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<document id="57E66A0C1F6D4EE3FF9A77BDE77FB7E3" ID-DOI="10.5281/zenodo.199052" ID-GBIF-Dataset="a07f4b3c-e602-4b4e-95dc-87fef597d912" ID-ISSN="1175-5326" ID-Zenodo-Dep="199052" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460321171440" checkinUser="plazi" docAuthor="Park, Jong-Seok, Carlton, Christopher E. &amp; Ferro, Michael L." docDate="2010" docId="03E2BD70FFF4806EFF5C2A2DFC4E4B8E" docLanguage="en" docName="zt02662p027.pdf" docOrigin="Zootaxa 2662" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptusa (Dysleptusa) gimmeli Park &amp; Carlton, new species" docType="treatment" docVersion="11" lastPageNumber="16" masterDocId="FFDBC508FFF98061FFCB282AFF954A52" masterDocTitle="Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="14" updateTime="1698256666968" updateUser="plazi">
<mods:mods id="5635A1AAED1B95F3895D6B1E30DE8F95" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C21DD9461C8A9E51A7ED5F660ECB739">
<mods:title id="4CD9BE7CBF78605FE15B2579B99D4F95">Diversity and taxonomic review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="AC7AA54F4FF54C7D7449055FD683D55A" type="personal">
<mods:role id="26B6C6379AA9F894396C31E058463170">
<mods:roleTerm id="297278C4F35E594D371BC65F6F451A4D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E26A76508388638FB751D41B0AC97002">Park, Jong-Seok</mods:namePart>
</mods:name>
<mods:name id="9F3052A52DC25E97930110A8FB7E6282" type="personal">
<mods:role id="F43B6A913ECF8276AFC571979B701D7D">
<mods:roleTerm id="045724AF56C8DB5A4382DA2DC7D0B667">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53AF786FAA5C982D9690BD635B421E7D">Carlton, Christopher E.</mods:namePart>
</mods:name>
<mods:name id="347DC017E78EBF298ECF29376115BA8E" type="personal">
<mods:role id="497631CDC1BEB98F5BF252F5B54AFAEF">
<mods:roleTerm id="C7978013A575D108EB249D5D6E232612">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9737BC0E46E68F4519EBEC0D08E8463B">Ferro, Michael L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A186A7472FC26246E9E140B13772C657">text</mods:typeOfResource>
<mods:relatedItem id="844AC27562106454580E8D2642CDF0E7" type="host">
<mods:titleInfo id="6A8902ECC1ED60C07A3C636D96FEC202">
<mods:title id="A910B12D76BDF72639DFF03C004D5B57">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4963E532A6CD2EA51473198467957105">
<mods:date id="ABF6AAC10F85D013AA1DCD599408DF05">2010</mods:date>
<mods:detail id="8ECEE8CABDE02ACD659126CC2FB436B1" type="volume">
<mods:number id="9F8E3EDCBDDEC2D55AD3BA98DC8A2C52">2662</mods:number>
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<mods:extent id="31CCF23F4EF8F657EDDBB4F9979CD519" unit="page">
<mods:start id="E7052DF5B193976DC2ACFE5A9BDDEDF4">1</mods:start>
<mods:end id="FD1CC8927C2932E2FD410C8E2347719E">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4B2EBFDDB3A9BF4B3CD61130514226BB">journal article</mods:classification>
<mods:identifier id="007C50AC894A13434D7A651567154B2E" type="DOI">10.5281/zenodo.199052</mods:identifier>
<mods:identifier id="3FA14089257AF1D49EA24953FBB5369D" type="GBIF-Dataset">a07f4b3c-e602-4b4e-95dc-87fef597d912</mods:identifier>
<mods:identifier id="1CADAF15982096375F242F2094333272" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E8644E13435769E8844C836CCD5F657C" type="Zenodo-Dep">199052</mods:identifier>
</mods:mods>
<treatment id="03E2BD70FFF4806EFF5C2A2DFC4E4B8E" ID-DOI="http://doi.org/10.5281/zenodo.5020591" ID-GBIF-Taxon="119402523" ID-Zenodo-Dep="5020591" LSID="urn:lsid:zoobank.org:act:3CC6148E-02A7-4805-92F6-7340BD568F88" httpUri="http://treatment.plazi.org/id/03E2BD70FFF4806EFF5C2A2DFC4E4B8E" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="C3515FEDFFF4806CFF5C2A2DFE364816" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF40C66FFF4806CFF5C2A2DFCFE4873" blockId="13.[151,875,519,580]" box="[151,875,519,545]" pageId="13" pageNumber="14">
<heading id="D0BCBB0AFFF4806CFF5C2A2DFCFE4873" bold="true" box="[151,875,519,545]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B93FD074FFF4806CFF5C2A2DFCFE4873" bold="true" box="[151,875,519,545]" pageId="13" pageNumber="14">
<taxonomicName id="4C4B77E5FFF4806CFF5C2A2DFD5B4873" ID-CoL="6PP8F" authority="Park &amp; Carlton" authorityName="Park &amp; Carlton" box="[151,718,519,545]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gimmeli" status="sp. nov." subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFF5C2A2DFF624873" bold="true" box="[151,247,519,545]" italics="true" pageId="13" pageNumber="14">Leptusa</emphasis>
(
<emphasis id="B93FD074FFF4806CFEC32A2DFE194873" bold="true" box="[264,396,519,545]" italics="true" pageId="13" pageNumber="14">Dysleptusa</emphasis>
)
<emphasis id="B93FD074FFF4806CFE562A2DFE6A4873" bold="true" box="[413,511,519,545]" italics="true" pageId="13" pageNumber="14">gimmeli</emphasis>
Park &amp; Carlton
</taxonomicName>
,
<taxonomicNameLabel id="A20C6D0FFFF4806CFD102A2DFCFE4873" box="[731,875,519,545]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF40C66FFF4806CFF5C2A00FE364816" blockId="13.[151,875,519,580]" box="[151,419,554,580]" pageId="13" pageNumber="14">
(
<figureCitation id="137010E3FFF4806CFF542A00FF7B4816" box="[159,238,554,580]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="13" pageNumber="14">
Figs.
<quantity id="4CB3A183FFF4806CFF292A00FF7B4816" box="[226,238,554,580]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="13" pageNumber="14" unit="g" value="1.0">1</quantity>
</figureCitation>
g, 7a
<emphasis id="B93FD074FFF4806CFEE22A00FEAD4811" box="[297,312,554,579]" italics="true" pageId="13" pageNumber="14">–</emphasis>
l, Map 1)
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF4806CFF5C2A5FFE494E56" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF40C66FFF4806CFF5C2A5FFE494E56" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF5C2A5FFD9248DD" bold="true" box="[151,519,629,655]" pageId="13" pageNumber="14">
<typeStatus id="54F0B2C4FFF4806CFF5C2A5FFE9848DD" box="[151,269,629,655]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F35C4CF6FFF4806CFED12A5FFEE548DD" box="[282,368,629,655]" name="United States of America" pageId="13" pageNumber="14">U.S.A.</collectingCountry>
: Tennessee:
</emphasis>
ɗ, “
<collectingCountry id="F35C4CF6FFF4806CFDF52A5FFD1548DD" box="[574,640,629,655]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFC802A5FFC2448DD" LSID="urn:lsid:biocol.org:col:13136" box="[843,945,629,655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFBB22A5FFA8848DD" box="[1145,1309,629,655]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFAEE2A5FFF4748E6" pageId="13" pageNumber="14" value="2006-10-05">5 October 2006</date>
<geoCoordinate id="EE7F6AA1FFF4806CFF102AB0FEE848E6" box="[219,381,666,692]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP3 Litter 3 of 3 –M Gimmel”, “
<typeStatus id="54F0B2C4FFF4806CFCD92AB0FC2048E6" box="[786,949,666,692]" pageId="13" pageNumber="14" type="holotype">
<collectionCode id="ED5A94A3FFF4806CFCD92AB0FC2048E6" box="[786,949,666,692]" pageId="13" pageNumber="14">HOLOTYPE</collectionCode>
</typeStatus>
,
<taxonomicName id="4C4B77E5FFF4806CFC092AB0FAF548E6" authority="Park and Carlton" authorityName="Park and Carlton" box="[962,1376,666,692]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gimmeli">
<emphasis id="B93FD074FFF4806CFC092AB0FB1D48E1" box="[962,1160,666,691]" italics="true" pageId="13" pageNumber="14">Leptusa gimmeli</emphasis>
Park and Carlton
</taxonomicName>
, des. 2010”.
<emphasis id="B93FD074FFF4806CFF3F2A95FD60488B" bold="true" box="[244,757,703,729]" pageId="13" pageNumber="14">
<typeStatus id="54F0B2C4FFF4806CFF3F2A95FEEF488B" box="[244,378,703,729]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
(n=107).
<collectingCountry id="F35C4CF6FFF4806CFE372A95FDC2488B" box="[508,599,703,729]" name="United States of America" pageId="13" pageNumber="14">U.S.A.</collectingCountry>
: Tennessee:
</emphasis>
13ɗɗ (2ɗɗ slide mounted), 15ΨΨ with same data as
<typeStatus id="54F0B2C4FFF4806CFF5C2ACFFF6B48AD" box="[151,254,741,767]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED5A94A3FFF4806CFEC72ACFFEF148AD" LSID="urn:lsid:biocol.org:col:33726" box="[268,356,741,767]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="13" pageNumber="14">LSAM</collectionCode>
); 4ɗɗ, 10ΨΨ, “
<collectingCountry id="F35C4CF6FFF4806CFDD22ACFFDCF48AD" box="[537,602,741,767]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFCD72ACFFC1448AD" LSID="urn:lsid:biocol.org:col:13136" box="[796,897,741,767]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFB8A2ACFFB7448AD" box="[1089,1249,741,767]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFB2C2ACFFA0948AD" box="[1255,1436,741,767]" pageId="13" pageNumber="14" value="2006-10-05">5 October 2006</date>
<geoCoordinate id="EE7F6AA1FFF4806CFF5C2B20FEA34976" box="[151,310,778,804]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP3 Litter 1 of 3 –M Gimmel” (
<collectionCode id="ED5A94A3FFF4806CFD7E2B20FC984976" LSID="urn:lsid:biocol.org:col:33726" box="[693,781,778,804]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="13" pageNumber="14">LSAM</collectionCode>
); 11ɗɗ, 15ΨΨ (3ΨΨ slide mounted), “
<collectingCountry id="F35C4CF6FFF4806CFB052B20FA9A4976" box="[1230,1295,778,804]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFF012B1AFEA74918" LSID="urn:lsid:biocol.org:col:13136" box="[202,306,816,842]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFE342B1AFD304918" box="[511,677,816,842]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFD652B1AFCF84918" box="[686,877,816,842]" pageId="13" pageNumber="14" value="2006-10-05">5 October 2006</date>
<geoCoordinate id="EE7F6AA1FFF4806CFCBE2B1AFB8F4918" box="[885,1050,816,842]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP3 Litter 2 of 3 –M Gimmel” (
<collectionCode id="ED5A94A3FFF4806CFF542B7FFE87493D" LSID="urn:lsid:biocol.org:col:13136" box="[159,274,853,879]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
); 6ɗɗ, 7ΨΨ, “
<collectingCountry id="F35C4CF6FFF4806CFE022B7FFD9B493D" box="[457,526,853,879]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFD2E2B7FFCDA493D" LSID="urn:lsid:biocol.org:col:13136" box="[741,847,853,879]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFBEE2B7FFB47493D" box="[1061,1234,853,879]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFB102B7FFA08493D" box="[1243,1437,853,879]" pageId="13" pageNumber="14" value="2007-03-31">31 March 2007</date>
<geoCoordinate id="EE7F6AA1FFF4806CFF5C2B50FEA349C6" box="[151,310,890,916]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP3 Litter 2 of 3 –M Gimmel”; 8ɗɗ, 17ΨΨ (Ψ slide mounted), “
<collectingCountry id="F35C4CF6FFF4806CFBFA2B50FBE749C6" box="[1073,1138,890,916]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFAFC2B50FA0949C6" LSID="urn:lsid:biocol.org:col:13136" box="[1335,1436,890,916]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFE962B8AFD9049E8" box="[349,517,928,954]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFDC62B8AFD5C49E8" box="[525,713,928,954]" pageId="13" pageNumber="14" value="2007-03-31">31 March 2007</date>
<geoCoordinate id="EE7F6AA1FFF4806CFD192B8AFCED49E8" box="[722,888,928,954]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP3 Litter 3 of 3 –M Gimmel” (
<collectionCode id="ED5A94A3FFF4806CFAC42B8AFAFF49E8" LSID="urn:lsid:biocol.org:col:33726" box="[1295,1386,928,954]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="13" pageNumber="14">LSAM</collectionCode>
); Ψ, “
<collectingCountry id="F35C4CF6FFF4806CFF692BEFFF76498D" box="[162,227,965,991]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
: TN: Cocke Co.
<collectionCode id="ED5A94A3FFF4806CFE612BEFFD9A498D" LSID="urn:lsid:biocol.org:col:13136" box="[426,527,965,991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="13" pageNumber="14">GSMNP</collectionCode>
Albright Grove
<geoCoordinate id="EE7F6AA1FFF4806CFD1F2BEFFCE0498D" box="[724,885,965,991]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="35.736217">N 35°44.173’</geoCoordinate>
<date id="FFF52AA6FFF4806CFCB72BEFFBAB498D" box="[892,1086,965,991]" pageId="13" pageNumber="14" value="1931-04-12" valueMax="1931-05-18" valueMin="1931-04-12">31 12 IV – 18 V</date>
<geoCoordinate id="EE7F6AA1FFF4806CFB8C2BEFFB73498D" box="[1095,1254,965,991]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-83.27745">W83°16.647’</geoCoordinate>
SP34A –CWD Rear 1 –M Ferro” (
<collectionCode id="ED5A94A3FFF4806CFEBC2BC0FE5A4E56" LSID="urn:lsid:biocol.org:col:33726" box="[375,463,1002,1028]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="13" pageNumber="14">LSAM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF4806CFF0D2C25FB4D4EB6" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BF40C66FFF4806CFF0D2C25FB4D4EB6" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF0D2C25FED34E7B" bold="true" box="[198,326,1039,1065]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
<taxonomicName id="4C4B77E5FFF4806CFE9A2C3AFD2C4E7B" box="[337,697,1040,1066]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gimmeli" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFE9A2C3AFE3B4E7B" box="[337,430,1040,1065]" italics="true" pageId="13" pageNumber="14">Leptusa</emphasis>
(
<emphasis id="B93FD074FFF4806CFE082C3AFDD04E7B" box="[451,581,1040,1065]" italics="true" pageId="13" pageNumber="14">Dysleptusa</emphasis>
)
<emphasis id="B93FD074FFF4806CFD912C3AFD2C4E7B" box="[602,697,1040,1065]" italics="true" pageId="13" pageNumber="14">gimmeli</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4B77E5FFF4806CFC952C3AFBDF4E7B" box="[862,1098,1040,1066]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="carolinensis" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFC952C3AFCFB4E7B" box="[862,878,1040,1065]" italics="true" pageId="13" pageNumber="14">L</emphasis>
. (
<emphasis id="B93FD074FFF4806CFC402C3AFC354E7B" box="[907,928,1040,1065]" italics="true" pageId="13" pageNumber="14">D</emphasis>
.)
<emphasis id="B93FD074FFF4806CFC772C3AFBDF4E7B" box="[956,1098,1040,1065]" italics="true" pageId="13" pageNumber="14">carolinensis</emphasis>
</taxonomicName>
by its smaller body length, shape of the spermatheca (
<figureCitation id="137010E3FFF4806CFE162C1FFDB24E1D" box="[477,551,1077,1103]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
l), short elytra (apparently as long as wide,
<figureCitation id="137010E3FFF4806CFBF02C1FFB154E1D" box="[1083,1152,1077,1103]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
f), and absence of hind wings. This species differs from
<taxonomicName id="4C4B77E5FFF4806CFD892C71FCD24E26" box="[578,839,1114,1140]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="smetanaiella" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFD892C71FDC74E26" box="[578,594,1115,1140]" italics="true" pageId="13" pageNumber="14">L</emphasis>
. (
<emphasis id="B93FD074FFF4806CFDBB2C71FD104E26" box="[624,645,1115,1140]" italics="true" pageId="13" pageNumber="14">D</emphasis>
.)
<emphasis id="B93FD074FFF4806CFD682C71FCD24E26" box="[675,839,1115,1140]" italics="true" pageId="13" pageNumber="14">smetanaiella</emphasis>
</taxonomicName>
by its shorter body length, color, subquadrate antennomere 4 (
<figureCitation id="137010E3FFF4806CFE912CAAFE364EC8" box="[346,419,1152,1178]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
a), and short elytra (apparently as long as wide,
<figureCitation id="137010E3FFF4806CFC212CAAFBBB4EC8" box="[1002,1070,1152,1178]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
f).
<taxonomicName id="4C4B77E5FFF4806CFB9E2CAAFAC24ECB" box="[1109,1367,1152,1178]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gimmeli" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFB9E2CAAFB214ECB" box="[1109,1204,1152,1177]" italics="true" pageId="13" pageNumber="14">Leptusa</emphasis>
(
<emphasis id="B93FD074FFF4806CFB0D2CAAFB764ECB" box="[1222,1251,1152,1177]" italics="true" pageId="13" pageNumber="14">D.</emphasis>
)
<emphasis id="B93FD074FFF4806CFB3D2CAAFAC24ECB" box="[1270,1367,1152,1177]" italics="true" pageId="13" pageNumber="14">gimmeli</emphasis>
</taxonomicName>
has a median lobe shaped similar to that of
<taxonomicName id="4C4B77E5FFF4806CFD9B2C8FFCA04EEC" box="[592,821,1189,1215]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="carolinensis" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFD9B2C8FFDFD4EEC" box="[592,616,1189,1214]" italics="true" pageId="13" pageNumber="14">L.</emphasis>
(
<emphasis id="B93FD074FFF4806CFDB22C8FFD1B4EEC" box="[633,654,1189,1214]" italics="true" pageId="13" pageNumber="14">D</emphasis>
.)
<emphasis id="B93FD074FFF4806CFD6C2C8FFCA04EEC" box="[679,821,1189,1214]" italics="true" pageId="13" pageNumber="14">carolinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4B77E5FFF4806CFCA52C8FFBCF4EEC" box="[878,1114,1189,1215]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="smetanaiella" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF4806CFCA52C8FFCEB4EEC" box="[878,894,1189,1214]" italics="true" pageId="13" pageNumber="14">L</emphasis>
. (
<emphasis id="B93FD074FFF4806CFC5D2C8FFC3E4EEC" box="[918,939,1189,1214]" italics="true" pageId="13" pageNumber="14">D</emphasis>
.)
<emphasis id="B93FD074FFF4806CFC0F2C8FFBCF4EEC" box="[964,1114,1189,1214]" italics="true" pageId="13" pageNumber="14">smetanaiella</emphasis>
</taxonomicName>
, but it can be distinguished from those species by the shape of the bulbus: the anterior parts of the median lobe (
<figureCitation id="137010E3FFF4806CFBB82CE0FB2C4EB6" box="[1139,1209,1226,1252]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
k).
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF4806EFF0D2CC5FB294AE0" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="8BF40C66FFF4806CFF0D2CC5FB784F28" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF0D2CC5FECA4F5B" bold="true" box="[198,351,1263,1289]" pageId="13" pageNumber="14">Description.</emphasis>
Length
<quantity id="4CB3A183FFF4806CFE072CDAFDF14F58" box="[460,612,1264,1290]" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="13" pageNumber="14" unit="mm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">
1.9
<emphasis id="B93FD074FFF4806CFE3A2CDAFD954F5B" box="[497,512,1264,1289]" italics="true" pageId="13" pageNumber="14">–</emphasis>
2.3 mm
</quantity>
. Body brown, glossy; head and abdominal tergites V
<emphasis id="B93FD074FFF4806CFB342CDAFA9B4F5B" box="[1279,1294,1264,1289]" italics="true" pageId="13" pageNumber="14">–</emphasis>
VI blackish brown; antennomeres 1
<emphasis id="B93FD074FFF4806CFE662D3FFE2E4F7C" box="[429,443,1301,1326]" italics="true" pageId="13" pageNumber="14">–</emphasis>
3, 11, mouthparts, legs, and abdominal tergite VIII yellowish brown (
<figureCitation id="137010E3FFF4806CFB3F2D3FFAA24F7D" box="[1268,1335,1301,1327]" captionStart="FIGURE 1" captionStartId="3.[151,255,1170,1194]" captionTargetBox="[153,1432,194,1147]" captionTargetId="figure@3.[151,1436,194,1149]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Leptusa. a: Leptusa (Adoxopisalia) opaca Casey; b: L. (A.) pseudosmokyiensis sp. nov.; c: L. (A.) smokyiensis Pace; d: L. (A.) stocksi sp. nov.; e: L. (Dysleptusa) carolinensis Pace; f: L. (D.) gimmeli sp. nov.; g: L. (D.) pusio (Casey); h: L. (Eucryptusa) brevicollis Casey; i: L. (E.) ferroi sp. nov.; j: L. (Ulitusa) cribratula (Casey). Scale bars = 1 mm." httpUri="https://zenodo.org/record/199053/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
f). Head pubescent and with microsculpture. Antennomeres 1
<emphasis id="B93FD074FFF4806CFCD92D11FCB44F06" box="[786,801,1339,1364]" italics="true" pageId="13" pageNumber="14">–</emphasis>
3 elongate, 4 subquadrate, 5
<emphasis id="B93FD074FFF4806CFBBE2D11FB114F06" box="[1141,1156,1339,1364]" italics="true" pageId="13" pageNumber="14">–</emphasis>
10 transverse (
<figureCitation id="137010E3FFF4806CFAFE2D10FAEB4F06" box="[1333,1406,1338,1364]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
a). Carina on each side of ventral surface of head incomplete, fading before attaining gular suture.
</paragraph>
<paragraph id="8BF40C66FFF4806CFF0D2DAFFAC54C5D" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF0D2DAFFEDA4FCC" box="[198,335,1413,1438]" italics="true" pageId="13" pageNumber="14">Mouthparts</emphasis>
. Labrum transverse, bearing 3 pairs of short setae, and 7 pairs of long setae, a-seta, b-seta, and pores (
<figureCitation id="137010E3FFF4806CFF2C2D80FEBA4F96" box="[231,303,1450,1476]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
b). Labium bearing two distal setae in a longitudinal row in median area; a pair of setal pores, 2 pairs of real pores and several pseudopores present in lateral area (
<figureCitation id="137010E3FFF4806CFC552DFAFC704FB8" box="[926,997,1488,1514]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
c). Labial palpi bearing 12 setae (a
<emphasis id="B93FD074FFF4806CFABC2DFAFA134FBB" box="[1399,1414,1488,1513]" italics="true" pageId="13" pageNumber="14">–</emphasis>
h, α
<emphasis id="B93FD074FFF4806CFF6D2DDFFF204C5C" box="[166,181,1525,1550]" italics="true" pageId="13" pageNumber="14">–</emphasis>
δ) (
<figureCitation id="137010E3FFF4806CFF172DDFFEB64C5D" box="[220,291,1525,1551]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
d). Mentum trapezoidal, bearing 4 pairs of main setae (b, u, v, w) and many pores (
<figureCitation id="137010E3FFF4806CFB272DDFFAA64C5D" box="[1260,1331,1525,1551]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
e).
</paragraph>
<paragraph id="8BF40C66FFF4806CFF0D2E31FC7A4CF6" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF0D2E31FEB64C66" box="[198,291,1563,1588]" italics="true" pageId="13" pageNumber="14">Thorax.</emphasis>
Pronotum wider than head, approximately 1.44 times wider than long, pubescent and with microsculpture. Mesoventrite with distinct reticulate microsculpture. Elytra as wide as pronotum, approximately 1.2 times longer than wide, approximately 1.06 times longer than pronotum, latero-posterior margin emarginate, and setigerous punctures present.
<collectingCountry id="F35C4CF6FFF4806CFCC62EA0FCDD4CF6" box="[781,840,1674,1700]" name="India" pageId="13" pageNumber="14">Hind</collectingCountry>
wings absent.
</paragraph>
<paragraph id="8BF40C66FFF4806CFF0D2E9AFDE24DA6" blockId="13.[151,1437,628,2036]" pageId="13" pageNumber="14">
<emphasis id="B93FD074FFF4806CFF0D2E9AFEA04C9B" box="[198,309,1712,1737]" italics="true" pageId="13" pageNumber="14">Abdomen</emphasis>
. Abdomen wider than elytra, widest at abdominal tergites V
<emphasis id="B93FD074FFF4806CFB822E9AFBC24C9B" box="[1097,1111,1712,1737]" italics="true" pageId="13" pageNumber="14">–</emphasis>
VI. Abdominal tergite VI approximately 1.88 times wider than long. Male abdominal tergites VII
<emphasis id="B93FD074FFF4806CFC302EFFFB9F4CBC" box="[1019,1034,1749,1774]" italics="true" pageId="13" pageNumber="14">–</emphasis>
VIII with single median elongate tubercle (
<figureCitation id="137010E3FFF4806CFEC72ED0FEC24D46" box="[268,343,1786,1812]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
f); tergite VIII bearing 4 pairs of main setae, and many additional short setae and pores; posterior margin with several tubercles, and shallowly emarginate (
<figureCitation id="137010E3FFF4806CFC602F0AFC644D68" box="[939,1009,1824,1850]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
f). Male abdominal sternite VII with many pores in anterior one-third; sternite VIII with posterior margin weakly produced, bearing 7 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFF4806CFD362F40FCDD4DD6" box="[765,840,1898,1924]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">
Fig.
<quantity id="4CB3A183FFF4806CFCFC2F40FCDD4DD6" box="[823,840,1898,1924]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="13" pageNumber="14" unit="g" value="7.0">7</quantity>
</figureCitation>
g). Female abdominal tergite VIII with posterior margin shallowly emarginate, bearing 4 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFF4806CFAA72FBAFF324D9D" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
h). Female abdominal sternite VIII with posterior margin rounded, bearing 6 pairs of main setae, and many additional short setae and pores (
<figureCitation id="137010E3FFF4806CFDD32FF0FDF54DA6" box="[536,608,2010,2036]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="13" pageNumber="14">
Fig.
<date id="FFF52AA6FFF4806CFD842FF0FDF54DA6" box="[591,608,2010,2036]" pageId="13" pageNumber="14">7</date>
</figureCitation>
i).
</paragraph>
<caption id="DF345CEEFFF7806FFF5C2FA2FBC94DB2" httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="14" pageNumber="15" targetBox="[165,1428,199,1907]" targetPageId="14">
<paragraph id="8BF40C66FFF7806FFF5C2FA2FBC94DB2" blockId="14.[151,1436,1928,2016]" pageId="14" pageNumber="15">
<emphasis id="B93FD074FFF7806FFF5C2FA2FE8A4DF2" bold="true" box="[151,287,1928,1952]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
Details of
<taxonomicName id="4C4B77E5FFF7806FFE572FA3FD794DF2" box="[412,748,1929,1952]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gimmeli" status="sp. nov." subGenus="Dysleptusa">
<emphasis id="B93FD074FFF7806FFE572FA3FE614DF2" box="[412,500,1929,1952]" italics="true" pageId="14" pageNumber="15">Leptusa</emphasis>
(
<emphasis id="B93FD074FFF7806FFDCE2FA3FD144DF2" box="[517,641,1929,1952]" italics="true" pageId="14" pageNumber="15">Dysleptusa</emphasis>
)
<emphasis id="B93FD074FFF7806FFD592FA3FD794DF2" box="[658,748,1929,1952]" italics="true" pageId="14" pageNumber="15">gimmeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C6D0FFFF7806FFD382FA2FCD84DF2" box="[755,845,1928,1952]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B93FD074FFF7806FFD382FA2FCD84DF2" bold="true" box="[755,845,1928,1952]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF40C66FFF6806EFF0D28B2FB294AE0" blockId="15.[151,1437,152,476]" box="[198,1212,152,178]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D28B2FEA04AE3" box="[198,309,152,177]" italics="true" pageId="15" pageNumber="16">Genitalia</emphasis>
. Paramere as in
<figureCitation id="137010E3FFF6806EFE3C28B2FDA84AE0" box="[503,573,152,178]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
j. Median lobe as in
<figureCitation id="137010E3FFF6806EFCE028B2FCFA4AE0" box="[811,879,152,178]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
k. Spermatheca as in
<figureCitation id="137010E3FFF6806EFBA228B2FB254AE0" box="[1129,1200,152,178]" captionStart="FIGURE 7" captionStartId="14.[151,258,1928,1952]" captionTargetBox="[165,1428,199,1907]" captionTargetId="figure@14.[151,1436,194,1907]" captionTargetPageId="14" captionText="FIGURE 7. Details of Leptusa (Dysleptusa) gimmeli sp. nov. a: antenna; b: labrum; c: labium; d: labial palpus; e: mentum; f: male abdominal tergite VIII; g: male abdominal sternite VIII; h: female abdominal tergite VIII; i: female abdominal sternite VIII; j: paramere; k: median lobe; l: spermatheca. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/199059/files/figure.png" pageId="15" pageNumber="16">
Fig.
<specimenCount id="9D4DC7EFFFF6806EFB5428B2FB254AE0" box="[1183,1200,152,178]" pageId="15" pageNumber="16" type="generic">7</specimenCount>
</figureCitation>
l.
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF6806EFF0D2897FF384AAE" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BF40C66FFF6806EFF0D2897FF384AAE" blockId="15.[151,1437,152,476]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2897FEF24A85" bold="true" box="[198,359,189,215]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="F35C4CF6FFF6806EFEA62897FE394A85" box="[365,428,189,215]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
: Tennessee. Locations where
<taxonomicName id="4C4B77E5FFF6806EFCCC2897FC2E4A84" box="[775,955,189,215]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gimmeli" subGenus="Dysleptusa">
<emphasis id="B93FD074FFF6806EFCCC2897FC8A4A84" box="[775,799,189,214]" italics="true" pageId="15" pageNumber="16">L.</emphasis>
(
<emphasis id="B93FD074FFF6806EFCE42897FCD94A84" box="[815,844,189,214]" italics="true" pageId="15" pageNumber="16">D.</emphasis>
)
<emphasis id="B93FD074FFF6806EFC972897FC2E4A84" box="[860,955,189,214]" italics="true" pageId="15" pageNumber="16">gimmeli</emphasis>
</taxonomicName>
has been collected in GSMNP as in Map 1.
</paragraph>
</subSubSection>
<subSubSection id="C3515FEDFFF6806EFF0D292DFC4E4B8E" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BF40C66FFF6806EFF0D292DFCC54B15" blockId="15.[151,1437,152,476]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D292DFEC64B73" bold="true" box="[198,339,263,289]" pageId="15" pageNumber="16">Etymology.</emphasis>
This species is named in honor of Matthew L. Gimmel (Louisiana State University, Baton Rouge), collector of all but one specimen in the
<typeStatus id="54F0B2C4FFF6806EFD002907FD6B4B15" box="[715,766,301,327]" pageId="15" pageNumber="16">type</typeStatus>
series.
</paragraph>
<paragraph id="8BF40C66FFF6806EFF0D2978FC4E4B8E" blockId="15.[151,1437,152,476]" pageId="15" pageNumber="16">
<emphasis id="B93FD074FFF6806EFF0D2978FEBB4B3E" bold="true" box="[198,302,338,364]" pageId="15" pageNumber="16">Habitat.</emphasis>
This species is known only from Albright Grove, an area of old growth forest in GSMNP and is represented by more specimens than any other species in the park. Over
<specimenCount id="9D4DC7EFFFF6806EFC3C2952FB334BC0" box="[1015,1190,376,402]" pageId="15" pageNumber="16" type="generic">100 specimens</specimenCount>
were collected from sifted leaf litter from both the spring and fall. Despite an extensive survey only a single specimen was collected from CWD in Albright Grove (emergent from decay class II).
</paragraph>
</subSubSection>
</treatment>
</document>
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