653 lines
76 KiB
XML
653 lines
76 KiB
XML
<document id="586847170B53326D16C20EDA6214A31E" ID-CLB-Dataset="42256" ID-DOI="10.11646/zootaxa.3881.6.2" ID-GBIF-Dataset="9487ed7a-9071-4418-942f-9d7cac01c9cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="230287" ID-ZooBank="9E627401-690E-4F2D-9334-5447248C940C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460960291231" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V. & Mal, Ahmad O." docDate="2014" docId="03EBC7135E2AFFDA7DB0FBECFEB9EA5F" docLanguage="en" docName="zt03881p531.pdf" docOrigin="Zootaxa 3881 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coryogalops pseudomonospilus Kovacic & Bogorodsky, sp. nov." docType="treatment" docUuid="868FD3F7-5661-421B-A75A-5B25629A483E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="527" masterDocId="FFD2BF6B5E23FFD57D27FFD3FFD9E863" masterDocTitle="Two new species of Coryogalops (Perciformes: Gobiidae) from the Red Sea" masterLastPageNumber="531" masterPageNumber="513" pageNumber="522" updateTime="1698586895071" updateUser="ExternalLinkService">
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<mods:mods id="2A966227BD01607A247ED769FE895234" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="90039788A5752C4359A43D52317252DA">
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<mods:title id="DB8C6B40391724CDB6C26A5C90C296D2">Two new species of Coryogalops (Perciformes: Gobiidae) from the Red Sea</mods:title>
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</mods:titleInfo>
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<mods:name id="6AD6871908E5F9FC6DB729DF98BA535E" type="personal">
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<mods:role id="24ACE33C1CF885F1CCD93D069A80733E">
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<mods:roleTerm id="11032D0177D31EBBA293788CB7A44BF2">Author</mods:roleTerm>
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<mods:namePart id="C8164E0696F3444D03894E304C8CB74A">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="078BEC387815DCC87F8EFC93ED14EFB1">Bogorodsky, Sergey V.</mods:namePart>
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<mods:roleTerm id="26DD2F2AA56DA86CCD2C41E22D1712D5">Author</mods:roleTerm>
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<mods:namePart id="4CA413FF85610066AF4BB84EA118B22C">Mal, Ahmad O.</mods:namePart>
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<mods:typeOfResource id="8D97CBA733F0BB520AD84E875E49277D">text</mods:typeOfResource>
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<mods:title id="3C17D5E021724690CD7768FA4779A3DD">Zootaxa</mods:title>
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<mods:part id="4C9116C3992C2664D39CF0EA6DA9F782">
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<mods:date id="41A698266CB474C23D086FD5F5A8640E">2014</mods:date>
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<mods:detail id="9E2E858F9D9373072EC45B47F50E43D5" type="volume">
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<mods:number id="8045A0281D51DB202F53CA81759A2D2C">3881</mods:number>
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<mods:detail id="FA4D15B96B70D7C0589328DCE2AB5B86" type="issue">
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<mods:number id="A83D470EBB04B6A0F09D714E415C72E0">6</mods:number>
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<mods:classification id="0B9E7C06C621CF3407A0D26D54E5C824">journal article</mods:classification>
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<mods:identifier id="8D996F31A019E73DEC7DB169683B51AA" type="DOI">10.11646/zootaxa.3881.6.2</mods:identifier>
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</mods:mods>
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<treatment id="03EBC7135E2AFFDA7DB0FBECFEB9EA5F" ID-DOI="http://doi.org/10.5281/zenodo.5691335" ID-GBIF-Taxon="119593069" ID-Zenodo-Dep="5691335" LSID="urn:lsid:zoobank.org:act:868FD3F7-5661-421B-A75A-5B25629A483E" httpUri="http://treatment.plazi.org/id/03EBC7135E2AFFDA7DB0FBECFEB9EA5F" lastPageId="15" lastPageNumber="527" pageId="9" pageNumber="522">
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<subSubSection id="C358258E5E2AFFDC7DB0FBECFE6FEC18" pageId="9" pageNumber="522" type="nomenclature">
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<paragraph id="8BFD76055E2AFFDC7DB0FBECFC71EC39" blockId="9.[151,936,1087,1147]" box="[151,936,1087,1114]" pageId="9" pageNumber="522">
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<heading id="D0B5C1695E2AFFDC7DB0FBECFC71EC39" bold="true" box="[151,936,1087,1114]" fontSize="11" level="1" pageId="9" pageNumber="522" reason="1">
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<emphasis id="B936AA175E2AFFDC7DB0FBECFC71EC39" bold="true" box="[151,936,1087,1114]" pageId="9" pageNumber="522">
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<taxonomicName id="4C420D865E2AFFDC7DB0FBECFCE7EC39" ID-CoL="YRWZ" authority="Kovacic & Bogorodsky" authorityName="Kovacic & Bogorodsky" box="[151,830,1087,1114]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="522" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
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<emphasis id="B936AA175E2AFFDC7DB0FBECFDCCEC3A" bold="true" box="[151,533,1087,1113]" italics="true" pageId="9" pageNumber="522">Coryogalops pseudomonospilus</emphasis>
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Kovačić & Bogorodsky
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</taxonomicName>
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,
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<taxonomicNameLabel id="A205176C5E2AFFDC7E6DFB93FC71EC39" box="[842,936,1088,1114]" pageId="9" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFD76055E2AFFDC7DB0FBB1FE6FEC18" blockId="9.[151,936,1087,1147]" box="[151,438,1122,1147]" pageId="9" pageNumber="522">
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(
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<figureCitation id="13796A805E2AFFDC7DB9FBB1FED3EC18" box="[158,266,1122,1147]" captionStart="FIGURE 3" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[179,1407,193,1897]" captionTargetId="figure@10.[179,1408,193,1899]" captionTargetPageId="10" captionText="FIGURE 3. Coryogalops pseudomonospilus sp. nov. A: SMF 35231, holotype, male, 21.5 + 5.5 mm, Al Khuraybah, Saudi Arabia, Red Sea, live specimen; B: the same, preserved specimen. C: KAUMM 260, paratype, female, 19.3 + 4.8 mm, Al Khuraybah, Saudi Arabia, Red Sea, preserved specimen. Photos by S. V. Bogorodsky (A), M. Kovačić (B and C)." httpUri="https://zenodo.org/record/230290/files/figure.png" pageId="9" pageNumber="522">Figures 3</figureCitation>
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,
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<figureCitation id="13796A805E2AFFDC7C30FBB0FEF1EC18" box="[279,296,1123,1147]" captionStart="FIGURE 4" captionStartId="11.[151,250,1353,1375]" captionTargetBox="[289,1436,538,1256]" captionTargetId="figure@11.[151,1436,439,1331]" captionTargetPageId="11" captionText="FIGURE 4. Cephalic sensory systems of Coryogalops pseudomonospilus sp. nov., KAUMM 260, paratype. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/230291/files/figure.png" pageId="9" pageNumber="522">4</figureCitation>
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; Tables 1, 2)
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</paragraph>
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</subSubSection>
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<subSubSection id="C358258E5E2AFFDC7DB0FB79FBEFED4C" pageId="9" pageNumber="522" type="materials_examined">
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<paragraph id="8BFD76055E2AFFDC7DB0FB79FBEFEC84" blockId="9.[151,1437,1194,2011]" pageId="9" pageNumber="522">
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<typeStatus id="54F9C8A75E2AFFDC7DB0FB79FEDDECA0" box="[151,260,1194,1219]" pageId="9" pageNumber="522" type="holotype">
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<emphasis id="B936AA175E2AFFDC7DB0FB79FEDDECA0" bold="true" box="[151,260,1194,1219]" pageId="9" pageNumber="522">Holotype</emphasis>
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</typeStatus>
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.
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<collectionCode id="ED53EEC05E2AFFDC7C3AFB78FE8CECA0" LSID="urn:lsid:biocol.org:col:34838" box="[285,341,1195,1219]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="522">SMF</collectionCode>
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35231, male, 21.5 +
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<quantity id="4CBADBE05E2AFFDC7F5DFB79FD04ECA1" box="[634,733,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="522" unit="mm" value="5.5">5.5 mm</quantity>
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, Red Sea,
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<collectingCountry id="F35536955E2AFFDC7E52FB79FBCFECA0" box="[885,1046,1194,1219]" name="Saudi Arabia" pageId="9" pageNumber="522">Saudi Arabia</collectingCountry>
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, Al Khuraybah,
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<geoCoordinate id="EE7610C25E2AFFDC79C8FB78FA41ECA0" box="[1263,1432,1195,1219]" direction="north" orientation="latitude" pageId="9" pageNumber="522" precision="1" value="28.069918">N28°04'11.70"</geoCoordinate>
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,
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<geoCoordinate id="EE7610C25E2AFFDC7DB0FB1DFEE4EC84" box="[151,317,1230,1255]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="1" value="35.0802">E35°04'48.72"</geoCoordinate>
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||
, coral patch in lagoon,
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||
<quantity id="4CBADBE05E2AFFDC7F65FB1CFD52EC85" box="[578,651,1231,1255]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="9" pageNumber="522" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">7–8 m</quantity>
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, coll. S.V. Bogorodsky,
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||
<date id="FFFC50C55E2AFFDC7EBEFB1CFBEAEC84" box="[921,1075,1231,1255]" pageId="9" pageNumber="522" value="2013-06-24">24 June 2013</date>
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.
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||
</paragraph>
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<paragraph id="8BFD76055E2AFFDC7DE0FB21FBEFED4C" blockId="9.[151,1437,1194,2011]" pageId="9" pageNumber="522">
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<typeStatus id="54F9C8A75E2AFFDC7DE0FB21FEEDED68" box="[199,308,1266,1291]" pageId="9" pageNumber="522" type="paratype">
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||
<emphasis id="B936AA175E2AFFDC7DE0FB21FEEDED68" bold="true" box="[199,308,1266,1291]" pageId="9" pageNumber="522">Paratype</emphasis>
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||
</typeStatus>
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||
.
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||
<collectionCode id="ED53EEC05E2AFFDC7C61FB20FE6DED68" box="[326,436,1267,1291]" pageId="9" pageNumber="522">KAUMM</collectionCode>
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||
260, female, 19.3 +
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||
<quantity id="4CBADBE05E2AFFDC7F96FB20FCD4ED69" box="[689,781,1267,1291]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="522" unit="mm" value="4.8">4.8 mm</quantity>
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, Red Sea,
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||
<collectingCountry id="F35536955E2AFFDC7EB7FB21FBF3ED68" box="[912,1066,1266,1291]" name="Saudi Arabia" pageId="9" pageNumber="522">Saudi Arabia</collectingCountry>
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||
, Al Khuraybah,
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||
<geoCoordinate id="EE7610C25E2AFFDC79C8FB20FA41ED68" box="[1263,1432,1267,1291]" direction="north" orientation="latitude" pageId="9" pageNumber="522" precision="1" value="28.069918">N28°04'11.70"</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE7610C25E2AFFDC7DB0FAC5FEE4ED4C" box="[151,317,1302,1327]" direction="east" orientation="longitude" pageId="9" pageNumber="522" precision="1" value="35.0802">E35°04'48.72"</geoCoordinate>
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||
, coral patch in lagoon,
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||
<quantity id="4CBADBE05E2AFFDC7F65FAC4FD52ED4D" box="[578,651,1303,1327]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="9" pageNumber="522" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">7–8 m</quantity>
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, coll. S.V. Bogorodsky,
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||
<date id="FFFC50C55E2AFFDC7EBEFAC4FBEAED4C" box="[921,1075,1303,1327]" pageId="9" pageNumber="522" value="2013-06-24">24 June 2013</date>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C358258E5E2AFFDC7DE0FAE9FE23EF60" pageId="9" pageNumber="522" type="diagnosis">
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<paragraph id="8BFD76055E2AFFDC7DE0FAE9FE23EF60" blockId="9.[151,1437,1194,2011]" pageId="9" pageNumber="522">
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<emphasis id="B936AA175E2AFFDC7DE0FAE9FEE3ED30" bold="true" box="[199,314,1338,1363]" pageId="9" pageNumber="522">Diagnosis</emphasis>
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. Dorsal-fin rays VI + I,11; second to sixth spines of first dorsal fin reaching spine of second fin in male, and at least fourth spine of first dorsal fin reaching spine of second fin in female; anal-fin rays I,10; pectoralfin rays 16–17, two upper rays with free tips; pelvic fins I,5, joined to form a disc, moderately emarginate (the shortest branches of fifth ray about 4/5 length of longest branches of fourth ray), with pelvic frenum welldeveloped, frenum height in midline 1/3–2/5 length of pelvic spine; longitudinal scale series 33–35; transverse scale rows to base of first dorsal fin and to base of second dorsal fin with same value, 9; circumpeduncular scale rows 12; head with seven transverse suborbital rows of sensory papillae, five rows in front of row
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<emphasis id="B936AA175E2AFFDC79C3F9C0FB2BEE48" box="[1252,1266,1555,1579]" italics="true" pageId="9" pageNumber="522">b</emphasis>
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and two above row
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<emphasis id="B936AA175E2AFFDC7DEDF9E5FF01EE2D" box="[202,216,1590,1614]" italics="true" pageId="9" pageNumber="522">b</emphasis>
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, sixth row interrupted by row
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<emphasis id="B936AA175E2AFFDC7F17F9E5FDE7EE2D" box="[560,574,1590,1614]" italics="true" pageId="9" pageNumber="522">b</emphasis>
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; no tentacle above upper eye margin; anterior nostril tubular, without flap at its tip, posterior nostril slightly raised; head pigmented except for pale band from lower margin of eye to rear part of mouth, including angle of jaw; first dorsal fin with a nearly square dark maroon, suffused with black, blotch larger than pupil on second and third membranes at base of fin, linked with first spine by a dark band across first membrane; preserved specimen also with oblique broad dark band visible above and behind the blotch, covering upper and posterior parts of fin.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358258E5E2AFFD87DE0F8DDFE78EACB" lastPageId="13" lastPageNumber="526" pageId="9" pageNumber="522" type="description">
|
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<paragraph id="8BFD76055E2AFFDE7DE0F8DDFD7AE906" blockId="9.[151,1437,1194,2011]" lastBlockId="11.[151,1437,151,357]" lastPageId="11" lastPageNumber="524" pageId="9" pageNumber="522">
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||
<emphasis id="B936AA175E2AFFDC7DE0F8DDFE8BEF44" bold="true" box="[199,338,1806,1831]" pageId="9" pageNumber="522">Description</emphasis>
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(all morphometrics values in the text are presented as male
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||
<typeStatus id="54F9C8A75E2AFFDC7EDCF8DDFB84EF44" box="[1019,1117,1806,1831]" pageId="9" pageNumber="522" type="holotype">holotype</typeStatus>
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||
first and female
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<typeStatus id="54F9C8A75E2AFFDC7839F8DCFAA6EF44" box="[1310,1407,1807,1831]" pageId="9" pageNumber="522" type="paratype">paratype</typeStatus>
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in parentheses) (Table 1). Body moderately elongate, depth at pelvic-fin origin 4.7 (4.7) in SL, at anal-fin origin 5.4 (5.1) in SL, and laterally compressed, body width at pectoral-fin origin 5.5 (6.0) in SL, 1.2 (1.3) in depth. Preanalfin length 1.6 (1.7) in SL, predorsal length 2.9 (2.6) in SL; prepelvic distance 3.0 (3.2) in SL. Head length 3.2 (3.3) in SL; dorsal profile of head almost smoothly convex, dorsal part of eye weakly projecting above contour; snout with convex sloping profile, its length 14.3 (14.9) in SL,
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||
<quantity id="4CBADBE05E2AFFDC7E02F811FCBFEFB8" box="[805,870,1986,2011]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" pageId="9" pageNumber="522" unit="in" value="4.5">4.5 in</quantity>
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head length. Eye diameter 10.8 (11.4) in SL, 3.4 (3.4) in head length; no tentacle above upper eye margin; interorbital space very narrow. Mouth oblique, terminal, lips anteriorly ending equally, upper lip width uniform, maxilla reaching to below pupil, upper jaw length 11.9 (12.0) in SL, 3.7 (3.6) in head length. Several rows of small pointed teeth in each jaw. Tongue truncate. Gill opening ending at level of lower base of pectoral fin. Anterior nostril tubular, without lappet on posterior rim; posterior nostril slightly raised. Caudal peduncle depth 8.3 (7.7) in SL, 2.6 (2.3) in head length; caudal peduncle length 4.8 (5.5) in SL, 1.5 (1.7) in head length.
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</paragraph>
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||
<caption id="DF3D268D5E29FFDF7DB0F853FAD6EFB9" httpUri="https://zenodo.org/record/230290/files/figure.png" pageId="10" pageNumber="523" targetBox="[179,1407,193,1897]" targetPageId="10">
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||
<paragraph id="8BFD76055E29FFDF7DB0F853FAD6EFB9" blockId="10.[151,1436,1920,2010]" pageId="10" pageNumber="523">
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||
<emphasis id="B936AA175E29FFDF7DB0F853FECDEFF6" bold="true" box="[151,276,1920,1942]" pageId="10" pageNumber="523">FIGURE 3.</emphasis>
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||
<taxonomicName id="4C420D865E29FFDF7C3AF853FDBAEFF5" box="[285,611,1920,1942]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="523" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
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<emphasis id="B936AA175E29FFDF7C3AF853FDBAEFF5" box="[285,611,1920,1942]" italics="true" pageId="10" pageNumber="523">Coryogalops pseudomonospilus</emphasis>
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</taxonomicName>
|
||
<emphasis id="B936AA175E29FFDF7F4CF853FD01EFF6" bold="true" box="[619,728,1920,1941]" pageId="10" pageNumber="523">
|
||
<taxonomicNameLabel id="A205176C5E29FFDF7F4CF853FD67EFF6" box="[619,702,1920,1941]" pageId="10" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
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||
A
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</emphasis>
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||
: SMF 35231, holotype, male, 21.5 + 5.5 mm, Al Khuraybah, Saudi Arabia, Red Sea, live specimen;
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||
<emphasis id="B936AA175E29FFDF7CD7F871FDD8EFD4" bold="true" box="[496,513,1954,1975]" pageId="10" pageNumber="523">B</emphasis>
|
||
: the same, preserved specimen.
|
||
<emphasis id="B936AA175E29FFDF7E70F871FCB0EFDB" bold="true" box="[855,873,1954,1976]" pageId="10" pageNumber="523">C</emphasis>
|
||
: KAUMM 260, paratype, female, 19.3 + 4.8 mm, Al Khuraybah, Saudi Arabia, Red Sea, preserved specimen. Photos by S.V. Bogorodsky (A), M. Kovačić (B and C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3D268D5E28FFDE7DB0FA9AFE1EED1D" httpUri="https://zenodo.org/record/230291/files/figure.png" pageId="11" pageNumber="524" targetBox="[289,1436,538,1256]" targetPageId="11">
|
||
<paragraph id="8BFD76055E28FFDE7DB0FA9AFE1EED1D" blockId="11.[151,1436,1353,1406]" pageId="11" pageNumber="524">
|
||
<emphasis id="B936AA175E28FFDE7DB0FA9AFECDED3D" bold="true" box="[151,276,1353,1375]" pageId="11" pageNumber="524">FIGURE 4.</emphasis>
|
||
Cephalic sensory systems of
|
||
<taxonomicName id="4C420D865E28FFDE7F6EFA9AFC56ED3C" box="[585,911,1353,1375]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="524" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
|
||
<emphasis id="B936AA175E28FFDE7F6EFA9AFC56ED3C" box="[585,911,1353,1375]" italics="true" pageId="11" pageNumber="524">Coryogalops pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A205176C5E28FFDE7EB0FA99FC30ED3C" box="[919,1001,1354,1375]" pageId="11" pageNumber="524" rank="species">
|
||
<emphasis id="B936AA175E28FFDE7EB0FA99FC30ED3C" bold="true" box="[919,1001,1354,1375]" pageId="11" pageNumber="524">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, KAUMM 260, paratype. Terminology in text. Drawing by M. Kovačić.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD76055E28FFDE7DE0FA78FC24EFD8" blockId="11.[151,1437,1450,2015]" pageId="11" pageNumber="524">
|
||
<emphasis id="B936AA175E28FFDE7DE0FA78FF20EDA1" box="[199,249,1451,1474]" italics="true" pageId="11" pageNumber="524">Fins</emphasis>
|
||
. First dorsal fin VI, second dorsal fin I,11, second to sixth spines of first dorsal fin reaching spine of second fin in male, and at least fourth spine of first dorsal fin reaching spine of second fin in female; anal fin I,10; pectoral-fin rays 16–17 (
|
||
<typeStatus id="54F9C8A75E28FFDE7C88FA21FDCAEE68" box="[431,531,1522,1547]" pageId="11" pageNumber="524" type="holotype">holotype</typeStatus>
|
||
16/16,
|
||
<typeStatus id="54F9C8A75E28FFDE7F4CFA20FD12EE68" box="[619,715,1523,1547]" pageId="11" pageNumber="524" type="paratype">paratype</typeStatus>
|
||
17/right fin missing), all rays branched, two upper pectoral-fin rays with well-developed free tips, tip of first ray about 1/2 ray length; branched caudal-fin rays 15, segmented rays 17; pelvic fins I,5, disc complete, moderately emarginate (shortest branches of fifth ray compared to longest branches of fourth ray: 80.0% in
|
||
<typeStatus id="54F9C8A75E28FFDE7F2EF98DFDB2EE14" box="[521,619,1630,1655]" pageId="11" pageNumber="524" type="holotype">holotype</typeStatus>
|
||
and 83.3% in
|
||
<typeStatus id="54F9C8A75E28FFDE7E2AF98CFCABEE14" box="[781,882,1631,1655]" pageId="11" pageNumber="524" type="paratype">paratype</typeStatus>
|
||
), pelvic frenum well-developed, frenum height in midline 1/3–2/5 length of pelvic spine. Origin of first dorsal fin behind vertical at pectoral-fin base, second dorsalfin spine longest in male
|
||
<typeStatus id="54F9C8A75E28FFDE7C95F975FDCEEEDC" box="[434,535,1702,1727]" pageId="11" pageNumber="524" type="holotype">holotype</typeStatus>
|
||
,
|
||
<quantity id="4CBADBE05E28FFDE7F04F975FDBAEEDC" box="[547,611,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="1.4731999999999998" pageId="11" pageNumber="524" unit="in" value="5.8">5.8 in</quantity>
|
||
SL,
|
||
<quantity id="4CBADBE05E28FFDE7FBEF974FD00EEDC" box="[665,729,1702,1727]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="11" pageNumber="524" unit="in" value="1.8">1.8 in</quantity>
|
||
head length, second and third equally long in female
|
||
<typeStatus id="54F9C8A75E28FFDE7812F97BFA41EEA3" box="[1333,1432,1704,1728]" pageId="11" pageNumber="524" type="paratype">paratype</typeStatus>
|
||
,
|
||
<quantity id="4CBADBE05E28FFDE7DB0F919FF03EE80" box="[151,218,1738,1763]" metricMagnitude="-1" metricUnit="m" metricValue="1.3208" pageId="11" pageNumber="524" unit="in" value="5.2">5.2 in</quantity>
|
||
SL,
|
||
<quantity id="4CBADBE05E28FFDE7C33F919FE8EEE80" box="[276,343,1738,1763]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" pageId="11" pageNumber="524" unit="in" value="1.6">1.6 in</quantity>
|
||
head length, remaining spines shorter; spine of second dorsal fin 7.7 (7.7) in SL; first to third dorsal-fin rays longest 6.1 (5.9) in SL; dorsal-fins base 2.2 (2.2) in SL. Origin of anal fin below base of first or second dorsal-fin soft ray; anal-fin spine 12.7 (11.4) in SL; tenth anal-fin soft ray longest, 6.9 (6.7) in SL; anal-fin base 4.6 (4.6) in SL. Caudal fin rounded and short (damaged in male
|
||
<typeStatus id="54F9C8A75E28FFDE7E83F8E5FBD2EF2C" box="[932,1035,1846,1871]" pageId="11" pageNumber="524" type="holotype">holotype</typeStatus>
|
||
), its length 3.9 (4.0) in SL. Pectoral fins more or less rounded, extending behind vertical to origin of second dorsal fin, tenth to eleventh ray longest, 3.4 (3.9) in SL. Origin of pelvic fins about at vertical to base of pectoral fins, pelvic spine 14.3 (16.1) in SL; pelvic-fins length 4.4 (4.1) in SL, reaching anus in female, almost reaching anus in male.
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E28FFD97DE0F814FD37E97F" blockId="11.[151,1437,1450,2015]" lastBlockId="12.[151,1437,151,2005]" lastPageId="12" lastPageNumber="525" pageId="11" pageNumber="524">
|
||
<emphasis id="B936AA175E28FFDE7DE0F814FE94EFBD" box="[199,333,1991,2014]" italics="true" pageId="11" pageNumber="524">Squamation</emphasis>
|
||
. Scales progressively larger posteriorly on body; scales ctenoid except cycloid anterior close to belly. Head naked, no predorsal scales and no scales on side of nape anterior to a line from origin of third spine of first dorsal fin to behind upper end of pectoral-fin base; no scales on prepectoral area, prepelvic area, central belly part and fins. Longitudinal scale series 33-35; transverse scale rows to base of first dorsal fin and to base of second dorsal fin same value, 9; circumpeduncular scales 12.
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E2FFFD97DE0FEFBFD02EC33" blockId="12.[151,1437,151,2005]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD97DE0FEFBFE33E923" box="[199,490,296,320]" italics="true" pageId="12" pageNumber="525">Cephalic sensory system</emphasis>
|
||
(
|
||
<figureCitation id="13796A805E2FFFD97F27FEF4FD94E923" box="[512,589,295,320]" captionStart="FIGURE 4" captionStartId="11.[151,250,1353,1375]" captionTargetBox="[289,1436,538,1256]" captionTargetId="figure@11.[151,1436,439,1331]" captionTargetPageId="11" captionText="FIGURE 4. Cephalic sensory systems of Coryogalops pseudomonospilus sp. nov., KAUMM 260, paratype. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/230291/files/figure.png" pageId="12" pageNumber="525">Fig. 4</figureCitation>
|
||
). Head with separate anterior and posterior oculoscapular canals, and preopercular canals, with pores
|
||
<emphasis id="B936AA175E2FFFD97CD9FE83FCD2E904" box="[510,779,331,359]" italics="true" pageId="12" pageNumber="525">σ, λ, κ, ω, α, β, ρ, ρ1, ρ2,</emphasis>
|
||
and
|
||
<emphasis id="B936AA175E2FFFD97E64FE83FC5AE904" box="[835,899,335,359]" italics="true" pageId="12" pageNumber="525">γ, δ, ε</emphasis>
|
||
respectively. Rows of head sensory papillae on left side of
|
||
<typeStatus id="54F9C8A75E2FFFD97C3DFEA1FEA5E9E8" box="[282,380,370,395]" pageId="12" pageNumber="525" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F9C8A75E2FFFD97C92FEA0FDC1E9E8" box="[437,536,371,395]" pageId="12" pageNumber="525" type="paratype">paratype</typeStatus>
|
||
, respectively: (1)
|
||
<emphasis id="B936AA175E2FFFD97FC1FEA0FC8FE9E8" box="[742,854,371,395]" italics="true" pageId="12" pageNumber="525">preorbital</emphasis>
|
||
: snout with four median preorbital series, row
|
||
<emphasis id="B936AA175E2FFFD97856FEA0FAA5E9E9" box="[1393,1404,371,394]" italics="true" pageId="12" pageNumber="525">r</emphasis>
|
||
(4 and 3) above and medially to posterior nostril, horizontal row
|
||
<emphasis id="B936AA175E2FFFD97E68FE49FCBBE9C7" box="[847,866,406,433]" italics="true" pageId="12" pageNumber="525">s1</emphasis>
|
||
(3 and 2) below pore
|
||
<emphasis id="B936AA175E2FFFD9797EFE49FBBEE9D2" box="[1113,1127,410,433]" italics="true" pageId="12" pageNumber="525">σ</emphasis>
|
||
, horizontal row
|
||
<emphasis id="B936AA175E2FFFD97807FE49FAEAE9C7" box="[1312,1331,406,433]" italics="true" pageId="12" pageNumber="525">s2</emphasis>
|
||
(3 and 2) medially to anterior nostril and vertical
|
||
<emphasis id="B936AA175E2FFFD97F7CFE12FDB7E9A8" box="[603,622,445,472]" italics="true" pageId="12" pageNumber="525">s3</emphasis>
|
||
(3 and 2) above upper lip. Lateral series
|
||
<emphasis id="B936AA175E2FFFD97965FE12FB97E9BB" box="[1090,1102,449,472]" italics="true" pageId="12" pageNumber="525">c</emphasis>
|
||
in four parts: superior (
|
||
<emphasis id="B936AA175E2FFFD97878FE12FAAAE9A8" box="[1375,1395,445,472]" italics="true" pageId="12" pageNumber="525">c2</emphasis>
|
||
) as two rows between anterior nostril and eye (2 + 2 and 3 + 2); middle (
|
||
<emphasis id="B936AA175E2FFFD97E80FE3BFC62E991" box="[935,955,484,511]" italics="true" pageId="12" pageNumber="525">c1</emphasis>
|
||
) (4 and 3) from anterior nostril downward and posteriorly to upper lip; inferior rows, upper horizontal
|
||
<emphasis id="B936AA175E2FFFD97E18FDDFFC8AEA4B" box="[831,851,524,552]" italics="true" pageId="12" pageNumber="525">c2</emphasis>
|
||
(4 and 3) and lower horizontal
|
||
<emphasis id="B936AA175E2FFFD9799BFDDFFB08EA4B" box="[1212,1233,524,552]" italics="true" pageId="12" pageNumber="525">c1</emphasis>
|
||
(2 and 2) starting anteriorly at upper lip. (2)
|
||
<emphasis id="B936AA175E2FFFD97C98FDE1FDEBEA29" box="[447,562,562,586]" italics="true" pageId="12" pageNumber="525">suborbital</emphasis>
|
||
: seven transverse suborbital rows (
|
||
<emphasis id="B936AA175E2FFFD97E9AFDE1FC31EA2A" box="[957,1000,562,586]" italics="true" pageId="12" pageNumber="525">1–7</emphasis>
|
||
) of sensory papillae, five rows in front of row
|
||
<emphasis id="B936AA175E2FFFD97DCFFD85FF2FEA0D" box="[232,246,598,622]" italics="true" pageId="12" pageNumber="525">b</emphasis>
|
||
and two above row
|
||
<emphasis id="B936AA175E2FFFD97CFCFD85FE30EA0D" box="[475,489,598,622]" italics="true" pageId="12" pageNumber="525">b</emphasis>
|
||
, sixth row interrupted by row
|
||
<emphasis id="B936AA175E2FFFD97E18FD85FC94EA0D" box="[831,845,598,622]" italics="true" pageId="12" pageNumber="525">b</emphasis>
|
||
; rows
|
||
<emphasis id="B936AA175E2FFFD97EBDFD85FC71EA0D" box="[922,936,598,622]" italics="true" pageId="12" pageNumber="525">2</emphasis>
|
||
and
|
||
<emphasis id="B936AA175E2FFFD97EF9FD85FC35EA0D" box="[990,1004,598,622]" italics="true" pageId="12" pageNumber="525">3</emphasis>
|
||
begin distant from orbit, rows
|
||
<emphasis id="B936AA175E2FFFD9786EFD85FA8EEA0D" box="[1353,1367,598,622]" italics="true" pageId="12" pageNumber="525">4</emphasis>
|
||
and
|
||
<emphasis id="B936AA175E2FFFD978A9FD85FA45EA0E" box="[1422,1436,598,621]" italics="true" pageId="12" pageNumber="525">5</emphasis>
|
||
short, inferior section of row
|
||
<emphasis id="B936AA175E2FFFD97CC7FDA9FE37EAF1" box="[480,494,634,658]" italics="true" pageId="12" pageNumber="525">6</emphasis>
|
||
well developed, extended to the level of row
|
||
<emphasis id="B936AA175E2FFFD97EC9FDA9FC25EAF1" box="[1006,1020,634,658]" italics="true" pageId="12" pageNumber="525">d</emphasis>
|
||
(
|
||
<emphasis id="B936AA175E2FFFD9792AFDA9FBC2EAF1" box="[1037,1051,634,658]" italics="true" pageId="12" pageNumber="525">1</emphasis>
|
||
: 6 and 6,
|
||
<emphasis id="B936AA175E2FFFD979ACFDA9FB40EAF1" box="[1163,1177,634,658]" italics="true" pageId="12" pageNumber="525">2</emphasis>
|
||
: 5 and 4,
|
||
<emphasis id="B936AA175E2FFFD9782FFDA9FACFEAF1" box="[1288,1302,634,658]" italics="true" pageId="12" pageNumber="525">3</emphasis>
|
||
: 5 and 4,
|
||
<emphasis id="B936AA175E2FFFD978A1FDA9FA4DEAF1" box="[1414,1428,634,658]" italics="true" pageId="12" pageNumber="525">4</emphasis>
|
||
: 5 and 4,
|
||
<emphasis id="B936AA175E2FFFD97DDEFD4DFEDEEAD6" box="[249,263,670,693]" italics="true" pageId="12" pageNumber="525">5</emphasis>
|
||
: 3 and 3,
|
||
<emphasis id="B936AA175E2FFFD97C5EFD4DFE4BEAD5" box="[377,402,670,694]" italics="true" pageId="12" pageNumber="525">6s</emphasis>
|
||
:
|
||
<date id="FFFC50C55E2FFFD97C85FD4DFE21EAD5" box="[418,504,669,694]" pageId="12" pageNumber="525">4 and 4</date>
|
||
,
|
||
<date id="FFFC50C55E2FFFD97F23FD4DFDC0EAD5" box="[516,537,670,694]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD97F23FD4DFDC0EAD5" box="[516,537,670,694]" italics="true" pageId="12" pageNumber="525">6i</emphasis>
|
||
</date>
|
||
: 7 and 7,
|
||
<emphasis id="B936AA175E2FFFD97FABFD4DFD43EAD6" box="[652,666,670,693]" italics="true" pageId="12" pageNumber="525">7</emphasis>
|
||
: 3 and 3). Longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97EC6FD4DFC36EAD5" box="[993,1007,670,694]" italics="true" pageId="12" pageNumber="525">b</emphasis>
|
||
short (5 and 5) extending forwards to row
|
||
<emphasis id="B936AA175E2FFFD97DF7FD11FF07EABA" box="[208,222,706,729]" italics="true" pageId="12" pageNumber="525">5</emphasis>
|
||
. Longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97CEEFD11FE0EEAB9" box="[457,471,706,730]" italics="true" pageId="12" pageNumber="525">d</emphasis>
|
||
(16 and 16) continuous, supralabial and horizontal part joined. (3)
|
||
<emphasis id="B936AA175E2FFFD9782BFD11FEC3EA9D" italics="true" pageId="12" pageNumber="525">preoperculomandibular</emphasis>
|
||
: external row
|
||
<emphasis id="B936AA175E2FFFD97CE7FD34FE15EA9D" box="[448,460,743,766]" italics="true" pageId="12" pageNumber="525">e</emphasis>
|
||
(24 + 18 and 22 + 12) and internal row
|
||
<emphasis id="B936AA175E2FFFD97EBAFD34FC7CEA9D" box="[925,933,743,766]" italics="true" pageId="12" pageNumber="525">i</emphasis>
|
||
(9 + 8 and 9 + 9) uniserial and divided into anterior and posterior sections; row
|
||
<emphasis id="B936AA175E2FFFD97F0AFCDEFDECEB46" box="[557,565,781,805]" italics="true" pageId="12" pageNumber="525">f</emphasis>
|
||
longitudinal (5 and 5). (4)
|
||
<emphasis id="B936AA175E2FFFD97E4EFCDEFBD0EB46" box="[873,1033,781,805]" italics="true" pageId="12" pageNumber="525">oculoscapular</emphasis>
|
||
: anterior longitudinal row
|
||
<date id="FFFC50C55E2FFFD9781EFCDEFA94EB74" box="[1337,1357,777,804]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD9781EFCDEFA94EB74" box="[1337,1357,777,804]" italics="true" pageId="12" pageNumber="525">x1</emphasis>
|
||
</date>
|
||
(7 and 5) above pore
|
||
<emphasis id="B936AA175E2FFFD97C10FCE7FE9DEB28" box="[311,324,820,843]" italics="true" pageId="12" pageNumber="525">ρ</emphasis>
|
||
, not extending forwards to pore
|
||
<emphasis id="B936AA175E2FFFD97F95FCE7FD19EB2F" box="[690,704,820,844]" italics="true" pageId="12" pageNumber="525">β</emphasis>
|
||
but ending above
|
||
<emphasis id="B936AA175E2FFFD97EA9FCE7FC40EB28" box="[910,921,820,843]" italics="true" pageId="12" pageNumber="525">z</emphasis>
|
||
, posterior longitudinal row
|
||
<date id="FFFC50C55E2FFFD979F5FCE7FB3EEB5D" box="[1234,1255,816,843]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD979F5FCE7FB3EEB5D" box="[1234,1255,816,843]" italics="true" pageId="12" pageNumber="525">x2</emphasis>
|
||
</date>
|
||
(3 and 3) above
|
||
<emphasis id="B936AA175E2FFFD97DB0FC88FF75EB06" box="[151,172,855,882]" italics="true" pageId="12" pageNumber="525">ρ2</emphasis>
|
||
; row
|
||
<emphasis id="B936AA175E2FFFD97DD7FC88FF22EB11" box="[240,251,859,882]" italics="true" pageId="12" pageNumber="525">z</emphasis>
|
||
(5 and 4) with lower end above pore
|
||
<emphasis id="B936AA175E2FFFD97F8BFC88FD6EEB11" box="[684,695,859,882]" italics="true" pageId="12" pageNumber="525">γ</emphasis>
|
||
, row
|
||
<emphasis id="B936AA175E2FFFD97FDDFC88FCD1EB11" box="[762,776,859,882]" italics="true" pageId="12" pageNumber="525">q</emphasis>
|
||
as single papilla behind pore
|
||
<emphasis id="B936AA175E2FFFD97979FC88FBB2EB11" box="[1118,1131,859,882]" italics="true" pageId="12" pageNumber="525">ρ</emphasis>
|
||
, vertical row
|
||
<emphasis id="B936AA175E2FFFD9782CFC88FAF5EB11" box="[1291,1324,859,882]" italics="true" pageId="12" pageNumber="525">trp</emphasis>
|
||
(2 and 2) behind
|
||
<date id="FFFC50C55E2FFFD97DCCFC51FF26EBEF" box="[235,255,894,921]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD97DCCFC51FF26EBEF" box="[235,255,894,921]" italics="true" pageId="12" pageNumber="525">x1</emphasis>
|
||
</date>
|
||
starting ventrally in front of
|
||
<emphasis id="B936AA175E2FFFD97F62FC51FD82EBEF" box="[581,603,894,921]" italics="true" pageId="12" pageNumber="525">ρ1</emphasis>
|
||
, row
|
||
<emphasis id="B936AA175E2FFFD97FBBFC51FD71EBFA" box="[668,680,898,921]" italics="true" pageId="12" pageNumber="525">y</emphasis>
|
||
(2 and 3) vertical behind pore
|
||
<emphasis id="B936AA175E2FFFD9792EFC51FBC7EBEF" box="[1033,1054,894,921]" italics="true" pageId="12" pageNumber="525">ρ2</emphasis>
|
||
. Axillary vertical rows
|
||
<emphasis id="B936AA175E2FFFD9780BFC51FA94EBEF" box="[1324,1357,894,921]" italics="true" pageId="12" pageNumber="525">as1</emphasis>
|
||
(5 and 4),
|
||
<emphasis id="B936AA175E2FFFD97D9BFC7AFF04EBD1" box="[188,221,932,960]" italics="true" pageId="12" pageNumber="525">as2</emphasis>
|
||
(4 and 4),
|
||
<emphasis id="B936AA175E2FFFD97C7FFC7AFEA0EBD1" box="[344,377,932,960]" italics="true" pageId="12" pageNumber="525">as3</emphasis>
|
||
(4 and 4), row
|
||
<emphasis id="B936AA175E2FFFD97F01FC7BFD9DEBD1" box="[550,580,932,960]" italics="true" pageId="12" pageNumber="525">la1</emphasis>
|
||
as single papilla above
|
||
<emphasis id="B936AA175E2FFFD97E77FC7AFCA8EBD1" box="[848,881,932,960]" italics="true" pageId="12" pageNumber="525">as2</emphasis>
|
||
, row
|
||
<emphasis id="B936AA175E2FFFD97E95FC7BFC09EBD1" box="[946,976,932,960]" italics="true" pageId="12" pageNumber="525">la2</emphasis>
|
||
as two papillae above
|
||
<emphasis id="B936AA175E2FFFD979F7FC7AFB28EBD1" box="[1232,1265,932,960]" italics="true" pageId="12" pageNumber="525">as3</emphasis>
|
||
. (5)
|
||
<emphasis id="B936AA175E2FFFD97800FC7BFA4DEBA3" box="[1319,1428,936,960]" italics="true" pageId="12" pageNumber="525">opercular</emphasis>
|
||
: transverse row
|
||
<emphasis id="B936AA175E2FFFD97C66FC1EFE8EEB87" box="[321,343,973,996]" italics="true" pageId="12" pageNumber="525">ot</emphasis>
|
||
(18 and 16); superior longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97E37FC1EFCF0EB87" box="[784,809,973,996]" italics="true" pageId="12" pageNumber="525">os</emphasis>
|
||
(7 and 6); and interior longitudinal row
|
||
<emphasis id="B936AA175E2FFFD979CBFC1EFADBEB87" box="[1260,1282,973,996]" italics="true" pageId="12" pageNumber="525">oi</emphasis>
|
||
(4 and 4). (6)
|
||
<emphasis id="B936AA175E2FFFD97DB0FC23FE99EC6B" box="[151,320,1008,1032]" italics="true" pageId="12" pageNumber="525">anterior dorsal</emphasis>
|
||
: anterior transverse row
|
||
<emphasis id="B936AA175E2FFFD97F71FC22FDBDEC6B" box="[598,612,1009,1032]" italics="true" pageId="12" pageNumber="525">n</emphasis>
|
||
(5 and 4) behind pore
|
||
<emphasis id="B936AA175E2FFFD97E47FC22FCA2EC6B" box="[864,891,1009,1032]" italics="true" pageId="12" pageNumber="525">ω,</emphasis>
|
||
transverse row
|
||
<emphasis id="B936AA175E2FFFD9790CFC22FBE0EC6B" box="[1067,1081,1009,1032]" italics="true" pageId="12" pageNumber="525">o</emphasis>
|
||
(3 and 3) distant from fellow in dorsal midline; longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97F2AFBC6FDC2EC4F" box="[525,539,1045,1068]" italics="true" pageId="12" pageNumber="525">g</emphasis>
|
||
(3 and 3) not reaching row
|
||
<emphasis id="B936AA175E2FFFD97E7BFBC6FCB3EC4F" box="[860,874,1045,1068]" italics="true" pageId="12" pageNumber="525">o</emphasis>
|
||
, longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97962FBC6FB80EC4F" box="[1093,1113,1045,1068]" italics="true" pageId="12" pageNumber="525">m</emphasis>
|
||
(3 and 3) below and behind row
|
||
<emphasis id="B936AA175E2FFFD97DEEFBEAFF06EC33" box="[201,223,1081,1104]" italics="true" pageId="12" pageNumber="525">g,</emphasis>
|
||
longitudinal row
|
||
<emphasis id="B936AA175E2FFFD97C81FBEBFE6DEC33" box="[422,436,1080,1104]" italics="true" pageId="12" pageNumber="525">h</emphasis>
|
||
(3 + 3 and 3 + 3) divided.
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E2FFFD97DE0FB8FFAC4EF43" blockId="12.[151,1437,151,2005]" pageId="12" pageNumber="525">
|
||
<emphasis id="B936AA175E2FFFD97DE0FB8FFEDEEC17" box="[199,263,1116,1140]" italics="true" pageId="12" pageNumber="525">Color</emphasis>
|
||
. In life only male (
|
||
<figureCitation id="13796A805E2FFFD97CF9FB8FFDF8EC17" box="[478,545,1116,1141]" captionStart="FIGURE 3" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[179,1407,193,1897]" captionTargetId="figure@10.[179,1408,193,1899]" captionTargetPageId="10" captionText="FIGURE 3. Coryogalops pseudomonospilus sp. nov. A: SMF 35231, holotype, male, 21.5 + 5.5 mm, Al Khuraybah, Saudi Arabia, Red Sea, live specimen; B: the same, preserved specimen. C: KAUMM 260, paratype, female, 19.3 + 4.8 mm, Al Khuraybah, Saudi Arabia, Red Sea, preserved specimen. Photos by S. V. Bogorodsky (A), M. Kovačić (B and C)." httpUri="https://zenodo.org/record/230290/files/figure.png" pageId="12" pageNumber="525">Fig. 3</figureCitation>
|
||
A): body yellowish white, densely mottled with brown. Six dark spots at lateral midline, two below first dorsal fin, three below second dorsal fin, and one in centre of caudal peduncle, pale interspace between spots covered with scattered melanophores; another dark spot at end of caudal peduncle is part of caudal fin-base mark. Internal black stripe visible along vertebral column. Six yellowish pale saddles along back, first and third saddles at origin of first and second dorsal fins, respectively, second and fourth at middle of each fin, fifth anteriorly on caudal peduncle, and last at end of caudal peduncle. Head densely mottled with dark brown, brown, and few, narrow, small whitish patches; an irregular whitish bar containing few tiny dark brown spots extending ventrally from eye across cheek and posterior part of lips, in front of this bar a triangular brown band across lips; two dark brown spots visible on lower opercle; pupil dark green with yellow outer ring, iris with five alternating brown and grey bands; first dorsal fin semitransparent with scattered melanophores on basal twothirds, and a nearly square, dark maroon blotch, suffused with black, and larger than pupil on second and third membranes at base of fin, linked to first spine by a dark band across first membrane (area below band on first membrane hyaline), and an oblique whitish band containing few melanophores above blotch; outer part of interspinous membranes of first dorsal fin whitish; second dorsal fin translucent with dark brown pigment extending basally onto several rays from dorsal midline and with 3 or 4 spots along each ray, scattered melanophores on basal half of fin; caudal fin transparent, with scattered melanophores, and with a dark brown ‘X’- shaped mark on base of rays, anterior branches of mark longer and reaching procurrent rays on upper and lower edge of fin; anal fin transparent, with many melanophores basally; pectoral fins transparent with scattered melanophores and two dark brown spots, each on upper and lower base of pectoral-fin rays, more or less contiguous with pigmentation of pectoral-fin base; pelvic fins transparent with scattered melanophores.
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E2FFFD87DE0F8FEFE78EACB" blockId="12.[151,1437,151,2005]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="526" pageId="12" pageNumber="525">
|
||
In alcohol (
|
||
<figureCitation id="13796A805E2FFFD97C6FF8FFFE4FEF27" box="[328,406,1836,1861]" captionStart="FIGURE 3" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[179,1407,193,1897]" captionTargetId="figure@10.[179,1408,193,1899]" captionTargetPageId="10" captionText="FIGURE 3. Coryogalops pseudomonospilus sp. nov. A: SMF 35231, holotype, male, 21.5 + 5.5 mm, Al Khuraybah, Saudi Arabia, Red Sea, live specimen; B: the same, preserved specimen. C: KAUMM 260, paratype, female, 19.3 + 4.8 mm, Al Khuraybah, Saudi Arabia, Red Sea, preserved specimen. Photos by S. V. Bogorodsky (A), M. Kovačić (B and C)." httpUri="https://zenodo.org/record/230290/files/figure.png" pageId="12" pageNumber="525">Figs. 3</figureCitation>
|
||
B, C): Head and body paler than in live specimen, light yellowish-brown, with brown color pattern. Melanophores on body more concentrated at scale edges. Six brown marks along lateral midline, less distinctive in male: two below first dorsal fin, three below second dorsal fin and one on caudal peduncle. Another dark spot at end of caudal peduncle is part of caudal fin base mark. Six pale yellowish saddles along back, first and third saddles below origin of first and second dorsal fins, respectively, second and fourth below middle of each fin, fifth anteriorly at caudal peduncle and last at end of caudal peduncle. Head covered with melanophores, pigment more intensive and darker in male. Differences in density of melanophores present on predorsal, cheek and opercle but without discernible pattern, except for pale bar from lower edge of eye over anterior cheek to rear part of mouth, including the angle of jaw and continuing over lips. Iris uniformly dark, without visible stripes present in live specimen. Underside of head uniformly pigmented with densely distributed melanophores in male and widely scattered in female. No gular spots, except for two brown spots present on lower opercle, less distinct in female. Chest pigmented only in male, whitish in female. Dark brown ‘┐’-shaped blotch somewhat larger than pupil size basally on first three membranes of first dorsal fin; oblique broad dark band visible above and behind blotch, covering upper and posterior parts of fin. Second dorsal fin dark, with scattered melanophores. Caudal fin transparent, with widely scattered melanophores and dark brown ‘X’-shaped mark at base of rays, anterior branches of mark longer and reaching procurrent rays on upper and lower edge of fin. Anal fin densely pigmented with melanophores. Pectoral fins transparent with scattered melanophores, two marks present on upper and lower base of pectoral fin rays, darker in male, still recognizable in female, more or less contiguous with pigmentation of pectoral-fin base. Pelvic fins transparent, moderately pigmented with scattered melanophores only in male, unpigmented in female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358258E5E2EFFD87DE0FD67FDE7EB76" pageId="13" pageNumber="526" type="etymology">
|
||
<paragraph id="8BFD76055E2EFFD87DE0FD67FDE7EB76" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="526">
|
||
<emphasis id="B936AA175E2EFFD87DE0FD67FE9FEAAE" bold="true" box="[199,326,692,717]" pageId="13" pageNumber="526">Etymology</emphasis>
|
||
. The species is named
|
||
<taxonomicName id="4C420D865E2EFFD87F79FD67FC10EAAF" box="[606,969,692,716]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">
|
||
<emphasis id="B936AA175E2EFFD87F79FD67FC10EAAF" box="[606,969,692,716]" italics="true" pageId="13" pageNumber="526">Coryogalops pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
, because of its close resemblance to
|
||
<taxonomicName id="4C420D865E2EFFD878A5FD67FECEEA93" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD878A5FD67FECEEA93" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
, particularly because of its similarity in coloration, including the superficial similarity of the black mark anteriorly on the first dorsal fin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358258E5E2EFFD87DE0FCCCFA5BEBA4" pageId="13" pageNumber="526" type="distribution">
|
||
<paragraph id="8BFD76055E2EFFD87DE0FCCCFA5BEBA4" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="526">
|
||
<emphasis id="B936AA175E2EFFD87DE0FCCCFE2DEB5B" bold="true" box="[199,500,799,824]" pageId="13" pageNumber="526">Distribution and habitat.</emphasis>
|
||
At present known from Al Khuraybah, northern
|
||
<collectingCountry id="F35536955E2EFFD87904FCCCFB63EB5B" box="[1059,1210,799,824]" name="Saudi Arabia" pageId="13" pageNumber="526">Saudi Arabia</collectingCountry>
|
||
, Red Sea.
|
||
<typeStatus id="54F9C8A75E2EFFD87813FCCCFA45EB5B" box="[1332,1436,799,824]" pageId="13" pageNumber="526" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54F9C8A75E2EFFD87DE1FC96FEFFEB3E" box="[198,294,837,861]" pageId="13" pageNumber="526" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C420D865E2EFFD87C6CFC97FDE2EB3F" box="[331,571,836,860]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">
|
||
<emphasis id="B936AA175E2EFFD87C6CFC97FDE2EB3F" box="[331,571,836,860]" italics="true" pageId="13" pageNumber="526">C. pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
were collected at the base of a small isolated coral patch at depth of
|
||
<quantity id="4CBADBE05E2EFFD8781EFC97FA59EB3F" box="[1337,1408,836,860]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="13" pageNumber="526" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">7–8 m</quantity>
|
||
in a large shallow-water lagoon. The lagoon is characterized by mixture of isolated coral reefs and broad flats covered with coral patches and sand. The gobies were collected at the island where reef flat ends by fringing reef with depth at base about
|
||
<quantity id="4CBADBE05E2EFFD87C17FC63FEA0EBA4" box="[304,377,944,968]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="526" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
|
||
, followed by the moderate sandy slope with scattered small coral patches down to
|
||
<quantity id="4CBADBE05E2EFFD8783FFC7CFAA7EBA4" box="[1304,1406,943,968]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="13" pageNumber="526" unit="m" value="13.5" valueMax="15.0" valueMin="12.0">
|
||
12–
|
||
<specimenCount id="9D44BD8C5E2EFFD87863FC7CFAA7EBA4" box="[1348,1406,943,968]" pageId="13" pageNumber="526" type="generic">15 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358258E5E2EFFDA7DE0FC07FEB9EA5F" lastPageId="15" lastPageNumber="528" pageId="13" pageNumber="526" type="discussion">
|
||
<paragraph id="8BFD76055E2EFFD87DE0FC07FC50EC1E" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="526">
|
||
<emphasis id="B936AA175E2EFFD87DE0FC07FEEDEB8E" bold="true" box="[199,308,980,1005]" pageId="13" pageNumber="526">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C420D865E2EFFD87C6BFC07FD65EB8F" box="[332,700,980,1004]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
|
||
<emphasis id="B936AA175E2EFFD87C6BFC07FD65EB8F" box="[332,700,980,1004]" italics="true" pageId="13" pageNumber="526">Coryogalops pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A205176C5E2EFFD87FEAFC07FCE8EB8E" box="[717,817,980,1005]" pageId="13" pageNumber="526" rank="species">
|
||
<emphasis id="B936AA175E2EFFD87FEAFC07FCE8EB8E" bold="true" box="[717,817,980,1005]" pageId="13" pageNumber="526">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from each of the nine known species of
|
||
<taxonomicName id="4C420D865E2EFFD87DB0FC2BFEF1EC73" box="[151,296,1016,1040]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="genus">
|
||
<emphasis id="B936AA175E2EFFD87DB0FC2BFEF1EC73" box="[151,296,1016,1040]" italics="true" pageId="13" pageNumber="526">Coryogalops</emphasis>
|
||
</taxonomicName>
|
||
and the new described
|
||
<taxonomicName id="4C420D865E2EFFD87F10FC2BFD6CEC73" box="[567,693,1016,1040]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="guttatus" status="sp. nov.">
|
||
<emphasis id="B936AA175E2EFFD87F10FC2BFD6CEC73" box="[567,693,1016,1040]" italics="true" pageId="13" pageNumber="526">C. guttatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A205176C5E2EFFD87F9BFC24FCC1EC73" box="[700,792,1015,1040]" pageId="13" pageNumber="526" rank="species">
|
||
<emphasis id="B936AA175E2EFFD87F9BFC24FCC1EC73" bold="true" box="[700,792,1015,1040]" pageId="13" pageNumber="526">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in several characters: absence of a supraorbital tentacle; the shape of the pelvic-fin disc; ray counts of the second dorsal, anal, and pectoral fins; scale counts of longitudinal and transverse rows; number and position of suborbital rows of the cephalic sensory systems; length and shape of nostrils (see Diagnosis for details, Tables 1 and 2 for comparisons).
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E2EFFD87DE0FB5BFCB1EEDF" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="526">
|
||
<taxonomicName id="4C420D865E2EFFD87DE0FB5BFDE8ECC3" box="[199,561,1160,1184]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
|
||
<emphasis id="B936AA175E2EFFD87DE0FB5BFDE8ECC3" box="[199,561,1160,1184]" italics="true" pageId="13" pageNumber="526">Coryogalops pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A205176C5E2EFFD87F1CFB54FD4EECC3" box="[571,663,1159,1184]" pageId="13" pageNumber="526" rank="species">
|
||
<emphasis id="B936AA175E2EFFD87F1CFB54FD4EECC3" bold="true" box="[571,663,1159,1184]" pageId="13" pageNumber="526">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
resembles
|
||
<taxonomicName id="4C420D865E2EFFD87E3CFB5BFC67ECC3" box="[795,958,1160,1184]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87E3CFB5BFC67ECC3" box="[795,958,1160,1184]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13796A805E2EFFD87EF6FB54FBCFECC3" box="[977,1046,1159,1184]" captionStart="FIGURE 5" captionStartId="14.[151,250,1681,1703]" captionTargetBox="[253,1333,195,1657]" captionTargetId="figure@14.[253,1333,195,1657]" captionTargetPageId="14" captionText="FIGURE 5. Coryogalops ocheticus. A: PMR VP 2476, male, 54.2 + 13.4 mm, off Suez, Gulf of Suez, Red Sea; B: SMF 13229, holotype of Cabillus anchialinae, male, 33.4 mm SL, Sinai coast near Ras Muhammad, Egypt, Red Sea; C: Coryogalops monospilus, BPBM 33235, holotype, 40.8 mm SL, Kuwait, Arabian Gulf. Photos by S. V. Bogorodsky (A & B), J. E. Randall (C)." httpUri="https://zenodo.org/record/230292/files/figure.png" pageId="13" pageNumber="526">Fig. 5</figureCitation>
|
||
C; known only from the
|
||
<typeStatus id="54F9C8A75E2EFFD8781DFB54FA45ECC3" box="[1338,1436,1159,1184]" pageId="13" pageNumber="526" type="holotype">holotype</typeStatus>
|
||
from the Arabian Gulf,
|
||
<collectingCountry id="F35536955E2EFFD87C8CFB7FFE26ECA6" box="[427,511,1196,1221]" name="Kuwait" pageId="13" pageNumber="526">Kuwait</collectingCountry>
|
||
) in body shape and coloration, but differs in having fewer pectoral-fin rays and fewer free tips, 16–17
|
||
<emphasis id="B936AA175E2EFFD87CBDFB02FE60EC8B" box="[410,441,1233,1256]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
19, and 2
|
||
<emphasis id="B936AA175E2EFFD87F13FB02FD8AEC8B" box="[564,595,1233,1256]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
3, respectively; fewer longitudinal scale series and transverse scale rows, 33–35
|
||
<emphasis id="B936AA175E2EFFD87DC3FB26FEDBED6F" box="[228,258,1269,1292]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
38 or 39, and 9
|
||
<emphasis id="B936AA175E2EFFD87C9EFB26FE0EED6F" box="[441,471,1269,1292]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
16, respectively (see Discussion for the explanation of the problem with the counts of the
|
||
<typeStatus id="54F9C8A75E2EFFD87DE7FAC4FEFBED53" box="[192,290,1303,1328]" pageId="13" pageNumber="526" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C420D865E2EFFD87C6FFACBFE30ED53" box="[328,489,1304,1328]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87C6FFACBFE30ED53" box="[328,489,1304,1328]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
); no scales on side of nape anterior to a line from origin of third spine of first dorsal fin to behind upper end of pectoral-fin base
|
||
<emphasis id="B936AA175E2EFFD87FA5FAEEFD78ED37" box="[642,673,1341,1364]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
small scales on side of nape ending above upper end of gill opening with a narrow single row of embedded scales reaching anteriorly nearly vertical above posterior margin of preopercle in
|
||
<taxonomicName id="4C420D865E2EFFD87C19FA57FE3DEDFF" box="[318,484,1412,1436]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87C19FA57FE3DEDFF" box="[318,484,1412,1436]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
; first dorsal fin with a nearly square dark maroon blotch on second and third membranes at base of fin, linked with first spine by a short dark band across first membrane vs. first dorsal fin with a black spot covers first two membranes, weakly extended downwards as brown oblique band at third membrane to base of fin; pectoral fins with two dark brown spots, each on upper and lower base of pectoral-fin rays
|
||
<emphasis id="B936AA175E2EFFD87832FA22FAEDEE6B" box="[1301,1332,1521,1544]" italics="true" pageId="13" pageNumber="526">vs.</emphasis>
|
||
only one black blotch visible on upper base of pectoral-fin rays in
|
||
<taxonomicName id="4C420D865E2EFFD87E34F9C7FC6BEE4F" box="[787,946,1556,1580]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87E34F9C7FC6BEE4F" box="[787,946,1556,1580]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
. Some morphometric values differ between the closest of two values of
|
||
<taxonomicName id="4C420D865E2EFFD87CFAF9EBFD16EE33" box="[477,719,1592,1616]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">
|
||
<emphasis id="B936AA175E2EFFD87CFAF9EBFD16EE33" box="[477,719,1592,1616]" italics="true" pageId="13" pageNumber="526">C. pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
and single value of
|
||
<taxonomicName id="4C420D865E2EFFD87E9AF9EBFB86EE33" box="[957,1119,1592,1616]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87E9AF9EBFB86EE33" box="[957,1119,1592,1616]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
by 1/10 or more (Table 1), however, to confirm the range of morphometric differences between two species larger samples of both sexes from both species are needed:
|
||
<taxonomicName id="4C420D865E2EFFD87C92F953FD8FEEFB" box="[437,598,1664,1688]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2EFFD87C92F953FD8FEEFB" box="[437,598,1664,1688]" italics="true" pageId="13" pageNumber="526">C. monospilus</emphasis>
|
||
</taxonomicName>
|
||
has larger eyes, shorter base of both dorsal fins, longer pelvic fin, longer dorsal-fin and pelvic-fin spines, and shorter upper jaw (Table 1).
|
||
</paragraph>
|
||
<paragraph id="8BFD76055E2EFFD87DE0F91BFA42EFBF" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="526">
|
||
<taxonomicName id="4C420D865E2EFFD87DE0F91BFDE8EE83" box="[199,561,1736,1760]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus" status="sp. nov.">
|
||
<emphasis id="B936AA175E2EFFD87DE0F91BFDE8EE83" box="[199,561,1736,1760]" italics="true" pageId="13" pageNumber="526">Coryogalops pseudomonospilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A205176C5E2EFFD87F1BF914FD40EE83" box="[572,665,1735,1760]" pageId="13" pageNumber="526" rank="species">
|
||
<emphasis id="B936AA175E2EFFD87F1BF914FD40EE83" bold="true" box="[572,665,1735,1760]" pageId="13" pageNumber="526">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="4C420D865E2EFFD87E5BF91BFBD2EE83" box="[892,1035,1736,1760]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2EFFD87E5BF91BFBD2EE83" box="[892,1035,1736,1760]" italics="true" pageId="13" pageNumber="526">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13796A805E2EFFD87939F914FBBDEE83" box="[1054,1124,1735,1760]" captionStart="FIGURE 5" captionStartId="14.[151,250,1681,1703]" captionTargetBox="[253,1333,195,1657]" captionTargetId="figure@14.[253,1333,195,1657]" captionTargetPageId="14" captionText="FIGURE 5. Coryogalops ocheticus. A: PMR VP 2476, male, 54.2 + 13.4 mm, off Suez, Gulf of Suez, Red Sea; B: SMF 13229, holotype of Cabillus anchialinae, male, 33.4 mm SL, Sinai coast near Ras Muhammad, Egypt, Red Sea; C: Coryogalops monospilus, BPBM 33235, holotype, 40.8 mm SL, Kuwait, Arabian Gulf. Photos by S. V. Bogorodsky (A & B), J. E. Randall (C)." httpUri="https://zenodo.org/record/230292/files/figure.png" pageId="13" pageNumber="526">Fig. 5</figureCitation>
|
||
A,
|
||
<tableCitation id="C6C043BE5E2EFFD879ADF914FB3AEE83" box="[1162,1251,1735,1760]" captionStart="TABLE 2" captionStartId="4.[136,217,539,560]" captionTargetBox="[136,1876,682,1445]" captionTargetPageId="4" captionText="TABLE 2. (Continued)" httpUri="http://table.plazi.org/id/DF3D268D5E27FFD17DAFFDC8FEB2EA53" pageId="13" pageNumber="526" tableUuid="DF3D268D5E27FFD17DAFFDC8FEB2EA53">Table 2</tableCitation>
|
||
), from which it differs in the following morphological and coloration characters: in males second to sixth spines of first dorsal fin reach spine of second fin, in females at least fourth spine reaches spine of second dorsal fin in
|
||
<emphasis id="B936AA175E2EFFD8798AF8C3FF6CEF2F" italics="true" pageId="13" pageNumber="526">
|
||
<taxonomicName id="4C420D865E2EFFD8798AF8C3FA45EF4B" box="[1197,1436,1808,1832]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
in males the appressed fourth spine of first dorsal fin may reach second dorsal fin, in females all spines do not reach second dorsal fin in
|
||
<taxonomicName id="4C420D865E2EFFD87CE7F88BFD92EF13" box="[448,587,1880,1904]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2EFFD87CE7F88BFD92EF13" box="[448,587,1880,1904]" italics="true" pageId="13" pageNumber="526">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; pelvic fins moderately emarginate (the shortest branches of fifth ray about 4/5 length of the longest branches of fourth ray), with pelvic frenum height in midline 1/3–2/5 length of pelvic spine in
|
||
<emphasis id="B936AA175E2EFFD87DD3F873FDD1EFDB" box="[244,520,1952,1976]" italics="true" pageId="13" pageNumber="526">
|
||
<taxonomicName id="4C420D865E2EFFD87DD3F873FE3AEFDB" box="[244,483,1952,1976]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
pelvic disc elliptical with rounded posterior edge, pelvic frenum height in midline about 1/2 length of pelvic spine in
|
||
<taxonomicName id="4C420D865E2EFFD87F10F817FD12EFBF" box="[567,715,1988,2012]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2EFFD87F10F817FD12EFBF" box="[567,715,1988,2012]" italics="true" pageId="13" pageNumber="526">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; longitudinal scale series
|
||
<quantity id="4CBADBE05E2EFFD87923F817FBB5EFBE" box="[1028,1132,1988,2013]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="13" pageNumber="526" unit="in" value="34.0" valueMax="35.0" valueMin="33.0">33–35 in</quantity>
|
||
<emphasis id="B936AA175E2EFFD87953F817FA42EFBF" box="[1140,1435,1988,2012]" italics="true" pageId="13" pageNumber="526">
|
||
<taxonomicName id="4C420D865E2EFFD87953F817FAAAEFBF" box="[1140,1395,1988,2012]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="526" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DF3D268D5E2DFFDB7DB0F942FF19EF60" httpUri="https://zenodo.org/record/230292/files/figure.png" pageId="14" pageNumber="527" targetBox="[253,1333,195,1657]" targetPageId="14">
|
||
<paragraph id="8BFD76055E2DFFDB7DB0F942FF19EF60" blockId="14.[151,1436,1681,1795]" pageId="14" pageNumber="527">
|
||
<emphasis id="B936AA175E2DFFDB7DB0F942FECBEEC4" bold="true" box="[151,274,1681,1703]" pageId="14" pageNumber="527">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C420D865E2DFFDB7C3FF942FDD9EEC4" box="[280,512,1681,1703]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2DFFDB7C3FF942FDD9EEC4" box="[280,512,1681,1703]" italics="true" pageId="14" pageNumber="527">Coryogalops ocheticus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B936AA175E2DFFDB7F2BF942FDFEEEC5" bold="true" box="[524,551,1681,1702]" pageId="14" pageNumber="527">A:</emphasis>
|
||
PMR VP2476, male, 54.2 + 13.4 mm, off Suez, Gulf of Suez, Red Sea;
|
||
<emphasis id="B936AA175E2DFFDB79D9F941FAC1EEC4" bold="true" box="[1278,1304,1682,1703]" pageId="14" pageNumber="527">B:</emphasis>
|
||
SMF 13229, holotype of
|
||
<taxonomicName id="4C420D865E2DFFDB7C3FF963FE36EEA5" box="[280,495,1712,1734]" class="Actinopterygii" family="Gobiidae" genus="Cabillus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="anchialinae">
|
||
<emphasis id="B936AA175E2DFFDB7C3FF963FE36EEA5" box="[280,495,1712,1734]" italics="true" pageId="14" pageNumber="527">Cabillus anchialinae</emphasis>
|
||
</taxonomicName>
|
||
, male, 33.4 mm SL, Sinai coast near Ras Muhammad, Egypt, Red Sea;
|
||
<emphasis id="B936AA175E2DFFDB79D3F963FAD6EEA5" bold="true" box="[1268,1295,1712,1734]" pageId="14" pageNumber="527">C:</emphasis>
|
||
<taxonomicName id="4C420D865E2DFFDB783EF963FED3EE86" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="monospilus">
|
||
<emphasis id="B936AA175E2DFFDB783EF963FED3EE86" italics="true" pageId="14" pageNumber="527">Coryogalops monospilus</emphasis>
|
||
</taxonomicName>
|
||
, BPBM 33235, holotype, 40.8 mm SL, Kuwait, Arabian Gulf. Photos by S.V. Bogorodsky (A & B), J.E. Randall (C).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD76055E2DFFDA7DB0F8E3FEB9EA5F" blockId="14.[151,1436,1840,2009]" lastBlockId="15.[151,1436,151,573]" lastPageId="15" lastPageNumber="528" pageId="14" pageNumber="527">
|
||
longitudinal scale series
|
||
<quantity id="4CBADBE05E2DFFDB7C99F8E3FDFEEF2A" box="[446,551,1840,1865]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.874" metricValueMin="6.604" pageId="14" pageNumber="527" unit="in" value="28.5" valueMax="31.0" valueMin="26.0">26–31 in</quantity>
|
||
<taxonomicName id="4C420D865E2DFFDB7F13F8E3FD1CEF2B" box="[564,709,1840,1864]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2DFFDB7F13F8E3FD1CEF2B" box="[564,709,1840,1864]" italics="true" pageId="14" pageNumber="527">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; anterior nostril tubular, without process from posterior rim, posterior nostril slightly raised in
|
||
<emphasis id="B936AA175E2DFFDB7F0BF886FC92EF0F" box="[556,843,1876,1900]" italics="true" pageId="14" pageNumber="527">
|
||
<taxonomicName id="4C420D865E2DFFDB7F0BF886FCF9EF0F" box="[556,800,1876,1900]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
anterior nostril short, tubular with oblique orifice bearing lappet or tentacle attaining length of tube, posterior nostril pore-like in
|
||
<taxonomicName id="4C420D865E2DFFDB797FF8ABFB30EFF3" box="[1112,1257,1912,1936]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2DFFDB797FF8ABFB30EFF3" box="[1112,1257,1912,1936]" italics="true" pageId="14" pageNumber="527">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; head laterally pigmented except for pale band from lower margin of eye to rear part of mouth, including the angle of jaw in
|
||
<emphasis id="B936AA175E2DFFDB78A5F84EFE54EFBB" italics="true" pageId="14" pageNumber="527">
|
||
<taxonomicName id="4C420D865E2DFFDB78A5F84EFEBFEFBB" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="527" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
head finely pigmented, with dark suborbital bar at angle of jaw and oblique posterior band from eye margin, with closely mottled cheek and opercle in
|
||
<taxonomicName id="4C420D865E2CFFDA7E79FF4BFC35E8D3" box="[862,1004,152,176]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7E79FF4BFC35E8D3" box="[862,1004,152,176]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; chest pigmented only in male in
|
||
<emphasis id="B936AA175E2CFFDA78A5FF4BFE54E8B7" italics="true" pageId="15" pageNumber="528">
|
||
<taxonomicName id="4C420D865E2CFFDA78A5FF4BFEBFE8B7" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
chest and branchiostegal membrane with melanophores in both sexes in
|
||
<taxonomicName id="4C420D865E2CFFDA79F3FF6FFAB8E8B7" box="[1236,1377,188,212]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA79F3FF6FFAB8E8B7" box="[1236,1377,188,212]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; two marks present in the upper and lower base of pectoral-fin rays, more distinctive in male in
|
||
<emphasis id="B936AA175E2CFFDA79AFFF33FA45E89B" box="[1160,1436,224,248]" italics="true" pageId="15" pageNumber="528">
|
||
<taxonomicName id="4C420D865E2CFFDA79AFFF33FAAEE89B" box="[1160,1399,224,248]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
oblique dark bar across proximal part of upper five pectoral-rays in
|
||
<taxonomicName id="4C420D865E2CFFDA7EB3FED7FBC6E97F" box="[916,1055,260,284]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7EB3FED7FBC6E97F" box="[916,1055,260,284]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; pelvic fin moderately pigmented only in male, unpigmented in female in
|
||
<emphasis id="B936AA175E2CFFDA7F47FEFBFCA1E923" box="[608,888,296,320]" italics="true" pageId="15" pageNumber="528">
|
||
<taxonomicName id="4C420D865E2CFFDA7F47FEFBFC88E923" box="[608,849,296,320]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
pelvic disc and anterior membrane dark in both sexes in
|
||
<taxonomicName id="4C420D865E2CFFDA7DDFFE9FFE5AE907" box="[248,387,332,356]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7DDFFE9FFE5AE907" box="[248,387,332,356]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
; first dorsal fin with dark brown spot anteriorly in basal part in
|
||
<emphasis id="B936AA175E2CFFDA7975FE9FFABEE907" box="[1106,1383,332,356]" italics="true" pageId="15" pageNumber="528">
|
||
<taxonomicName id="4C420D865E2CFFDA7975FE9FFA9BE907" box="[1106,1346,332,356]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="pseudomonospilus">C. pseudomonospilus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
first dorsal fin with indistinct oblique dark band (no spots present) in
|
||
<taxonomicName id="4C420D865E2CFFDA7E50FEA3FBDBE9EB" box="[887,1026,368,392]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7E50FEA3FBDBE9EB" box="[887,1026,368,392]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
. During our study, the
|
||
<typeStatus id="54F9C8A75E2CFFDA782EFEBCFAB2E9EB" box="[1289,1387,367,392]" pageId="15" pageNumber="528" type="holotype">holotype</typeStatus>
|
||
and three
|
||
<typeStatus id="54F9C8A75E2CFFDA7DF2FE46FE99E9CE" box="[213,320,405,429]" pageId="15" pageNumber="528" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C420D865E2CFFDA7C42FE47FD96E9CF" box="[357,591,404,428]" class="Actinopterygii" family="Gobiidae" genus="Cabillus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="anchialinae">
|
||
<emphasis id="B936AA175E2CFFDA7C42FE47FD96E9CF" box="[357,591,404,428]" italics="true" pageId="15" pageNumber="528">Cabillus anchialinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13796A805E2CFFDA7F79FE47FD78E9CE" box="[606,673,404,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1681,1703]" captionTargetBox="[253,1333,195,1657]" captionTargetId="figure@14.[253,1333,195,1657]" captionTargetPageId="14" captionText="FIGURE 5. Coryogalops ocheticus. A: PMR VP 2476, male, 54.2 + 13.4 mm, off Suez, Gulf of Suez, Red Sea; B: SMF 13229, holotype of Cabillus anchialinae, male, 33.4 mm SL, Sinai coast near Ras Muhammad, Egypt, Red Sea; C: Coryogalops monospilus, BPBM 33235, holotype, 40.8 mm SL, Kuwait, Arabian Gulf. Photos by S. V. Bogorodsky (A & B), J. E. Randall (C)." httpUri="https://zenodo.org/record/230292/files/figure.png" pageId="15" pageNumber="528">Fig. 5</figureCitation>
|
||
B) were re-examined in comparison with our material examined and the redescription of
|
||
<taxonomicName id="4C420D865E2CFFDA7CA6FE6BFDD6E9B3" box="[385,527,440,464]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7CA6FE6BFDD6E9B3" box="[385,527,440,464]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
given in
|
||
<bibRefCitation id="EFD30BF45E2CFFDA7FA5FE64FCC7E9B3" author="Miller" box="[642,798,439,464]" pageId="15" pageNumber="528" refString="Miller, P. J. (1978) The systematic position and origin of Gobius ocheticus Norman, 1927, from the Suez Canal. Zoological Journal of the Linnean Society, 62 (1), 39 - 58. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1978. tb 00522. x" type="journal article" year="1978">Miller (1978)</bibRefCitation>
|
||
, and confirmed that it is conspecific with
|
||
<taxonomicName id="4C420D865E2CFFDA7829FE6BFA45E9B3" box="[1294,1436,440,464]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="ocheticus">
|
||
<emphasis id="B936AA175E2CFFDA7829FE6BFA45E9B3" box="[1294,1436,440,464]" italics="true" pageId="15" pageNumber="528">C. ocheticus</emphasis>
|
||
</taxonomicName>
|
||
based on identical morphology. Another species of the genus sympatric in the Red Sea with both of the new species is
|
||
<taxonomicName id="4C420D865E2CFFDA7D95FDD3FE98EA7B" box="[178,321,512,536]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="528" phylum="Chordata" rank="species" species="anomolus">
|
||
<emphasis id="B936AA175E2CFFDA7D95FDD3FE98EA7B" box="[178,321,512,536]" italics="true" pageId="15" pageNumber="528">C. anomolus</emphasis>
|
||
</taxonomicName>
|
||
but it can be easily distinguished by its having separate pelvic fins and different coloration (
|
||
<figureCitation id="13796A805E2CFFDA7843FE2CFF7DEA5E" captionStart="FIGURE 6" captionStartId="15.[151,250,1938,1960]" captionTargetBox="[217,1369,661,1916]" captionTargetId="figure@15.[217,1369,661,1917]" captionTargetPageId="15" captionText="FIGURE 6. Coryogalops anomolus. A: PMR VP 3052, male, 24.3 + 6.2 mm, Sharm el Moya, southern Sinai, Red Sea; B: uncatalogued, Brayka Bay, Marsa Alam, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/230293/files/figure.png" pageId="15" pageNumber="528">Figs. 6</figureCitation>
|
||
A & B,
|
||
<tableCitation id="C6C043BE5E2CFFDA7DDAFDF7FE8AEA5F" box="[253,339,548,573]" captionStart="TABLE 2" captionStartId="4.[136,217,539,560]" captionTargetBox="[136,1876,682,1445]" captionTargetPageId="4" captionText="TABLE 2. (Continued)" httpUri="http://table.plazi.org/id/DF3D268D5E27FFD17DAFFDC8FEB2EA53" pageId="15" pageNumber="528" tableUuid="DF3D268D5E27FFD17DAFFDC8FEB2EA53">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |