B1A4B78" box="[775,855,1316,1342]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 24</figureCitation>
), or large and almost reaching the suture (
<figureCitation id="13797AB8FFFFFFCAF72E74330D314B78" box="[1324,1404,1316,1342]" captionStart="Figures 2025" captionStartId="12.[192,279,1833,1859]" captionTargetBox="[192,1440,229,1802]" captionTargetId="figure-53@12.[192,1440,229,1802]" captionTargetPageId="12" captionText="Figures 2025. Leptostylus cristulatus Bates, 1872. 2023) Female from Belize; 20) Dorsal habitus; 21) Ventral habitus; 22) Lateral habitus; 23) Head, frontal view. 24) Holotype, by Jesus Santiago Moure. 25) Specimen from Panama, by Larry G. Bezark." figureDoi="http://doi.org/10.5281/zenodo.10793380" httpUri="https://zenodo.org/record/10793380/files/figure.png" pageId="13" pageNumber="12">Fig. 20</figureCitation>
);
</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C2745109114BC7" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">7. Dark pubescent band close to the inner apex of the oblique whitish pubescent band from absent to very distinct;</paragraph>
<paragraph id="8BFD663DFFFFFFCAF2C2749E09134B82" blockId="13.[192,1442,912,1476]" pageId="13" pageNumber="12">8. Centrobasal crest on the elytra from slightly elevated to distinctly elevated, with three or four tubercles dorsally.</paragraph>
<paragraph id="8BFD663DFFFFFFCAF30274C10BCF4875" blockId="13.[192,1442,1494,1588]" pageId="13" pageNumber="12">
Currently,
<taxonomicName id="4C421DBEFFFFFFCAF37874C00A474BB6" authorityName="Bates" authorityYear="1872" box="[378,522,1494,1520]" class="Insecta" family="Cerambycidae" genus="Leptostylus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cristulatus">
<emphasis id="B936BA2FFFFFFFCAF37874C00A474BB6" box="[378,522,1494,1520]" italics="true" pageId="13" pageNumber="12">L. cristulatus</emphasis>
</taxonomicName>
is formally known from
<collectingCountry id="F35526ADFFFFFFCAF12F74C10BD24BB6" box="[813,927,1494,1520]" name="Honduras" pageId="13" pageNumber="12">Honduras</collectingCountry>
,
<collectingCountry id="F35526ADFFFFFFCAF1AE74C10C6C4BB6" box="[940,1057,1494,1520]" name="Nicaragua" pageId="13" pageNumber="12">Nicaragua</collectingCountry>
,
<collectingCountry id="F35526ADFFFFFFCAF62F74C10CE74BB6" box="[1069,1194,1494,1520]" name="Costa Rica" pageId="13" pageNumber="12">Costa Rica</collectingCountry>
, and
<collectingCountry id="F35526ADFFFFFFCAF6EB74C00D0E4BB7" box="[1257,1347,1495,1521]" name="Panama" pageId="13" pageNumber="12">Panama</collectingCountry>
(
<bibRefCitation id="EFD31BCCFFFFFFCAF75174C109444854" author="Bezark LG" pageId="13" pageNumber="12" refId="ref10065" refString="Bezark LG. 2023 b. Checklist of the Oxypeltidae, Vesperidae, Disteniidae and Cerambycidae (Coleoptera) of the Western Hemisphere. 2023 Edition (updated through 31 December 2022). Available athttp: // bezbycids. com / byciddb / wdefault. asp? w = n (Last accessed 30 June 2023.)" type="book" year="2023">Bezark 2023b</bibRefCitation>
;
<bibRefCitation id="EFD31BCCFFFFFFCAF31174EF09EF4854" author="Monne MA" box="[275,418,1528,1554]" pageId="13" pageNumber="12" refId="ref10246" refString="Monne MA. 2023. Catalogue of the Cerambycidae (Coleoptera) of the Neotropical region. Part II. Subfamily Lamiinae. Available at https: // cerambycids. com / catalog / (Last accessed 30 June 2023.)" type="url" year="2023">Monné 2023</bibRefCitation>
;
<bibRefCitation id="EFD31BCCFFFFFFCAF3AF74EF0B584854" author="Tavakilian GL &amp; Chevillotte H." box="[429,789,1528,1554]" pageId="13" pageNumber="12" refId="ref10390" refString="Tavakilian GL, Chevillotte H. 2023. Titan: base de donnees internationales sur les Cerambycidae ou Longicornes. Available at http: // titan. gbif. fr / (Last accessed 7 August 2023.)" type="url" year="2023">Tavakilian and Chevillotte 2023</bibRefCitation>
). Although we have examined a photograph of a specimen from
<collectingCountry id="F35526ADFFFFFFCAF2FC770E091C4875" box="[254,337,1561,1587]" name="Mexico" pageId="13" pageNumber="12">Mexico</collectingCountry>
(
<bibRefCitation id="EFD31BCCFFFFFFCAF35D770E09B44872" author="Bezark LG" box="[351,505,1561,1588]" pageId="13" pageNumber="12" refId="ref10017" refString="Bezark LG. 2023 a. A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available at http: // bezbycids. com / byciddb / wdefault. asp? w = n / (Last accessed 30 June 2023.)" type="url" year="2023">Bezark 2023a</bibRefCitation>
) we are not formalizing the record.
</paragraph>
</subSubSection>
<subSubSection id="C35835B6FFFFFFCAF2C277540B7E4839" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="8BFD663DFFFFFFCAF2C277540B7E4839" blockId="13.[192,1440,1603,1663]" pageId="13" pageNumber="12">
<emphasis id="B936BA2FFFFFFFCAF2C2775409EE481B" bold="true" box="[192,419,1603,1629]" pageId="13" pageNumber="12">Material examined.</emphasis>
<collectingCountry id="F35526ADFFFFFFCAF3A977530A4A4818" box="[427,519,1604,1630]" name="Belize" pageId="13" pageNumber="12">BELIZE</collectingCountry>
,
<emphasis id="B936BA2FFFFFFFCAF01377540A14481B" box="[529,601,1603,1629]" italics="true" pageId="13" pageNumber="12">
<collectingRegion id="4986A8DFFFFFFFCAF01377540A14481B" box="[529,601,1603,1629]" country="Belize" name="Toledo" pageId="13" pageNumber="12">Toledo</collectingRegion>
</emphasis>
: BARC San Pedro Columbia,
<geoCoordinate id="EE7600FAFFFFFFCAF1A877540C664818" box="[938,1067,1603,1631]" degrees="15" direction="north" minutes="16" orientation="latitude" pageId="13" pageNumber="12" precision="15" seconds="43" value="15.278611">15°1643″N</geoCoordinate>
<geoCoordinate id="EE7600FAFFFFFFCAF63077540CF54818" box="[1074,1208,1603,1631]" degrees="88" direction="west" minutes="57" orientation="longitude" pageId="13" pageNumber="12" precision="15" seconds="49" value="-88.96361">88°5749″W</geoCoordinate>
, on dead tree trunks at night,
<specimenCount id="9D44ADB4FFFFFFCAF321777209CE4839" box="[291,387,1637,1663]" count="1" pageId="13" pageNumber="12" type="female">1 female</specimenCount>
,
<date id="FFFC40FDFFFFFFCAF38C77720A474839" box="[398,522,1637,1663]" pageId="13" pageNumber="12" value="2004-09-23">23.IX.2004</date>
, P.W. Kovarik leg. (
<collectionCode id="ED53FEF8FFFFFFCAF0DC77720B6A4839" box="[734,807,1637,1663]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="13" pageNumber="12" type="Museum">FSCA</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>