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<document id="FA685CB991D979735E1B4F4BEF5AFEC9" ID="10.11646/zootaxa.4188.1.1" ID-DOI="10.11646/zootaxa.4188.1.1" ID-GBIF-Dataset="03a7cb89-747a-411a-855c-f46ad7df47b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="165406" ID-ZooBank="AA85050E-7653-44BE-9330-AC617BFE6DF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478593795583" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Kottelat, Maurice" docDate="2016" docId="03E0BE0EFF895400FF58FC1BFF3E54FF" docLanguage="en" docName="zootaxa.4188.1.1.pdf" docOrigin="Zootaxa 4188 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Glyptothorax exodon Ng &amp; Rachmatika 2005" docType="treatment" docVersion="11" lastPageNumber="20" masterDocId="FFD9C676FF995413FFCFFFF2FFE65323" masterDocTitle="The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="17" updateTime="1698392689045" updateUser="plazi">
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<mods:title id="F2143AE2105D1A418CBEA99E372BAFCA">The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae)</mods:title>
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<mods:namePart id="C18015B18544340C016B03C016ABFAE1">Ng, Heok Hee</mods:namePart>
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<mods:namePart id="B0C901F75BB099E9DBDB6A447A648394">Kottelat, Maurice</mods:namePart>
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<mods:typeOfResource id="AA8740D7D136CE4957EAB0A47F72F8D7">text</mods:typeOfResource>
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<mods:title id="7A145956ED227FF968208EA163F77F13">Zootaxa</mods:title>
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<mods:date id="C345C0B9025432D9E42026DE4D25BA41">2016</mods:date>
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<mods:number id="1AB7F966B4105905737BF8E51DB58771">4188</mods:number>
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<mods:identifier id="AFDD01F52624A4FB28FFC1677A73BA34" type="DOI">10.11646/zootaxa.4188.1.1</mods:identifier>
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<treatment id="03E0BE0EFF895400FF58FC1BFF3E54FF" ID-DOI="http://doi.org/10.5281/zenodo.6063748" ID-GBIF-Taxon="124624015" ID-Zenodo-Dep="6063748" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF" httpUri="http://treatment.plazi.org/id/03E0BE0EFF895400FF58FC1BFF3E54FF" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="C3535C93FF895403FF58FC1BFF1E5706" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BF60F18FF895403FF58FC1BFD255727" blockId="16.[151,707,1001,1061]" box="[151,707,1001,1028]" pageId="16" pageNumber="17">
<heading id="D0BEB874FF895403FF58FC1BFD255727" bold="true" box="[151,707,1001,1028]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C49749BFF895403FF58FC1BFD255727" ID-CoL="3GJCB" authority="Ng &amp; Rachmatika 2005" authorityName="Ng &amp; Rachmatika" authorityYear="2005" box="[151,707,1001,1028]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF895403FF58FC1BFD255727" bold="true" box="[151,707,1001,1028]" pageId="16" pageNumber="17">
<emphasis id="B93DD30AFF895403FF58FC1BFE705720" bold="true" box="[151,406,1001,1027]" italics="true" pageId="16" pageNumber="17">Glyptothorax exodon</emphasis>
Ng &amp;
<bibRefCitation id="EFD872E9FF895403FE27FC18FD255727" author="Rachmatika" box="[488,707,1002,1028]" pageId="16" pageNumber="17" refString="Ng, H. H. &amp; Rachmatika, I. (2005) Glyptothorax exodon, a new species of rheophilic catfish from Borneo (Teleostei: Sisoridae). The Raffles Bulletin of Zoology, 53, 251 - 255." type="journal article" year="2005">Rachmatika 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF60F18FF895403FF58FBFEFF1E5706" blockId="16.[151,707,1001,1061]" box="[151,248,1036,1061]" pageId="16" pageNumber="17">
(
<figureCitation id="1372139DFF895403FF50FBFEFF175706" box="[159,241,1036,1061]" captionStart="FIGURE 10" captionStartId="18.[151,250,1509,1531]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@18.[151,1436,193,1487]" captionTargetPageId="18" captionText="FIGURE 10. Glyptothorax exodon, BMNH 1982.3. 29.190, paratype, 63.2 mm SL; Borneo: Kalimantan Barat, Sungai Pinoh. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/266204/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF895403FF58FBBDFC6F57E2" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BF60F18FF895403FF58FBBDFD1E57A7" blockId="16.[151,1375,1103,1217]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB592836FF895403FF58FBBDFD1E57A7" pageId="16" pageNumber="17">
<taxonomicName id="4C49749BFF895403FF58FBBDFD465746" ID-CoL="3GJCB" authority="Ng &amp; Rachmatika, 2005: 251" authorityName="Ng &amp; Rachmatika" authorityPageNumber="251" authorityYear="2005" box="[151,672,1103,1125]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF895403FF58FBBDFE8B5746" box="[151,365,1103,1125]" italics="true" pageId="16" pageNumber="17">Glyptothorax exodon</emphasis>
Ng &amp;
<treatmentCitation id="0AE82909FF895403FE7CFBBDFD465746" author="Rachmatika" box="[435,672,1103,1125]" page="251" pageId="16" pageNumber="17" year="2005">
<bibRefCitation id="EFD872E9FF895403FE7CFBBDFD885746" author="Rachmatika" box="[435,622,1103,1125]" pageId="16" pageNumber="17" refString="Ng, H. H. &amp; Rachmatika, I. (2005) Glyptothorax exodon, a new species of rheophilic catfish from Borneo (Teleostei: Sisoridae). The Raffles Bulletin of Zoology, 53, 251 - 255." type="journal article" year="2005">Rachmatika, 2005</bibRefCitation>
: 251
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="1372139DFF895403FD64FBBDFD165746" box="[683,752,1103,1125]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[230,1355,767,1931]" captionTargetId="figure@3.[228,1358,767,1931]" captionTargetPageId="3" captionText="FIGURE 1. Schematic illustration of thoracic adhesive apparatus of Glyptothorax, showing structures defined and mentioned in this study: anterolateral striae (ALS); anteromedial striae (AMS); medial pit (MP) (based on G. plectilis, MZB 4530, 50.2 mm SL)." httpUri="https://zenodo.org/record/165407/files/figure.png" pageId="16" pageNumber="17">Figs. 1</figureCitation>
,
<figureCitation id="1372139DFF895403FD33FBBDFCEF5746" box="[764,777,1103,1125]" captionStart="FIGURE 2" captionStartId="5.[151,250,1720,1742]" captionTargetBox="[150,1435,301,1698]" captionTargetId="figure@5.[150,1436,301,1699]" captionTargetPageId="5" captionText="FIGURE 2. Glyptothorax amnestus, MZB 17214, holotype, 88.1 mm SL; Sumatra: Padang Panjang. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/165408/files/figure.png" pageId="16" pageNumber="17">2</figureCitation>
(type locality: Sungai Tekelan, Kalimantan Barat, Borneo)
<taxonomicName id="4C49749BFF895403FF58FB9DFEB257A7" ID-CoL="KDCB" box="[151,340,1134,1156]" class="Actinopterygii" family="Sisoridae" genus="Bagarius" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="bagarius">
<emphasis id="B93DD30AFF895403FF58FB9DFEB257A7" box="[151,340,1134,1156]" italics="true" pageId="16" pageNumber="17">Bagarius bagarius</emphasis>
</taxonomicName>
(non Hamilton, 1822)
<treatmentCitation id="0AE82909FF895403FDF3FB9CFD1257A7" author="Vaillant" box="[572,756,1134,1156]" page="72" pageId="16" pageNumber="17" year="1902">
<bibRefCitation id="EFD872E9FF895403FDF3FB9CFD2857A7" author="Vaillant" box="[572,718,1134,1156]" pageId="16" pageNumber="17" refString="Vaillant, L. L. (1902) Resultats zoologiques de l'expedition scientifique Neerlandaise au Borneo central. Poissons. Notes from the Leyden Museum, 24, 1 - 166." type="journal article" year="1902">Vaillant, 1902</bibRefCitation>
: 72
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF60F18FF895403FF58FB7EFC6F57E2" blockId="16.[151,1375,1103,1217]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB592836FF895403FF58FB7EFC6F57E2" pageId="16" pageNumber="17">
<taxonomicName id="4C49749BFF895403FF58FB7EFE2D5781" ID-CoL="6KM52" box="[151,459,1164,1186]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="platypogonoides">
<emphasis id="B93DD30AFF895403FF58FB7EFE2D5781" box="[151,459,1164,1186]" italics="true" pageId="16" pageNumber="17">Glyptothorax platypogonoides</emphasis>
</taxonomicName>
(non Bleeker, 1855)—Roberts, 1989: 136, Fig. 104 lower (in part).
<taxonomicName id="4C49749BFF895403FF58FB59FE0D57E2" ID-CoL="6KM52" box="[151,491,1195,1217]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="platypogonoides">
<emphasis id="B93DD30AFF895403FF58FB59FEF857E2" box="[151,286,1195,1217]" italics="true" pageId="16" pageNumber="17">Glyptothorax</emphasis>
cf.
<emphasis id="B93DD30AFF895403FE8BFB59FE0D57E2" box="[324,491,1195,1217]" italics="true" pageId="16" pageNumber="17">platypogonoides</emphasis>
</taxonomicName>
—Ng &amp;
<treatmentCitation id="0AE82909FF895403FD8BFB59FCD457E2" author="Rachmatika" box="[580,818,1195,1217]" page="182" pageId="16" pageNumber="17" year="1999">
<bibRefCitation id="EFD872E9FF895403FD8BFB59FCE657E2" author="Rachmatika" box="[580,768,1195,1217]" pageId="16" pageNumber="17" refString="Ng, H. H. &amp; Rachmatika, I. (1999) The catfishes (Teleostei: Siluriformes) of Bentuang Karimun National Park, West Kalimantan, Indonesia. The Raffles Bulletin of Zoology, 47, 167 - 183." type="journal article" year="1999">Rachmatika, 1999</bibRefCitation>
: 182
</treatmentCitation>
,
<figureCitation id="1372139DFF895403FCF3FB59FC9157E2" box="[828,887,1195,1217]" captionStart="FIGURE 8" captionStartId="14.[151,250,1807,1829]" captionTargetBox="[151,1434,193,1785]" captionTargetId="figure@14.[151,1436,193,1785]" captionTargetPageId="14" captionText="FIGURE 8. Glyptothorax decussatus, MZB 17216, holotype, 74.5 mm SL; Borneo: Kalimantan Timur, Enggeng Bio. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/266203/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
B
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF895403FF58FB1AFAC456FA" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BF60F18FF895403FF58FB1AFAC456FA" blockId="16.[151,1437,1256,2037]" pageId="16" pageNumber="17">
<materialsCitation id="3B210545FF895403FF58FB1AFE7B5606" ID-GBIF-Occurrence="1324948229" collectionCode="MZB" location="Kapuas River" municipality="Sungai Tekelan" pageId="16" pageNumber="17" specimenCode="MZB 9940" specimenCount="1" stateProvince="KALIMANTAN BARAT" typeStatus="holotype">
<emphasis id="B93DD30AFF895403FF58FB1AFE645622" bold="true" box="[151,386,1256,1281]" pageId="16" pageNumber="17">Material examined.</emphasis>
BORNEO:
<collectingRegion id="498DC1FAFF895403FDC1FB1BFCD75622" box="[526,817,1257,1281]" country="Indonesia" name="Kalimantan Barat" pageId="16" pageNumber="17">KALIMANTAN BARAT</collectingRegion>
:
<specimenCode id="DBEFA763FF895403FCF2FB1BFC5C5622" box="[829,954,1257,1281]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FCF2FB1BFC9C5623" box="[829,890,1257,1280]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">MZB</collectionCode>
9940
</specimenCode>
(
<typeStatus id="54F2B1BAFF895403FC04FB1AFBD55622" box="[971,1075,1256,1281]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
),
<quantity id="4CB1A2FDFF895403FB88FB1AFB4B5623" box="[1095,1197,1256,1281]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" pageId="16" pageNumber="17" unit="mm" value="57.5">57.5 mm</quantity>
SL;
<collectingMunicipality id="6B929562FF895403FB29FB1AFA7E5622" box="[1254,1432,1256,1281]" pageId="16" pageNumber="17">Sungai Tekelan</collectingMunicipality>
,
<location id="8E9659C3FF895403FF58FAFFFED75606" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FF58FAFFFED75606" box="[151,305,1292,1317]" municipality="Sungai Tekelan" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage
</materialsCitation>
. BMNH 1982.3.29.190 (1 paratype),
<quantity id="4CB1A2FDFF895403FC89FAFEFC4D5607" box="[838,939,1292,1317]" metricMagnitude="-2" metricUnit="m" metricValue="6.32" pageId="16" pageNumber="17" unit="mm" value="63.2">63.2 mm</quantity>
SL; CAS 49419 (1 paratype), 59.0 mm SL;
<materialsCitation id="3B210545FF895403FF58FAC3FE7A564E" ID-GBIF-Occurrence="1324948225" collectionCode="RMNH" county="Sungai Pinoh" location="Kapuas River" municipality="Nangapinoh" pageId="16" pageNumber="17" specimenCode="RMNH 28907" specimenCount="1" stateProvince="KALIMANTAN BARAT" typeStatus="paratype">
<specimenCode id="DBEFA763FF895403FF58FAC3FEDD566A" box="[151,315,1329,1353]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FF58FAC3FF0D566B" box="[151,235,1329,1352]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
28907
</specimenCode>
(
<specimenCount id="9D4FC491FF895403FE9FFAC3FE2D566A" box="[336,459,1329,1353]" pageId="16" pageNumber="17" type="generic" typeStatus="paratype">1 paratype</specimenCount>
),
<quantity id="4CB1A2FDFF895403FE2EFAC2FDAE566B" box="[481,584,1328,1353]" metricMagnitude="-2" metricUnit="m" metricValue="5.72" pageId="16" pageNumber="17" unit="mm" value="57.2">57.2 mm</quantity>
SL; mainstream of
<collectingCounty id="62977794FF895403FCFCFAC2FC29566A" box="[819,975,1328,1353]" pageId="16" pageNumber="17">Sungai Pinoh</collectingCounty>
,
<quantity id="4CB1A2FDFF895403FC12FAC2FBB7566A" box="[989,1105,1328,1353]" metricMagnitude="4" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="16" pageNumber="17" unit="km" value="40.0" valueMax="60.0" valueMin="20.0">2060 km</quantity>
upstream from
<collectingMunicipality id="6B929562FF895403FAC3FAC2FA7E566A" box="[1292,1432,1328,1353]" pageId="16" pageNumber="17">Nangapinoh</collectingMunicipality>
,
<location id="8E9659C3FF895403FF58FAA7FED6564E" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FF58FAA7FED6564E" box="[151,304,1364,1389]" county="Sungai Pinoh" municipality="Nangapinoh" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage
</materialsCitation>
.
<materialsCitation id="3B210545FF895403FE68FAA7FAF3564E" ID-GBIF-Occurrence="1324948227" box="[423,1301,1364,1389]" collectionCode="RMNH" location="Kapuas River" municipality="Nanga Raun" pageId="16" pageNumber="17" specimenCode="RMNH 7845" specimenCount="1" stateProvince="KALIMANTAN BARAT" typeStatus="paratype">
<specimenCode id="DBEFA763FF895403FE68FAA7FDDD564E" box="[423,571,1364,1389]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FE68FAA7FE1D564F" box="[423,507,1365,1388]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
7845
</specimenCode>
(
<specimenCount id="9D4FC491FF895403FD82FAA7FD20564E" box="[589,710,1365,1389]" pageId="16" pageNumber="17" type="generic" typeStatus="paratype">1 paratype</specimenCount>
),
<quantity id="4CB1A2FDFF895403FD16FAA6FCD8564F" box="[729,830,1364,1389]" metricMagnitude="-2" metricUnit="m" metricValue="6.02" pageId="16" pageNumber="17" unit="mm" value="60.2">60.2 mm</quantity>
SL;
<collectingMunicipality id="6B929562FF895403FCBAFAA7FBE2564F" box="[885,1028,1365,1389]" pageId="16" pageNumber="17">Nanga Raun</collectingMunicipality>
,
<location id="8E9659C3FF895403FBDFFAA7FB4F564E" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FBDFFAA7FB4F564E" box="[1040,1193,1364,1389]" municipality="Nanga Raun" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage
</materialsCitation>
.
<materialsCitation id="3B210545FF895403FAEFFAA7FBD856B2" ID-GBIF-Occurrence="1324948221" collectionCode="MZB" county="Sungai Jaket" location="Kapuas River" municipality="Embaloh" pageId="16" pageNumber="17" specimenCode="MZB 9412" specimenCount="2" stateProvince="KALIMANTAN BARAT" typeStatus="paratype">
<specimenCode id="DBEFA763FF895403FAEFFAA7FA7A564E" box="[1312,1436,1365,1389]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FAEFFAA7FABA564F" box="[1312,1372,1365,1388]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">MZB</collectionCode>
9412
</specimenCode>
(
<specimenCount id="9D4FC491FF895403FF50FA8BFECE56B2" box="[159,296,1401,1425]" pageId="16" pageNumber="17" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
), 55.160.0 mm SL;
<collectingCounty id="62977794FF895403FDD1FA8AFD5456B2" box="[542,690,1400,1425]" pageId="16" pageNumber="17">Sungai Jaket</collectingCounty>
,
<collectingMunicipality id="6B929562FF895403FD70FA8AFCC156B2" box="[703,807,1400,1425]" pageId="16" pageNumber="17">Embaloh</collectingMunicipality>
,
<location id="8E9659C3FF895403FCFAFA8BFC3656B2" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FCFAFA8BFC3656B2" box="[821,976,1400,1425]" county="Sungai Jaket" municipality="Embaloh" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage
</materialsCitation>
.
<materialsCitation id="3B210545FF895403FB84FA8BFD8B5696" ID-GBIF-Occurrence="1324948220" collectionCode="MZB" location="Kapuas River" municipality="Sungai Embaloh" pageId="16" pageNumber="17" specimenCode="MZB 9204" specimenCount="1" stateProvince="KALIMANTAN BARAT" typeStatus="paratype">
<specimenCode id="DBEFA763FF895403FB84FA8BFB2F56B2" box="[1099,1225,1401,1425]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FB84FA8BFB6156B3" box="[1099,1159,1401,1424]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">MZB</collectionCode>
9204
</specimenCode>
(1),
<quantity id="4CB1A2FDFF895403FACBFA8BFA8D56B3" box="[1284,1387,1401,1425]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="16" pageNumber="17" unit="mm" value="42.9">42.9 mm</quantity>
SL;
<collectingMunicipality id="6B929562FF895403FF58FA6EFEB05696" box="[151,342,1436,1461]" pageId="16" pageNumber="17">Sungai Embaloh</collectingMunicipality>
,
<location id="8E9659C3FF895403FEABFA6FFE195696" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FEABFA6FFE195696" box="[356,511,1436,1461]" municipality="Sungai Embaloh" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage
</materialsCitation>
.
<materialsCitation id="3B210545FF895403FDB5FA6FFAC456FA" ID-GBIF-Occurrence="1324948218" collectionCode="MZB" county="Sungai Peyang" location="Sungai Sibau" municipality="Kapuas River" pageId="16" pageNumber="17" specimenCode="MZB 6633" specimenCount="1" stateProvince="KALIMANTAN BARAT" typeStatus="paratype">
<specimenCode id="DBEFA763FF895403FDB5FA6FFD1E5696" box="[634,760,1436,1461]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">
<collectionCode id="ED5897DDFF895403FDB5FA6FFD515697" box="[634,695,1437,1460]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="16" pageNumber="17">MZB</collectionCode>
6633
</specimenCode>
(1),
<quantity id="4CB1A2FDFF895403FCFCFA6FFC7C5697" box="[819,922,1437,1461]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="16" pageNumber="17" unit="mm" value="42.9">42.9 mm</quantity>
SL;
<collectingCounty id="62977794FF895403FC1AFA6EFB675696" box="[981,1153,1436,1461]" pageId="16" pageNumber="17">Sungai Peyang</collectingCounty>
,
<collectingMunicipality id="6B929562FF895403FB40FA6FFACD5696" box="[1167,1323,1436,1461]" pageId="16" pageNumber="17">Kapuas River</collectingMunicipality>
drainage. CMK 6905 (1),
<quantity id="4CB1A2FDFF895403FE84FA33FE4956FB" box="[331,431,1473,1497]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="16" pageNumber="17" unit="mm" value="29.1">29.1 mm</quantity>
SL;
<location id="8E9659C3FF895403FE2AFA32FD9C56FA" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FE2AFA32FD9C56FA" box="[485,634,1472,1497]" county="Sungai Peyang" municipality="Kapuas River" name="Sungai Sibau" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Sungai Sibau</location>
about
<quantity id="4CB1A2FDFF895403FD09FA33FD1956FA" box="[710,767,1472,1497]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="km" value="2.0">2 km</quantity>
upstream of
<location id="8E9659C3FF895403FC5DFA32FBE856FA" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FC5DFA32FBE856FA" box="[914,1038,1472,1497]" county="Sungai Peyang" municipality="Kapuas River" name="Putussibau" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Putussibau</location>
,
<location id="8E9659C3FF895403FBD6FA33FB5456FA" LSID="urn:lsid:plazi:treatment:03E0BE0EFF895400FF58FC1BFF3E54FF:8E9659C3FF895403FBD6FA33FB5456FA" box="[1049,1202,1472,1497]" county="Sungai Peyang" municipality="Kapuas River" name="Kapuas River" pageId="16" pageNumber="17" stateProvince="KALIMANTAN BARAT">Kapuas River</location>
drainage.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF895403FF08FA16FE17543E" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BF60F18FF895403FF08FA16FE17543E" blockId="16.[151,1437,1256,2037]" pageId="16" pageNumber="17">
<emphasis id="B93DD30AFF895403FF08FA16FEA756DE" bold="true" box="[199,321,1508,1533]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="4C49749BFF895403FE83FA17FDD856DE" box="[332,574,1509,1533]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF895403FE83FA17FDD856DE" box="[332,574,1509,1533]" italics="true" pageId="16" pageNumber="17">Glyptothorax exodon</emphasis>
</taxonomicName>
can be distinguished from Sundaic congeners in having the premaxillary toothband almost entirely (vs. two-thirds or less) exposed when the mouth is closed. The following unique combination of characters further distinguishes
<taxonomicName id="4C49749BFF895403FD0FF9DFFCD55566" box="[704,819,1581,1605]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF895403FD0FF9DFFCD55566" box="[704,819,1581,1605]" italics="true" pageId="16" pageNumber="17">G. exodon</emphasis>
</taxonomicName>
from Sundaic congeners: eye diameter 1013% HL; head width 16.018.1% SL; head depth 13.815.1% SL; anteromedial striae in thoracic adhesive apparatus absent; prepectoral length 20.625.4% SL; predorsal length 36.739.2% SL; dorsal-fin spine length 15.020.0% SL; margin of dorsal fin concave; smooth posterior margin of dorsal-fin spine; dorsal-to-adipose distance 21.926.3% SL; body depth at anus 13.815.8% SL; prepelvic length 48.652.0% SL; post-adipose distance 21.524.0% SL; caudal peduncle depth 6.87.3% SL; straight dorsoposterior margin of adipose fin; and mottled body pattern lacking prominent pale stripes.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF895400FF08F8DAFEFC511E" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF60F18FF895403FF08F8DAFBD854D6" blockId="16.[151,1437,1256,2037]" pageId="16" pageNumber="17">
<emphasis id="B93DD30AFF895403FF08F8DAFEBF5462" bold="true" box="[199,345,1832,1857]" pageId="16" pageNumber="17">Description.</emphasis>
Morphometric data in
<tableCitation id="C6CB3AA3FF895403FD94F8DAFD495462" box="[603,687,1832,1857]" captionStart="TABLE 3" captionStartId="17.[151,239,152,175]" captionTargetBox="[159,1147,202,1506]" captionTargetPageId="17" captionText="TABLE 3. Morphometric data for Glyptothorax exodon (n = 7)." httpUri="http://table.plazi.org/id/DF365F90FF885402FF58FF6AFCD2538C" pageId="16" pageNumber="17" tableUuid="DF365F90FF885402FF58FF6AFCD2538C">Table 3</tableCitation>
. Head depressed; body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin prominently tuberculate, with ovoid tubercles particularly visible on sides of body. Lateral line complete and midlateral. Vertebrae 17+18=35 (1), 17+19=36 (2), 18+18=36 (2) or 18+19=37 (2).
</paragraph>
<caption id="DF365F90FF885402FF58FF6AFCD2538C" ID-Table-UUID="DF365F90FF885402FF58FF6AFCD2538C" box="[151,820,151,175]" httpUri="http://table.plazi.org/id/DF365F90FF885402FF58FF6AFCD2538C" pageId="17" pageNumber="18" targetBox="[159,1147,202,1506]" targetIsTable="true" targetPageId="17">
<paragraph id="8BF60F18FF885402FF58FF6AFCD2538C" blockId="17.[151,820,151,175]" box="[151,820,151,175]" pageId="17" pageNumber="18">
<emphasis id="B93DD30AFF885402FF58FF6AFEEF538C" bold="true" box="[151,265,152,175]" pageId="17" pageNumber="18">TABLE 3.</emphasis>
Morphometric data for
<taxonomicName id="4C49749BFF885402FDC5FF65FD0A538C" box="[522,748,151,175]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF885402FDC5FF65FD0A538C" box="[522,748,151,175]" italics="true" pageId="17" pageNumber="18">Glyptothorax exodon</emphasis>
</taxonomicName>
(n=7).
</paragraph>
</caption>
<paragraph id="8BF60F18FF885402FD4BFF39FBBD56C1" pageId="17" pageNumber="18">
<table id="F949FDB8FF88ABECFF50FF38FB9D56C1" box="[159,1147,202,1506]" gridcols="3" gridrows="32" pageId="17" pageNumber="18">
<tr id="35790D5AFF88ABECFF50FF38FB9D53C3" box="[159,1147,202,224]" gridrow="0" pageId="17" pageNumber="18" rowspan-0="1">
<th id="76A86426FF88ABECFD4BFF38FD0C53C3" box="[644,746,202,224]" gridcol="1" gridrow="0" pageId="17" pageNumber="18">Range</th>
<th id="76A86426FF88ABECFBDBFF38FB9D53C3" box="[1044,1147,202,224]" gridcol="2" gridrow="0" pageId="17" pageNumber="18">Mean±SD</th>
</tr>
<tr id="35790D5AFF88ABECFF50FF06FB9D5229" box="[159,1147,244,266]" gridrow="1" pageId="17" pageNumber="18" rowspan-2="1">
<th id="76A86426FF88ABECFF50FF06FE3B5229" box="[159,477,244,266]" gridcol="0" gridrow="1" pageId="17" pageNumber="18">Standard length (mm)</th>
<td id="76A86426FF88ABECFD4BFF06FD0C5229" box="[644,746,244,266]" gridcol="1" gridrow="1" pageId="17" pageNumber="18">55.163.2</td>
</tr>
<tr id="35790D5AFF88ABECFF50FEEFFB9D5210" box="[159,1147,285,307]" gridrow="2" pageId="17" pageNumber="18" rowspan-1="1" rowspan-2="1">
<th id="76A86426FF88ABECFF50FEEFFE3B5210" box="[159,477,285,307]" gridcol="0" gridrow="2" pageId="17" pageNumber="18">%SL</th>
</tr>
<tr id="35790D5AFF88ABECFF50FEB4FB9D527F" box="[159,1147,326,348]" gridrow="3" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FEB4FE3B527F" box="[159,477,326,348]" gridcol="0" gridrow="3" pageId="17" pageNumber="18">Predorsal length</th>
<td id="76A86426FF88ABECFD4BFEB4FD0C527F" box="[644,746,326,348]" gridcol="1" gridrow="3" pageId="17" pageNumber="18">36.739.2</td>
<td id="76A86426FF88ABECFBDBFEB4FB9D527F" box="[1044,1147,326,348]" gridcol="2" gridrow="3" pageId="17" pageNumber="18">37.9±0.96</td>
</tr>
<tr id="35790D5AFF88ABECFF50FE82FB9D52A5" box="[159,1147,368,390]" gridrow="4" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FE82FE3B52A5" box="[159,477,368,390]" gridcol="0" gridrow="4" pageId="17" pageNumber="18">Preanal length</th>
<td id="76A86426FF88ABECFD4BFE82FD0C52A5" box="[644,746,368,390]" gridcol="1" gridrow="4" pageId="17" pageNumber="18">64.167.8</td>
<td id="76A86426FF88ABECFBDBFE82FB9D52A5" box="[1044,1147,368,390]" gridcol="2" gridrow="4" pageId="17" pageNumber="18">66.4±1.40</td>
</tr>
<tr id="35790D5AFF88ABECFF50FE6BFB9D528C" box="[159,1147,409,431]" gridrow="5" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FE6BFE3B528C" box="[159,477,409,431]" gridcol="0" gridrow="5" pageId="17" pageNumber="18">Prepelvic length</th>
<td id="76A86426FF88ABECFD4BFE6BFD0C528C" box="[644,746,409,431]" gridcol="1" gridrow="5" pageId="17" pageNumber="18">48.653.0</td>
<td id="76A86426FF88ABECFBDBFE6BFB9D528C" box="[1044,1147,409,431]" gridcol="2" gridrow="5" pageId="17" pageNumber="18">49.7±1.69</td>
</tr>
<tr id="35790D5AFF88ABECFF50FE30FB9D52FB" box="[159,1147,450,472]" gridrow="6" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FE30FE3B52FB" box="[159,477,450,472]" gridcol="0" gridrow="6" pageId="17" pageNumber="18">Prepectoral length</th>
<td id="76A86426FF88ABECFD4BFE30FD0C52FB" box="[644,746,450,472]" gridcol="1" gridrow="6" pageId="17" pageNumber="18">20.625.4</td>
<td id="76A86426FF88ABECFBDBFE30FB9D52FB" box="[1044,1147,450,472]" gridcol="2" gridrow="6" pageId="17" pageNumber="18">22.7±2.16</td>
</tr>
<tr id="35790D5AFF88ABECFF50FE1EFB9D5121" box="[159,1147,492,514]" gridrow="7" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FE1EFE3B5121" box="[159,477,492,514]" gridcol="0" gridrow="7" pageId="17" pageNumber="18">Length of dorsal-fin base</th>
<td id="76A86426FF88ABECFD4BFE1EFD0C5121" box="[644,746,492,514]" gridcol="1" gridrow="7" pageId="17" pageNumber="18">10.614.9</td>
<td id="76A86426FF88ABECFBDBFE1EFB9D5121" box="[1044,1147,492,514]" gridcol="2" gridrow="7" pageId="17" pageNumber="18">12.8±1.43</td>
</tr>
<tr id="35790D5AFF88ABECFF50FDE7FB9D5108" box="[159,1147,533,555]" gridrow="8" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FDE7FE3B5108" box="[159,477,533,555]" gridcol="0" gridrow="8" pageId="17" pageNumber="18">Dorsal-fin spine length</th>
<td id="76A86426FF88ABECFD4BFDE7FD0C5108" box="[644,746,533,555]" gridcol="1" gridrow="8" pageId="17" pageNumber="18">15.020.0</td>
<td id="76A86426FF88ABECFBDBFDE7FB9D5108" box="[1044,1147,533,555]" gridcol="2" gridrow="8" pageId="17" pageNumber="18">16.7±1.95</td>
</tr>
<tr id="35790D5AFF88ABECFF50FDCCFB9D5177" box="[159,1147,574,596]" gridrow="9" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FDCCFE3B5177" box="[159,477,574,596]" gridcol="0" gridrow="9" pageId="17" pageNumber="18">Length of anal-fin base</th>
<td id="76A86426FF88ABECFD4BFDCCFD0C5177" box="[644,746,574,596]" gridcol="1" gridrow="9" pageId="17" pageNumber="18">14.215.3</td>
<td id="76A86426FF88ABECFBDBFDCCFB9D5177" box="[1044,1147,574,596]" gridcol="2" gridrow="9" pageId="17" pageNumber="18">14.9±0.46</td>
</tr>
<tr id="35790D5AFF88ABECFF50FD9AFB9D515D" box="[159,1147,616,638]" gridrow="10" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FD9AFE3B515D" box="[159,477,616,638]" gridcol="0" gridrow="10" pageId="17" pageNumber="18">Pelvic-fin length</th>
<td id="76A86426FF88ABECFD4BFD9AFD0C515D" box="[644,746,616,638]" gridcol="1" gridrow="10" pageId="17" pageNumber="18">15.216.5</td>
<td id="76A86426FF88ABECFBDBFD9AFB9D515D" box="[1044,1147,616,638]" gridcol="2" gridrow="10" pageId="17" pageNumber="18">15.7±0.46</td>
</tr>
<tr id="35790D5AFF88ABECFF50FD63FB9D5184" box="[159,1147,657,679]" gridrow="11" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FD63FE3B5184" box="[159,477,657,679]" gridcol="0" gridrow="11" pageId="17" pageNumber="18">Pectoral-fin length</th>
<td id="76A86426FF88ABECFD4BFD63FD0C5184" box="[644,746,657,679]" gridcol="1" gridrow="11" pageId="17" pageNumber="18">22.125.0</td>
<td id="76A86426FF88ABECFBDBFD63FB9D5184" box="[1044,1147,657,679]" gridcol="2" gridrow="11" pageId="17" pageNumber="18">23.7±1.06</td>
</tr>
<tr id="35790D5AFF88ABECFF50FD48FB9D51F3" box="[159,1147,698,720]" gridrow="12" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FD48FE3B51F3" box="[159,477,698,720]" gridcol="0" gridrow="12" pageId="17" pageNumber="18">Pectoral-fin spine length</th>
<td id="76A86426FF88ABECFD4BFD48FD0C51F3" box="[644,746,698,720]" gridcol="1" gridrow="12" pageId="17" pageNumber="18">16.220.3</td>
<td id="76A86426FF88ABECFBDBFD48FB9D51F3" box="[1044,1147,698,720]" gridcol="2" gridrow="12" pageId="17" pageNumber="18">18.7±1.51</td>
</tr>
<tr id="35790D5AFF88ABECFF50FD16FB9D51D9" box="[159,1147,740,762]" gridrow="13" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FD16FE3B51D9" box="[159,477,740,762]" gridcol="0" gridrow="13" pageId="17" pageNumber="18">Caudal-fin length</th>
<td id="76A86426FF88ABECFD4BFD16FD0C51D9" box="[644,746,740,762]" gridcol="1" gridrow="13" pageId="17" pageNumber="18">28.031.0</td>
<td id="76A86426FF88ABECFBDBFD16FB9D51D9" box="[1044,1147,740,762]" gridcol="2" gridrow="13" pageId="17" pageNumber="18">29.2±1.02</td>
</tr>
<tr id="35790D5AFF88ABECFF50FCFFFB9D5000" box="[159,1147,781,803]" gridrow="14" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FCFFFE3B5000" box="[159,477,781,803]" gridcol="0" gridrow="14" pageId="17" pageNumber="18">Length of adipose-fin base</th>
<td id="76A86426FF88ABECFD4BFCFFFD0C5000" box="[644,746,781,803]" gridcol="1" gridrow="14" pageId="17" pageNumber="18">11.014.0</td>
<td id="76A86426FF88ABECFBDBFCFFFB9D5000" box="[1044,1147,781,803]" gridcol="2" gridrow="14" pageId="17" pageNumber="18">13.0±1.13</td>
</tr>
<tr id="35790D5AFF88ABECFF50FCC4FB9D506F" box="[159,1147,822,844]" gridrow="15" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FCC4FE3B506F" box="[159,477,822,844]" gridcol="0" gridrow="15" pageId="17" pageNumber="18">Dorsal to adipose distance</th>
<td id="76A86426FF88ABECFD4BFCC4FD0C506F" box="[644,746,822,844]" gridcol="1" gridrow="15" pageId="17" pageNumber="18">21.926.3</td>
<td id="76A86426FF88ABECFBDBFCC4FB9D506F" box="[1044,1147,822,844]" gridcol="2" gridrow="15" pageId="17" pageNumber="18">23.4±2.51</td>
</tr>
<tr id="35790D5AFF88ABECFF50FC92FB9D5055" box="[159,1147,864,886]" gridrow="16" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FC92FE3B5055" box="[159,477,864,886]" gridcol="0" gridrow="16" pageId="17" pageNumber="18">Post-adipose distance</th>
<td id="76A86426FF88ABECFD4BFC92FD0C5055" box="[644,746,864,886]" gridcol="1" gridrow="16" pageId="17" pageNumber="18">21.524.0</td>
<td id="76A86426FF88ABECFBDBFC92FB9D5055" box="[1044,1147,864,886]" gridcol="2" gridrow="16" pageId="17" pageNumber="18">22.5±1.32</td>
</tr>
<tr id="35790D5AFF88ABECFF50FC7BFB9D50BC" box="[159,1147,905,927]" gridrow="17" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FC7BFE3B50BC" box="[159,477,905,927]" gridcol="0" gridrow="17" pageId="17" pageNumber="18">Length of caudal peduncle</th>
<td id="76A86426FF88ABECFD4BFC7BFD0C50BC" box="[644,746,905,927]" gridcol="1" gridrow="17" pageId="17" pageNumber="18">18.020.6</td>
<td id="76A86426FF88ABECFBDBFC7BFB9D50BC" box="[1044,1147,905,927]" gridcol="2" gridrow="17" pageId="17" pageNumber="18">19.3±1.07</td>
</tr>
<tr id="35790D5AFF88ABECFF50FC40FB9D50EB" box="[159,1147,946,968]" gridrow="18" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FC40FE3B50EB" box="[159,477,946,968]" gridcol="0" gridrow="18" pageId="17" pageNumber="18">Depth of caudal peduncle</th>
<td id="76A86426FF88ABECFD4BFC40FD0C50EB" box="[644,746,946,968]" gridcol="1" gridrow="18" pageId="17" pageNumber="18">6.87.3</td>
<td id="76A86426FF88ABECFBDBFC40FB9D50EB" box="[1044,1147,946,968]" gridcol="2" gridrow="18" pageId="17" pageNumber="18">7.0±0.21</td>
</tr>
<tr id="35790D5AFF88ABECFF50FC2EFB9D50D1" box="[159,1147,988,1010]" gridrow="19" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FC2EFE3B50D1" box="[159,477,988,1010]" gridcol="0" gridrow="19" pageId="17" pageNumber="18">Body depth at anus</th>
<td id="76A86426FF88ABECFD4BFC2EFD0C50D1" box="[644,746,988,1010]" gridcol="1" gridrow="19" pageId="17" pageNumber="18">13.815.8</td>
<td id="76A86426FF88ABECFBDBFC2EFB9D50D1" box="[1044,1147,988,1010]" gridcol="2" gridrow="19" pageId="17" pageNumber="18">14.5±0.88</td>
</tr>
<tr id="35790D5AFF88ABECFF50FBF7FB9D5738" box="[159,1147,1029,1051]" gridrow="20" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FBF7FE3B5738" box="[159,477,1029,1051]" gridcol="0" gridrow="20" pageId="17" pageNumber="18">Body depth at dorsal-fin origin</th>
<td id="76A86426FF88ABECFD4BFBF7FD0C5738" box="[644,746,1029,1051]" gridcol="1" gridrow="20" pageId="17" pageNumber="18">18.920.0</td>
<td id="76A86426FF88ABECFBDBFBF7FB9D5738" box="[1044,1147,1029,1051]" gridcol="2" gridrow="20" pageId="17" pageNumber="18">19.6±0.59</td>
</tr>
<tr id="35790D5AFF88ABECFF50FBDCFB9D5767" box="[159,1147,1070,1092]" gridrow="21" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FBDCFE3B5767" box="[159,477,1070,1092]" gridcol="0" gridrow="21" pageId="17" pageNumber="18">Head length</th>
<td id="76A86426FF88ABECFD4BFBDCFD0C5767" box="[644,746,1070,1092]" gridcol="1" gridrow="21" pageId="17" pageNumber="18">25.729.2</td>
<td id="76A86426FF88ABECFBDBFBDCFB9D5767" box="[1044,1147,1070,1092]" gridcol="2" gridrow="21" pageId="17" pageNumber="18">26.8±1.28</td>
</tr>
<tr id="35790D5AFF88ABECFF50FBAAFB9D574D" box="[159,1147,1112,1134]" gridrow="22" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FBAAFE3B574D" box="[159,477,1112,1134]" gridcol="0" gridrow="22" pageId="17" pageNumber="18">Head width</th>
<td id="76A86426FF88ABECFD4BFBAAFD0C574D" box="[644,746,1112,1134]" gridcol="1" gridrow="22" pageId="17" pageNumber="18">16.018.1</td>
<td id="76A86426FF88ABECFBDBFBAAFB9D574D" box="[1044,1147,1112,1134]" gridcol="2" gridrow="22" pageId="17" pageNumber="18">17.0±0.85</td>
</tr>
<tr id="35790D5AFF88ABECFF50FB73FB9D57B4" box="[159,1147,1153,1175]" gridrow="23" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FB73FE3B57B4" box="[159,477,1153,1175]" gridcol="0" gridrow="23" pageId="17" pageNumber="18">Head depth</th>
<td id="76A86426FF88ABECFD4BFB73FD0C57B4" box="[644,746,1153,1175]" gridcol="1" gridrow="23" pageId="17" pageNumber="18">13.815.1</td>
<td id="76A86426FF88ABECFBDBFB73FB9D57B4" box="[1044,1147,1153,1175]" gridcol="2" gridrow="23" pageId="17" pageNumber="18">14.3±0.56</td>
</tr>
<tr id="35790D5AFF88ABECFF50FB58FB9D57E3" box="[159,1147,1194,1216]" gridrow="24" pageId="17" pageNumber="18" rowspan-1="1" rowspan-2="1">
<th id="76A86426FF88ABECFF50FB58FE3B57E3" box="[159,477,1194,1216]" gridcol="0" gridrow="24" pageId="17" pageNumber="18">%HL</th>
</tr>
<tr id="35790D5AFF88ABECFF50FB26FB9D57C9" box="[159,1147,1236,1258]" gridrow="25" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FB26FE3B57C9" box="[159,477,1236,1258]" gridcol="0" gridrow="25" pageId="17" pageNumber="18">Snout length</th>
<td id="76A86426FF88ABECFD4BFB26FD0C57C9" box="[644,746,1236,1258]" gridcol="1" gridrow="25" pageId="17" pageNumber="18">4447</td>
<td id="76A86426FF88ABECFBDBFB26FB9D57C9" box="[1044,1147,1236,1258]" gridcol="2" gridrow="25" pageId="17" pageNumber="18">46±1.0</td>
</tr>
<tr id="35790D5AFF88ABECFF50FB0FFB9D5630" box="[159,1147,1277,1299]" gridrow="26" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FB0FFE3B5630" box="[159,477,1277,1299]" gridcol="0" gridrow="26" pageId="17" pageNumber="18">Interorbital distance</th>
<td id="76A86426FF88ABECFD4BFB0FFD0C5630" box="[644,746,1277,1299]" gridcol="1" gridrow="26" pageId="17" pageNumber="18">2427</td>
<td id="76A86426FF88ABECFBDBFB0FFB9D5630" box="[1044,1147,1277,1299]" gridcol="2" gridrow="26" pageId="17" pageNumber="18">25±1.2</td>
</tr>
<tr id="35790D5AFF88ABECFF50FAD4FB9D561F" box="[159,1147,1318,1340]" gridrow="27" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FAD4FE3B561F" box="[159,477,1318,1340]" gridcol="0" gridrow="27" pageId="17" pageNumber="18">Eye diameter</th>
<td id="76A86426FF88ABECFD4BFAD4FD0C561F" box="[644,746,1318,1340]" gridcol="1" gridrow="27" pageId="17" pageNumber="18">1013</td>
<td id="76A86426FF88ABECFBDBFAD4FB9D561F" box="[1044,1147,1318,1340]" gridcol="2" gridrow="27" pageId="17" pageNumber="18">12±1.0</td>
</tr>
<tr id="35790D5AFF88ABECFF50FAA2FB9D5645" box="[159,1147,1360,1382]" gridrow="28" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FAA2FE3B5645" box="[159,477,1360,1382]" gridcol="0" gridrow="28" pageId="17" pageNumber="18">Nasal barbel length</th>
<td id="76A86426FF88ABECFD4BFAA2FD0C5645" box="[644,746,1360,1382]" gridcol="1" gridrow="28" pageId="17" pageNumber="18">1520</td>
<td id="76A86426FF88ABECFBDBFAA2FB9D5645" box="[1044,1147,1360,1382]" gridcol="2" gridrow="28" pageId="17" pageNumber="18">18±2.1</td>
</tr>
<tr id="35790D5AFF88ABECFF50FA8BFB9D56AC" box="[159,1147,1401,1423]" gridrow="29" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FA8BFE3B56AC" box="[159,477,1401,1423]" gridcol="0" gridrow="29" pageId="17" pageNumber="18">Maxillary barbel length</th>
<td id="76A86426FF88ABECFD4BFA8BFD0C56AC" box="[644,746,1401,1423]" gridcol="1" gridrow="29" pageId="17" pageNumber="18">92111</td>
<td id="76A86426FF88ABECFBDBFA8BFB9D56AC" box="[1044,1147,1401,1423]" gridcol="2" gridrow="29" pageId="17" pageNumber="18">99±9.3</td>
</tr>
<tr id="35790D5AFF88ABECFF50FA50FB9D569B" box="[159,1147,1442,1464]" gridrow="30" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FA50FE3B569B" box="[159,477,1442,1464]" gridcol="0" gridrow="30" pageId="17" pageNumber="18">Inner mandibular barbel length</th>
<td id="76A86426FF88ABECFD4BFA50FD0C569B" box="[644,746,1442,1464]" gridcol="1" gridrow="30" pageId="17" pageNumber="18">3039</td>
<td id="76A86426FF88ABECFBDBFA50FB9D569B" box="[1044,1147,1442,1464]" gridcol="2" gridrow="30" pageId="17" pageNumber="18">34±4.5</td>
</tr>
<tr id="35790D5AFF88ABECFF50FA3EFB9D56C1" box="[159,1147,1484,1506]" gridrow="31" pageId="17" pageNumber="18">
<th id="76A86426FF88ABECFF50FA3EFE3B56C1" box="[159,477,1484,1506]" gridcol="0" gridrow="31" pageId="17" pageNumber="18">Outer mandibular barbel length</th>
<td id="76A86426FF88ABECFD4BFA3EFD0C56C1" box="[644,746,1484,1506]" gridcol="1" gridrow="31" pageId="17" pageNumber="18">4867</td>
<td id="76A86426FF88ABECFBDBFA3EFB9D56C1" box="[1044,1147,1484,1506]" gridcol="2" gridrow="31" pageId="17" pageNumber="18">55±8.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF60F18FF885402FF08F9D4FDA755EC" blockId="17.[151,1437,1574,1995]" pageId="17" pageNumber="18">Head depressed and narrow, triangular when viewed laterally and with acute snout margin when viewed from above. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill opening broad, extending from immediately ventral to post-temporal to isthmus. First branchial arch with 2+6 (1) rakers. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head.</paragraph>
<paragraph id="8BF60F18FF885402FF08F928FEDD547C" blockId="17.[151,1437,1574,1995]" pageId="17" pageNumber="18">Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to midway between its base and anterior orbital margin. Inner mandibular barbel extending to two-thirds of distance between its base and that of pectoral spine. Outer mandibular barbel extending to base of pectoral spine.</paragraph>
<paragraph id="8BF60F18FF885402FF08F898FB1C54E8" blockId="17.[151,1437,1574,1995]" pageId="17" pageNumber="18">Mouth inferior, premaxillary tooth band almost wholly exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in single broad semilunate band. Dentary teeth in a single crescentic band, consisting of two separate halves tightly bound at midline.</paragraph>
<caption id="DF365F90FF8B5401FF58FA17FE00553A" httpUri="https://zenodo.org/record/266204/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,193,1486]" targetPageId="18">
<paragraph id="8BF60F18FF8B5401FF58FA17FE00553A" blockId="18.[151,1437,1508,1561]" pageId="18" pageNumber="19">
<emphasis id="B93DD30AFF8B5401FF58FA17FEC656D9" bold="true" box="[151,288,1509,1531]" pageId="18" pageNumber="19">FIGURE 10.</emphasis>
<taxonomicName id="4C49749BFF8B5401FEE8FA16FE1956D9" box="[295,511,1508,1530]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8B5401FEE8FA16FE1956D9" box="[295,511,1508,1530]" italics="true" pageId="18" pageNumber="19">Glyptothorax exodon</emphasis>
</taxonomicName>
, BMNH 1982.3.29.190, paratype, 63.2 mm SL; Borneo: Kalimantan Barat, Sungai Pinoh. Dorsal, lateral and ventral views.
</paragraph>
</caption>
<paragraph id="8BF60F18FF8B5401FF08F9B7FF2C5585" blockId="18.[151,1436,1605,1990]" pageId="18" pageNumber="19">
Thoracic adhesive apparatus consisting of keratinized striae in an elliptical field (
<figureCitation id="1372139DFF8B5401FB4BF9B7FB2C557D" box="[1156,1226,1605,1630]" captionStart="FIGURE 3" captionStartId="6.[151,250,1784,1806]" captionTargetBox="[228,1361,193,1761]" captionTargetId="figure@6.[228,1362,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. Thoracic adhesive apparatus of: a. Glyptothorax amnestus, ZRC 54231, paratype, 51.2 mm SL; b. G. decussatus, MZB 17216, holotype, 74.5 mm SL; c. G. exodon, BMNH 1982.3. 29.190, paratype, 63.2 mm SL; d. G. famelicus, USNM 193015, holotype, 73.3 mm SL; e. G. f u s c u s, ZRC 42123, 54.3 mm SL; f. G. keluk, MZB 17217, holotype, 52.5 mm SL; g. G. ketambe, MZB 8694, holotype, 100.0 mm SL; h. G. major, ZRC 38761, 59.1 mm SL; i. G. nieuwenhuisi, MZB 16475, 61.0 mm SL; j. G. pictus, MZB 17218, holotype, 54.2 mm SL; k. G. platypogon, ZRC 48376, 56.3 mm SL; l. G. platypogonides, UF 166626, 52.4 mm SL; m. G. plectilis, ZRC 46420, paratype, 44.3 mm SL; n. G. prashadi, CMK 25369, 50.7 mm SL; o. G. robustus, ZRC 48377, 59.9 mm SL; p. G. schmidti, CMK 4536, 74.6 mm SL; q. G. stibaros, MZB 17215, holotype, 55.3 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/266202/files/figure.png" pageId="18" pageNumber="19">Fig. 3</figureCitation>
c) extending from isthmus to posterior limit of pectoral-fin base. Anteromedial striae absent. Narrow, v-shaped medial pit on posterior half.
</paragraph>
<paragraph id="8BF60F18FF8B5401FF08F940FC4E54E5" blockId="18.[151,1436,1605,1990]" pageId="18" pageNumber="19">Dorsal fin located above anterior third of body, with I,6 (7) rays; fin margin concave; spine short and gently curved. Adipose fin with anterior margin concave and posterior margin straight. Caudal fin strongly forked, with equal upper and lower lobes and i,7,8,i (7) principal rays. Procurrent rays symmetrical and extending only slightly anterior to fin base. Anal-fin base ventral to adipose-fin origin. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with iv,8 (2) or iv,9* (5) rays. Pelvic-fin origin immediately posterior to vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (7) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,8 (7) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 1012 serrations.</paragraph>
<paragraph id="8BF60F18FF8A5400FF08FF65FB6E52D6" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
<emphasis id="B93DD30AFF8A5400FF08FF65FEAA5393" bold="true" box="[199,332,151,176]" pageId="19" pageNumber="20">Coloration.</emphasis>
In 70% ethanol: Dorsal and lateral surfaces of head, and body brown to brownish gray, fading to beige on ventral surfaces. A dark, diffuse midlateral stripe running along lateral line; lateral line without such pigmentation in some individuals, causing it to show up as paler stripe within darker stripe. Body with three long, elliptical pale patches: first above lateral line and spanning region between dorsal and adipose fins, second on anterior half of caudal peduncle above lateral line and third on anterior half of caudal peduncle below lateral line. Dorsal fin with brown base and broad subdistal triangular brown mark. Pectoral fin hyaline, with brown melanophores forming a diffuse broad, transverse band subdistally. Pelvic fin hyaline, with faint transverse brown band in middle. Anal fin with brown base and brown melanophores forming V-shaped subdistal mark on anterior half of fin; rest of fin hyaline. Adipose fin brown to brownish gray, with hyaline distal margin. Caudal fin brown to brownish gray, tips and posterior margins of lobes hyaline. Maxillary barbel brown to brownish gray dorsally, beige ventrally.
</paragraph>
<paragraph id="8BF60F18FF8A5400FF08FE0DFEFC511E" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
<emphasis id="B93DD30AFF8A5400FF08FE0DFECC513B" bold="true" box="[199,298,511,536]" pageId="19" pageNumber="20">Habitat.</emphasis>
<taxonomicName id="4C49749BFF8A5400FEFEFDF2FDF8513B" box="[305,542,512,536]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FEFEFDF2FDF8513B" box="[305,542,512,536]" italics="true" pageId="19" pageNumber="20">Glyptothorax exodon</emphasis>
</taxonomicName>
is found in swift, clear streams with a substrate consisting of gravel, cobbles or large rocks.
</paragraph>
</subSubSection>
<subSubSection id="C3535C93FF8A5400FF08FDB5FF3E54FF" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BF60F18FF8A5400FF08FDB5FC3651EE" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
<emphasis id="B93DD30AFF8A5400FF08FDB5FE865143" bold="true" box="[199,352,583,608]" pageId="19" pageNumber="20">Distribution.</emphasis>
Known only from the upper reaches of the Kapuas River drainage in western Borneo (
<figureCitation id="1372139DFF8A5400FA87FDB5FA685143" box="[1352,1422,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,250,1990,2012]" captionTargetBox="[170,1433,949,1972]" captionTargetId="figure@9.[170,1433,949,1972]" captionTargetPageId="9" captionText="FIGURE 5. Map showing collection localities of Glyptothorax amnestus, G. decussatus, G. exodon and G. famelicus." httpUri="https://zenodo.org/record/165411/files/figure.png" pageId="19" pageNumber="20">Fig. 5</figureCitation>
).
<taxonomicName id="4C49749BFF8A5400FF58FD9EFE6C51A7" box="[151,394,620,644]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FF58FD9EFE6C51A7" box="[151,394,620,644]" italics="true" pageId="19" pageNumber="20">Glyptothorax exodon</emphasis>
</taxonomicName>
has been collected from Sungai Pinoh (a southern tributary of the Kapuas River), the headwaters of the northern tributaries in the Bentuang-Karimun National Park, and Putussibau and is expected to occur in other headwater tributaries in the Kapuas River drainage as well.
</paragraph>
<paragraph id="8BF60F18FF8A5400FF08FD25FAD65716" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
<emphasis id="B93DD30AFF8A5400FF08FD25FE8B51D3" bold="true" box="[199,365,727,752]" pageId="19" pageNumber="20">Comparisons.</emphasis>
Besides the exposure of the premaxillary teeth,
<taxonomicName id="4C49749BFF8A5400FC57FD2AFBEF51D3" box="[920,1033,728,752]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FC57FD2AFBEF51D3" box="[920,1033,728,752]" italics="true" pageId="19" pageNumber="20">G. exodon</emphasis>
</taxonomicName>
differs from all Sundaic congeners except for
<taxonomicName id="4C49749BFF8A5400FEDCFD0EFE785037" box="[275,414,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8A5400FEDCFD0EFE785037" box="[275,414,764,788]" italics="true" pageId="19" pageNumber="20">G. famelicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8A5400FE61FD0EFD855037" box="[430,611,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFF8A5400FE61FD0EFD855037" box="[430,611,764,788]" italics="true" pageId="19" pageNumber="20">G. nieuwenhuisi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8A5400FDBCFD0EFCD95037" box="[627,831,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="platypogonides">
<emphasis id="B93DD30AFF8A5400FDBCFD0EFCD95037" box="[627,831,764,788]" italics="true" pageId="19" pageNumber="20">G. platypogonides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8A5400FC81FD0EFC285037" box="[846,974,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B93DD30AFF8A5400FC81FD0EFC285037" box="[846,974,764,788]" italics="true" pageId="19" pageNumber="20">G. robustus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8A5400FC10FD0EFBB85037" box="[991,1118,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8A5400FC10FD0EFBB85037" box="[991,1118,764,788]" italics="true" pageId="19" pageNumber="20">G. schmidti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8A5400FB56FD0EFAF45037" box="[1177,1298,764,788]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="stibaros">
<emphasis id="B93DD30AFF8A5400FB56FD0EFAF45037" box="[1177,1298,764,788]" italics="true" pageId="19" pageNumber="20">G. stibaros</emphasis>
</taxonomicName>
in having a more slender caudal peduncle (6.87.3% SL vs. 7.411.4). It is further distinguished from
<taxonomicName id="4C49749BFF8A5400FB75FCD2FAAE501B" box="[1210,1352,800,824]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="famelicus">
<emphasis id="B93DD30AFF8A5400FB75FCD2FAAE501B" box="[1210,1352,800,824]" italics="true" pageId="19" pageNumber="20">G. famelicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8A5400FA4BFCD2FF10507F" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B93DD30AFF8A5400FA4BFCD2FF10507F" italics="true" pageId="19" pageNumber="20">G. schmidti</emphasis>
</taxonomicName>
in having a larger eye (diameter 1013% HL vs. 5.10) and a mottled body lacking a prominent pale midlateral stripe (vs. a uniform dark brown body with a prominent pale midlateral stripe), from
<taxonomicName id="4C49749BFF8A5400FB03FC9AFA9950A3" box="[1228,1407,872,896]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFF8A5400FB03FC9AFA9950A3" box="[1228,1407,872,896]" italics="true" pageId="19" pageNumber="20">G. nieuwenhuisi</emphasis>
</taxonomicName>
in having a narrower head (16.018.1% SL vs. 18.521.4) and a longer post-adipose distance (21.524.0% SL vs. 16.819.7), from
<emphasis id="B93DD30AFF8A5400FEACFC42FE0650EB" box="[355,480,944,968]" italics="true" pageId="19" pageNumber="20">G. ro b u s t u s</emphasis>
and
<emphasis id="B93DD30AFF8A5400FDEDFC42FD7F50EB" box="[546,665,944,968]" italics="true" pageId="19" pageNumber="20">G. s t i b a ro s</emphasis>
in having a larger eye (diameter 1013% HL vs. 710), a longer dorsal-to-adipose distance (21.926.3% SL vs. 15.121.3) and a more contrasting color pattern on the body (compare
<figureCitation id="1372139DFF8A5400FEC3FC05FE8A5733" box="[268,364,1015,1040]" captionStart="FIGURE 10" captionStartId="18.[151,250,1509,1531]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@18.[151,1436,193,1487]" captionTargetPageId="18" captionText="FIGURE 10. Glyptothorax exodon, BMNH 1982.3. 29.190, paratype, 63.2 mm SL; Borneo: Kalimantan Barat, Sungai Pinoh. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/266204/files/figure.png" pageId="19" pageNumber="20">Figs. 10</figureCitation>
,
<figureCitation id="1372139DFF8A5400FEB3FC0AFE7E5733" box="[380,408,1016,1040]" captionStart="FIGURE 37" captionStartId="75.[151,250,1957,1979]" captionTargetBox="[151,1434,445,1935]" captionTargetId="figure@75.[151,1436,445,1936]" captionTargetPageId="75" captionText="FIGURE 37. Glyptothorax robustus, ZRC 48377, 59.9 mm SL; Java: Jawa Barat, Cisadane River. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/165442/files/figure.png" pageId="19" pageNumber="20">37</figureCitation>
and
<figureCitation id="1372139DFF8A5400FE18FC0AFE105733" box="[471,502,1016,1040]" captionStart="FIGURE 39" captionStartId="83.[151,250,1896,1918]" captionTargetBox="[151,1434,193,1856]" captionTargetId="figure@83.[151,1436,193,1857]" captionTargetPageId="83" captionText="FIGURE 39. Glyptothorax stibaros, MZB 17215, holotype, 55.3 mm SL; Borneo: Kalimantan Barat, Sanggau. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/266216/files/figure.png" pageId="19" pageNumber="20">39</figureCitation>
), from
<taxonomicName id="4C49749BFF8A5400FD80FC0AFD375733" box="[591,721,1016,1040]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B93DD30AFF8A5400FD80FC0AFD375733" box="[591,721,1016,1040]" italics="true" pageId="19" pageNumber="20">G. robustus</emphasis>
</taxonomicName>
in having (vs. lacking) a medial pit in the thoracic adhesive apparatus, and from
<taxonomicName id="4C49749BFF8A5400FEB2FBEEFE135717" box="[381,501,1052,1076]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="stibaros">
<emphasis id="B93DD30AFF8A5400FEB2FBEEFE135717" box="[381,501,1052,1076]" italics="true" pageId="19" pageNumber="20">G. stibaros</emphasis>
</taxonomicName>
in having a smooth (vs. serrated) posterior margin of the dorsal-fin spine.
</paragraph>
<paragraph id="8BF60F18FF8A5400FF08FBCDFB375613" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
There are three other congeners known from Borneo besides
<taxonomicName id="4C49749BFF8A5400FCBEFBB2FBC4577B" box="[881,1058,1088,1112]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="nieuwenhuisi">
<emphasis id="B93DD30AFF8A5400FCBEFBB2FBC4577B" box="[881,1058,1088,1112]" italics="true" pageId="19" pageNumber="20">G. nieuwenhuisi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8A5400FB96FBB2FB36577B" box="[1113,1232,1088,1112]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="stibaros">
<emphasis id="B93DD30AFF8A5400FB96FBB2FB36577B" box="[1113,1232,1088,1112]" italics="true" pageId="19" pageNumber="20">G. stibaros</emphasis>
</taxonomicName>
(which have been compared with
<taxonomicName id="4C49749BFF8A5400FE8AFB96FE53575F" box="[325,437,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FE8AFB96FE53575F" box="[325,437,1124,1148]" italics="true" pageId="19" pageNumber="20">G. exodon</emphasis>
</taxonomicName>
in the diagnosis and above):
<taxonomicName id="4C49749BFF8A5400FD30FB96FC71575F" box="[767,919,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="decussatus">
<emphasis id="B93DD30AFF8A5400FD30FB96FC71575F" box="[767,919,1124,1148]" italics="true" pageId="19" pageNumber="20">G. decussatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C49749BFF8A5400FC6AFB96FBE1575F" box="[933,1031,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="major">
<emphasis id="B93DD30AFF8A5400FC6AFB96FBE1575F" box="[933,1031,1124,1148]" italics="true" pageId="19" pageNumber="20">G. major</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49749BFF8A5400FBF1FB96FB79575F" box="[1086,1183,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B93DD30AFF8A5400FBF1FB96FB79575F" box="[1086,1183,1124,1148]" italics="true" pageId="19" pageNumber="20">G. pictus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C49749BFF8A5400FB61FB96FA7A575F" box="[1198,1436,1124,1148]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FB61FB96FA7A575F" box="[1198,1436,1124,1148]" italics="true" pageId="19" pageNumber="20">Glyptothorax exodon</emphasis>
</taxonomicName>
further differs from these three species in having a longer post-adipose distance (21.524.0% SL vs. 13.619.8); it is additionally distinguished from
<taxonomicName id="4C49749BFF8A5400FDD5FB5EFD5557E7" box="[538,691,1196,1220]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="decussatus">
<emphasis id="B93DD30AFF8A5400FDD5FB5EFD5557E7" box="[538,691,1196,1220]" italics="true" pageId="19" pageNumber="20">G. decussatus</emphasis>
</taxonomicName>
and
<emphasis id="B93DD30AFF8A5400FD24FB5EFCAC57E7" box="[747,842,1196,1220]" italics="true" pageId="19" pageNumber="20">G. m a j o r</emphasis>
in having a larger eye (diameter 1013% HL vs. 7 10), narrower head (16.018.1% SL vs. 19.623.8) and lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, from
<taxonomicName id="4C49749BFF8A5400FE74FB06FDB3562F" box="[443,597,1268,1292]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="decussatus">
<emphasis id="B93DD30AFF8A5400FE74FB06FDB3562F" box="[443,597,1268,1292]" italics="true" pageId="19" pageNumber="20">G. decussatus</emphasis>
</taxonomicName>
in having a longer post-adipose distance (21.524.0% SL vs. 16.217.5), and from
<taxonomicName id="4C49749BFF8A5400FECBFAEAFE83560C" box="[260,357,1304,1327]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B93DD30AFF8A5400FECBFAEAFE83560C" box="[260,357,1304,1327]" italics="true" pageId="19" pageNumber="20">G. pictus</emphasis>
</taxonomicName>
in having a longer dorsal-to-adipose distance (21.926.3% SL vs. 14.724.1).
</paragraph>
<paragraph id="8BF60F18FF8A5400FF08FACEFF3E54FF" blockId="19.[151,1437,151,2012]" pageId="19" pageNumber="20">
Among the other Sundaic congeners,
<taxonomicName id="4C49749BFF8A5400FD47FACEFD1B5677" box="[648,765,1340,1364]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B93DD30AFF8A5400FD47FACEFD1B5677" box="[648,765,1340,1364]" italics="true" pageId="19" pageNumber="20">G. exodon</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C49749BFF8A5400FC34FACEFB615677" box="[1019,1159,1340,1364]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="amnestus">
<emphasis id="B93DD30AFF8A5400FC34FACEFB615677" box="[1019,1159,1340,1364]" italics="true" pageId="19" pageNumber="20">G. amnestus</emphasis>
</taxonomicName>
in having a larger eye (diameter 1013% HL vs. 610), a narrower head (16.018.1% SL vs. 19.923.0), a more slender body (depth at anus 13.815.8% SL vs. 15.320.6) and a longer post-adipose distance (21.524.0% SL vs. 15.819.7), from
<taxonomicName id="4C49749BFF8A5400FA4BFA76FF3B56E3" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="B93DD30AFF8A5400FA4BFA76FF3B56E3" italics="true" pageId="19" pageNumber="20">G. fuscus</emphasis>
</taxonomicName>
in having a narrower head (16.018.1% SL vs. 18.323.3), a concave (vs. straight) margin of the dorsal fin, a longer dorsal-to-adipose distance (21.926.3% SL vs. 14.820.6), a longer post-adipose distance (21.524.0% SL vs. 15.720.8) and a more slender body (depth at anus 13.815.8% SL vs. 15.620.7), from
<emphasis id="B93DD30AFF8A5400FB77FA02FAF6552B" box="[1208,1296,1520,1544]" italics="true" pageId="19" pageNumber="20">G. ke l u k</emphasis>
in having a narrower head (16.018.1% SL vs. 18.820.4), a longer post-adipose distance (21.524.0% SL vs. 15.216.7) and a straight (vs. convex) dorsoposterior margin of the adipose fin, and from
<emphasis id="B93DD30AFF8A5400FBC4F9CAFB6E5573" box="[1035,1160,1592,1616]" italics="true" pageId="19" pageNumber="20">G. ke t a m b e</emphasis>
in having a larger eye (diameter 1013% HL vs. 810), a longer prepectoral length (20.625.4% SL vs. 18.220.6), a greater predorsal length (36.739.2% SL vs. 33.335.9), a greater prepelvic length (48.653.0% SL vs. 45.447.8), a longer dorsalfin spine (15.020.0% SL vs. 12.514.8) and a body with a mottled color pattern (vs. uniformly dark with pale middorsal and midlateral stripes). It is further distinguished from
<taxonomicName id="4C49749BFF8A5400FC98F93AFC1355C3" box="[855,1013,1736,1760]" family="Sisoridae" genus="Glyptosternum" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="platypogon">
<emphasis id="B93DD30AFF8A5400FC98F93AFC1355C3" box="[855,1013,1736,1760]" italics="true" pageId="19" pageNumber="20">G. platypogon</emphasis>
</taxonomicName>
in having a larger eye (diameter 10 13% HL vs. 79), a narrower head (16.018.1% SL vs. 19.422.3), a straight (vs. convex) dorsoposterior margin of the adipose fin, a longer post-adipose distance (21.524.0% SL vs. 14.617.6), a longer caudal fin (28.031.0% SL vs. 20.127.0) and a body with a mottled (vs. uniform) color pattern, from
<taxonomicName id="4C49749BFF8A5400FC2CF8C6FBBD546F" box="[995,1115,1844,1868]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="B93DD30AFF8A5400FC2CF8C6FBBD546F" box="[995,1115,1844,1868]" italics="true" pageId="19" pageNumber="20">G. plectilis</emphasis>
</taxonomicName>
in having a narrower (16.0 18.1% SL vs. 18.423.5) and more slender (13.815.1% SL vs. 15.219.4) head and a longer post-adipose distance (21.524.0% SL vs. 14.017.6), and from
<taxonomicName id="4C49749BFF8A5400FDBFF88EFD1754B7" box="[624,753,1916,1940]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="prashadi">
<emphasis id="B93DD30AFF8A5400FDBFF88EFD1754B7" box="[624,753,1916,1940]" italics="true" pageId="19" pageNumber="20">G. prashadi</emphasis>
</taxonomicName>
in having a narrower head (16.018.1% SL vs. 20.222.8), a concave (vs. straight) margin of the dorsal fin and a more slender body (depth at anus 13.815.8% SL vs. 15.8 18.8).
</paragraph>
</subSubSection>
</treatment>
</document>