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<mods:title id="EE3A7E75F051D18E8BA5B09936992E7B">Description of female nymphal instars and adult female of Kermes echinatus Balachowsky (Hemiptera, Coccoidea, Kermesidae) based on specimens from Crete and mainland Greece, with a discussion on geographical variation</mods:title>
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<treatment id="03DE1418FFE94302FF1FFB5E7115BB4E" ID-DOI="http://doi.org/10.5281/zenodo.5695417" ID-GBIF-Taxon="127670609" ID-Zenodo-Dep="5695417" LSID="urn:lsid:plazi:treatment:03DE1418FFE94302FF1FFB5E7115BB4E" httpUri="http://treatment.plazi.org/id/03DE1418FFE94302FF1FFB5E7115BB4E" lastPageId="4" lastPageNumber="64" pageId="1" pageNumber="62">
<subSubSection id="C36DF685FFE94307FF1FFB5E7206BC61" box="[151,620,1275,1301]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8BC8A50EFFE94307FF1FFB5E7206BC61" blockId="1.[151,620,1275,1301]" box="[151,620,1275,1301]" pageId="1" pageNumber="62">
<heading id="D0801262FFE94307FF1FFB5E7206BC61" bold="true" box="[151,620,1275,1301]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<taxonomicName id="4C77DE8DFFE94307FF1FFB5E7206BC61" ID-CoL="3R64Z" authority="Balachowsky" authorityName="Balachowsky" box="[151,620,1275,1301]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFE94307FF1FFB5E7206BC61" bold="true" box="[151,620,1275,1301]" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF1FFB5E71AABC61" bold="true" box="[151,448,1275,1301]" italics="true" pageId="1" pageNumber="62">KERMES ECHINATUS</emphasis>
Balachowsky
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36DF685FFE94304FF1FFAE7733FBB6C" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="reference_group">
<paragraph id="8BC8A50EFFE94307FF1FFAE772EFBC23" blockId="1.[151,645,1345,1367]" box="[151,645,1345,1367]" pageId="1" pageNumber="62">
<treatmentCitationGroup id="AB678220FFE94307FF1FFAE772EFBC23" box="[151,645,1345,1367]" pageId="1" pageNumber="62">
<taxonomicName id="4C77DE8DFFE94307FF1FFAE77221BC23" ID-CoL="3R64Z" authority="Balachowsky, 1953: 183" authorityName="Balachowsky" authorityPageNumber="183" authorityYear="1953" box="[151,587,1345,1367]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFE94307FF1FFAE77126BC23" box="[151,332,1345,1367]" italics="true" pageId="1" pageNumber="62">Kermes echinatus</emphasis>
<treatmentCitation id="0AD6831FFFE94307FEDAFAE47221BC23" author="Balachowsky" box="[338,587,1345,1367]" page="183" pageId="1" pageNumber="62" year="1953">
<bibRefCitation id="EFE6D8FFFFE94307FEDAFAE47271BC23" author="Balachowsky" box="[338,539,1345,1367]" pageId="1" pageNumber="62" refString="Balachowsky, A. S. (1953) Sur les Kermes Boitard (Hom. Coccoidea) des chenes du bassin oriental de la Mediterranee. Revue de Pathologie Vegetale et d'Entomologie Agricole de France, 32, 181 - 189." type="journal article" year="1953">Balachowsky, 1953</bibRefCitation>
: 183
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184.
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</paragraph>
<paragraph id="8BC8A50EFFE94307FF1FFA2173BABCB5" blockId="1.[151,1437,1412,2018]" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF1FFA2171A5BCE9" bold="true" box="[151,463,1412,1437]" pageId="1" pageNumber="62">FIRST INSTAR NYMPH</emphasis>
(crawler) (
<figureCitation id="134CB98BFFE94307FDC7FA2172E6BCE9" box="[591,652,1412,1437]" captionStart="FIGURE 1" captionStartId="5.[151,250,1958,1980]" captionTargetBox="[281,1302,204,1922]" captionTargetId="figure@5.[272,1314,195,1934]" captionTargetPageId="5" captionText="FIGURE 1. Kermes echinatus Balachowsky: first instar nymph." httpUri="https://zenodo.org/record/287344/files/figure.png" pageId="1" pageNumber="62">Fig.1</figureCitation>
). Described from
<specimenCount id="9D716E87FFE94307FCD4FA217382BCE9" box="[860,1000,1412,1437]" pageId="1" pageNumber="62" type="generic">5 specimens</specimenCount>
in good condition; details checked on another
<specimenCount id="9D716E87FFE94307FF7AFA0D71E0BCB5" box="[242,394,1448,1473]" pageId="1" pageNumber="62" type="generic">20 specimens</specimenCount>
(11 from Crete, 1 from Athens, 13 from Kalamata).
</paragraph>
<paragraph id="8BC8A50EFFE94307FF4FFA69736CBC91" blockId="1.[151,1437,1412,2018]" box="[199,774,1484,1509]" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF4FFA6971E0BC91" bold="true" box="[199,394,1484,1509]" pageId="1" pageNumber="62">Living specimen</emphasis>
: oval and almost flat, orange-red.
</paragraph>
<paragraph id="8BC8A50EFFE94307FF4FFA55708CBF59" blockId="1.[151,1437,1412,2018]" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF4FFA5571C6BF7D" bold="true" box="[199,428,1520,1545]" pageId="1" pageNumber="62">Mounted specimen</emphasis>
: cuticle membranous, body oval, 500 (440 640) Μm long, 230 (200300) Μm wide at thorax.
</paragraph>
<paragraph id="8BC8A50EFFE94307FF4FF99D7319BE71" blockId="1.[151,1437,1412,2018]" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF4FF99D7145BF25" bold="true" box="[199,303,1592,1617]" pageId="1" pageNumber="62">Dorsum.</emphasis>
Marginal setae 3234 on each side in two lines almost superimposed, each seta spine-like and slightly bent, 818 Μm long, basal socket 46 Μm wide; usually with a line of longer spines 17 (1618) Μm long alternating with a line of smaller spines, 11 (814) Μm long, (plate 1, b, c, d). Dorsal setae: four trichoid setae, each 79 Μm long, in a submedial longitudinal row on each side of head and thorax. Microtubular ducts detectable near each inter-segmental fold and present also on head. Anal lobes small, lightly sclerotized, each with two spine-like setae on inner margin. Apical setae each 216 (210224) Μm long.
</paragraph>
<paragraph id="8BC8A50EFFE94304FF4FF8B5733FBB6C" blockId="1.[151,1437,1412,2018]" lastBlockId="2.[151,1437,151,2015]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
<emphasis id="B903791CFFE94307FF4FF8B5717EBE5D" bold="true" box="[199,276,1808,1833]" pageId="1" pageNumber="62">Venter</emphasis>
. Dermal spinules present medially on abdomen and thorax. Eyes found near margin. Antennae 6- segmented, each 9496 Μm long; scape with 2 trichoid seta; pedicel with 2 trichoid setae; 3rd segment with 1 trichoid seta; 4th segment with one chaetic seta; 5th with 2 trichoid and 1 chaetic setae; apical segment with 4 or 5 long trichoid setae and 2 chaetic setae. One small trilocular pore present near the scape of each antenna. Labium triangular in shape, 3-segmented, 6667 Μm long, basal and second segments each with one pair of setae, third segment with 4 pairs. Stylets longer than body, looped within a crumena, shorter arm extending anteriorly as far as anterior coxa. Legs well developed, with two diamond-shaped sensilla on each trochanter. Tarsal and claw digituli knobbed, longer than claw. Thoracic spiracles small and narrow, with one associated spiracular pore, each about 3.5 Μm wide with 5, rarely 7, loculi; two bilocular pores, circular and about 23 Μm wide, present on body margin between spiracles; one trilocular (rarely pentalocular) pore present near margin of prothorax and another on body margin opposite anterior spiracle. Pairs of trilocular pores, each about 3 Μm wide, found medially on head (one pair), on each thorax segment and on abdominal segments VVII. With 3 pairs of trichoid setae present medially on frons and one pair of short setae medially on each thoracic segment; other trichoid setae in 6 longitudinal rows on abdominal segments: setae in medial rows each 1016 Μm long; sub-medial setae each 810 Μm long and submarginal setae each 4.5 Μm long. Anal lobes small, anal ring horse-shoe shaped, 14 Μm wide, with a few cells and 6 setae, each 1012 Μm long; with one anterolateral seta on each side of anal ring, each 10 Μm long, plus one postero-lateral seta on each side of anal ring, each 16 Μm long.
</paragraph>
</subSubSection>
<subSubSection id="C36DF685FFEA4305FF4FFD81732FBF75" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63" type="discussion">
<paragraph id="8BC8A50EFFEA4304FF4FFD81708EBB84" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFD817125BB49" bold="true" box="[199,335,548,573]" pageId="2" pageNumber="63">Comments.</emphasis>
Among the Mediterranean and European
<taxonomicName id="4C77DE8DFFEA4304FCCAFD8073FDBB48" box="[834,919,549,572]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B903791CFFEA4304FCCAFD8073FDBB48" box="[834,919,549,572]" italics="true" pageId="2" pageNumber="63">Kermes</emphasis>
</taxonomicName>
, the first-instar nymphs of
<emphasis id="B903791CFFEA4304FB57FD80709EBB14" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="4C77DE8DFFEA4304FB57FD807518BB48" box="[1247,1394,548,572]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="echinatus">K. echinatus</taxonomicName>
,
<taxonomicName id="4C77DE8DFFEA4304FA0AFD80709EBB14" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vermilio">K. vermilio</taxonomicName>
</emphasis>
and
<taxonomicName id="4C77DE8DFFEA4304FEA4FDEC72AEBB14" authority="Ben-Dov &amp; Spodek" authorityName="Ben-Dov &amp; Spodek" box="[300,708,583,608]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B903791CFFEA4304FEA4FDEC71B1BB14" box="[300,475,584,608]" italics="true" pageId="2" pageNumber="63">K. hermonensis</emphasis>
Ben-Dov &amp; Spodek
</taxonomicName>
are easily distinguishable from other first-instar
<taxonomicName id="4C77DE8DFFEA4304FB65FDEC7529BB14" box="[1261,1347,585,608]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B903791CFFEA4304FB65FDEC7529BB14" box="[1261,1347,585,608]" italics="true" pageId="2" pageNumber="63">Kermes</emphasis>
</taxonomicName>
species by the presence of marginal spine-like setae. In
<taxonomicName id="4C77DE8DFFEA4304FD34FDC87351BBF0" box="[700,827,620,644]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEA4304FD34FDC87351BBF0" box="[700,827,620,644]" italics="true" pageId="2" pageNumber="63">K. vermilio</emphasis>
</taxonomicName>
the marginal spine-like setae are conical, stouter and straight, whereas in
<taxonomicName id="4C77DE8DFFEA4304FEFFFD347269BBDC" box="[375,515,656,680]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEA4304FEFFFD347269BBDC" box="[375,515,656,680]" italics="true" pageId="2" pageNumber="63">K. echinatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77DE8DFFEA4304FDB0FD34728DBBDC" box="[568,743,656,680]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B903791CFFEA4304FDB0FD34728DBBDC" box="[568,743,656,680]" italics="true" pageId="2" pageNumber="63">K. hermonensis</emphasis>
</taxonomicName>
the marginal spine-like setae are distinctly longer and slightly bent. Moreover,
<taxonomicName id="4C77DE8DFFEA4304FEDEFD10718CBBB8" box="[342,486,692,716]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEA4304FEDEFD10718CBBB8" box="[342,486,692,716]" italics="true" pageId="2" pageNumber="63">K. echinatus</emphasis>
</taxonomicName>
has 6268 marginal spines whereas
<taxonomicName id="4C77DE8DFFEA4304FC19FD10742BBBB8" box="[913,1089,692,716]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B903791CFFEA4304FC19FD10742BBBB8" box="[913,1089,692,716]" italics="true" pageId="2" pageNumber="63">K. hermonensis</emphasis>
</taxonomicName>
differs by having only 4248 spines.
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFD5972D9BAF7" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
The presence of microtubular ducts near the inter-segmental folds is also reported by
<bibRefCitation id="EFE6D8FFFFEA4304FB09FD5875F6BA61" author="Baer" box="[1153,1436,764,789]" pageId="2" pageNumber="63" refString="Baer, R. G. &amp; Kosztarab, M. (1985) Studies on the morphology and systematics of scale insects - No. 12. II. A morphological and systematic study of the first and second instars of the family Kermesidae in the Nearctic region (Homoptera: Coccoidea). Bulletin of the Virginia Polytechnic Institute and University Research Division, 85, 119 - 261." type="journal article" year="1985">Baer &amp; Kosztarab (1985)</bibRefCitation>
in Nearctic
<taxonomicName id="4C77DE8DFFEA4304FE91FC8771F5BA4F" box="[281,415,802,827]" class="Insecta" family="Kermesidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="family">Kermesidae</taxonomicName>
1st -instars (e.g.
<taxonomicName id="4C77DE8DFFEA4304FDC7FC867307BA4F" authority="Cockerell" authorityName="Cockerell" box="[591,877,802,827]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="concinnulus">
<emphasis id="B903791CFFEA4304FDC7FC86729CBA4F" box="[591,758,803,827]" italics="true" pageId="2" pageNumber="63">K. concinnulus</emphasis>
Cockerell
</taxonomicName>
,
<taxonomicName id="4C77DE8DFFEA4304FCFFFC867400BA4F" authority="Ehrhorn" authorityName="Ehrhorn" box="[887,1130,802,827]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="B903791CFFEA4304FCFFFC86746FBA4F" box="[887,1029,803,827]" italics="true" pageId="2" pageNumber="63">K. cockerelli</emphasis>
Ehrhorn
</taxonomicName>
,
<taxonomicName id="4C77DE8DFFEA4304FBFDFC86752BBA4F" authority="Ferris" authorityName="Ferris" box="[1141,1345,802,827]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="rimarum">
<emphasis id="B903791CFFEA4304FBFDFC86749DBA4E" box="[1141,1271,803,826]" italics="true" pageId="2" pageNumber="63">K. rimarum</emphasis>
Ferris
</taxonomicName>
) but are referred as “dorsal simple pores”. However, according to the drawings of
<bibRefCitation id="EFE6D8FFFFEA4304FC5FFCED70A7BAF7" author="Baer" pageId="2" pageNumber="63" refString="Baer, R. G. &amp; Kosztarab, M. (1985) Studies on the morphology and systematics of scale insects - No. 12. II. A morphological and systematic study of the first and second instars of the family Kermesidae in the Nearctic region (Homoptera: Coccoidea). Bulletin of the Virginia Polytechnic Institute and University Research Division, 85, 119 - 261." type="journal article" year="1985">Baer &amp; Kosztarab (1985: 131, 175, 184, 197)</bibRefCitation>
, these pores are clearly microtubular ducts.
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFC2B7447BABF" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFC2B72CDBAD3" bold="true" box="[199,679,910,935]" pageId="2" pageNumber="63">SECOND-INSTAR FEMALE NYMPH</emphasis>
(
<figureCitation id="134CB98BFFEA4304FD32FC2B7369BAD3" box="[698,771,910,935]" captionStart="FIGURE 2" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[161,1430,203,1865]" captionTargetId="figure@6.[151,1436,193,1876]" captionTargetPageId="6" captionText="FIGURE 2. Kermes echinatus Balachowsky: second instar nymph, female." httpUri="https://zenodo.org/record/287345/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
). Described from
<specimenCount id="9D716E87FFEA4304FC52FC2B7402BAD3" box="[986,1128,910,935]" pageId="2" pageNumber="63" type="generic">5 specimens</specimenCount>
in good condition; details checked on another
<specimenCount id="9D716E87FFEA4304FEFFFC177268BABF" box="[375,514,946,971]" pageId="2" pageNumber="63" type="generic">5 specimens</specimenCount>
(1 from Crete, 2 from Athens, 7 from Kalamata).
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFC73719CBA9B" blockId="2.[151,1437,151,2015]" box="[199,502,982,1007]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFC7371E0BA9B" bold="true" box="[199,394,982,1007]" pageId="2" pageNumber="63">Living specimen</emphasis>
: not seen
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFC5F74DBBD67" blockId="2.[151,1437,151,2015]" box="[199,1201,1018,1043]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFC5F71C3BD67" bold="true" box="[199,425,1018,1043]" pageId="2" pageNumber="63">Mounted specimen</emphasis>
: body oval,
<quantity id="4C8F08EBFFEA4304FDBDFC5E721CBD66" box="[565,630,1019,1043]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="63" unit="mm" value="1.0">1 mm</quantity>
(0.81.2) long,
<quantity id="4C8F08EBFFEA4304FCA2FC5E73EABD66" box="[810,896,1019,1043]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="63" unit="mm" value="0.7">0.7 mm</quantity>
(0.50.9) wide (plate 1, e).
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFBBB7177BDB3" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFBBB7144BD43" bold="true" box="[199,302,1054,1079]" pageId="2" pageNumber="63">Dorsum.</emphasis>
Marginal setae conical, spine-like and stout, of one size, each 11 (1014) µm long, 6 Μm wide at base, basal socket 8 µm wide; with about 2932 on each margin. Small conical setae, each about 56 µm long, sometimes bent, sparse; also a few small trichoid setae, sparse. Tubular ducts, similar to ventral tubular ducts, present on body margin and sparsely throughout. Microtubular ducts numerous and scattered throughout. Anal lobes fused.
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FFB777264BE5F" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FFB777173BD9F" bold="true" box="[199,281,1234,1259]" pageId="2" pageNumber="63">Venter.</emphasis>
Cuticle crenulated medially on abdomen, thorax and head. Eyes located antero-laterally to each antenna, misshapen in mounted specimens. Antennae 13 segmented, sometimes with indistinct segmentation; 3- segmented antennae about 4648 Μm long and 16 Μm wide at base, scape usually with 2 very short setae, second segment with one seta; third segment with 6 or 7 chaetic setae on apex; monomerous antennae 1620 µm long, with 6 or 7 chaetic setae on apex. Frontal lobes present, “sausage-shaped”, about as long as 3-segmented antennae (plate 2, g). Labium triangular, 112 (96130) Μm long, 3 segmented, basal and second segments each with one pair of setae, third segment with 4 pairs. Legs very reduced, tubercle-like, usually with 2 or 3 short setae. Spiracles well developed, sclerotized; each anterior spiracle with peritreme 24 (1630) Μm wide, usually with 2 associated spiracular disc-pores, each 4 Μm wide with 5 loculi; each posterior spiracle with peritreme 23 (1330) Μm wide, with 1 or 2 associated spiracular pores. Bilocular pores, each about 2 Μm wide, sparsely distributed on submargin of abdomen. Tubular ducts each 12 (816) Μm long and 3 wide, with thin inner ductule, distributed in a wide marginal band, sparse on thorax and head and present on each abdominal segment; and thinner tubular ducts, with the opening of outer ductule narrower than its inner end (plate 2, p), each 16 Μm long and 1.5 Μm wide, inner ductule about 12 Μm long, present mostly on head near labium. Ventral trichoid setae of variable length, 514 Μm long, irregularly distributed on abdominal segments and sparse on head and thorax. Anal lobes small, each with one small conical seta and one apical seta, 40 (2764) Μm long. Anal ring rounded, 2430 Μm wide, with pores and 6 setae, each about 2734 Μm long.
</paragraph>
<paragraph id="8BC8A50EFFEA4304FF4FF89372D7BEAB" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="63">
<emphasis id="B903791CFFEA4304FF4FF8937125BE3B" bold="true" box="[199,335,1846,1871]" pageId="2" pageNumber="63">Comments.</emphasis>
<taxonomicName id="4C77DE8DFFEA4304FEDEF89D7275BE3B" box="[342,543,1847,1871]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEA4304FEDEF89D7275BE3B" box="[342,543,1847,1871]" italics="true" pageId="2" pageNumber="63">Kermes echinatus</emphasis>
</taxonomicName>
second-instar females mainly differ from those of
<taxonomicName id="4C77DE8DFFEA4304FBD4F89D70B2BE07" authority="Pellizzari et al., 2012" authorityName="Pellizzari et al." authorityYear="2012" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEA4304FBD4F89D74B3BE3B" box="[1116,1241,1847,1871]" italics="true" pageId="2" pageNumber="63">K. vermilio</emphasis>
(
<bibRefCitation id="EFE6D8FFFFEA4304FB61F89370BBBE07" author="Pellizzari" pageId="2" pageNumber="63" refString="Pellizzari, G., Porcelli, F., Convertini, S. &amp; Marotta, S. (2012) Description of nymphal instars and adult female of Kermes vermilio Planchon (Hemiptera, Coccoidea, Kermesidae), with a synopsis of the European and Mediterranean species Zootaxa, 3336, 36 - 50." type="journal article" year="2012">
Pellizzari
<emphasis id="B903791CFFEA4304FAD4F89D75E7BE3B" box="[1372,1421,1847,1871]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
., 2012
</bibRefCitation>
)
</taxonomicName>
as follows (characters of
<taxonomicName id="4C77DE8DFFEA4304FE76F8FE7217BE07" box="[510,637,1883,1907]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEA4304FE76F8FE7217BE07" box="[510,637,1883,1907]" italics="true" pageId="2" pageNumber="63">K. vermilio</emphasis>
</taxonomicName>
in brackets): 13-segmented antennae (5-segmented); and with 5864 marginal spine-like setae (6874). Both species have frontal lobes and small, tubercle-like legs.
<taxonomicName id="4C77DE8DFFEA4304FB63F82575F1BEE3" box="[1259,1435,1919,1943]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B903791CFFEA4304FB63F82575F1BEE3" box="[1259,1435,1919,1943]" italics="true" pageId="2" pageNumber="63">K. hermonensis</emphasis>
</taxonomicName>
second-instar females differ from those of both
<taxonomicName id="4C77DE8DFFEA4304FD39F8067345BECF" box="[689,815,1955,1979]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEA4304FD39F8067345BECF" box="[689,815,1955,1979]" italics="true" pageId="2" pageNumber="63">K. vermilio</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77DE8DFFEA4304FCEFF806739FBECF" box="[871,1013,1955,1979]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEA4304FCEFF806739FBECF" box="[871,1013,1955,1979]" italics="true" pageId="2" pageNumber="63">K. echinatus</emphasis>
</taxonomicName>
in having 3-segmented legs and 36- segmented antennae (Spodek &amp; Ben-Dov, 2014).
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFF327520B868" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
Other described second-instar females of
<taxonomicName id="4C77DE8DFFEB4305FD17FF3C729EB9C4" box="[671,756,153,176]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B903791CFFEB4305FD17FF3C729EB9C4" box="[671,756,153,176]" italics="true" pageId="3" pageNumber="64">Kermes</emphasis>
</taxonomicName>
species from Europe and the Mediterranean are those of
<taxonomicName id="4C77DE8DFFEB4305FA0AFF3C709CB9A0" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="bytinskii">
<emphasis id="B903791CFFEB4305FA0AFF3C709CB9A0" italics="true" pageId="3" pageNumber="64">K. bytinskii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77DE8DFFEB4305FEB0FF1971DCB9A7" box="[312,438,188,211]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="quercus">
<emphasis id="B903791CFFEB4305FEB0FF1971DCB9A7" box="[312,438,188,211]" italics="true" pageId="3" pageNumber="64">K. quercus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE6D8FFFFEB4305FE42FF1E72E2B9A0" author="Sternlicht" box="[458,648,187,212]" pageId="3" pageNumber="64" refString="Sternlicht, M. (1969) Kermes bytinskii n. spec. (Coccoidea, Kermesidae) in Israel and observations on its life history. Israel Journal of Entomology, 4, 251 - 270." type="journal article" year="1969">Sternlicht, 1969</bibRefCitation>
;
<bibRefCitation id="EFE6D8FFFFEB4305FD11FF1E733DB9A0" author="Podsiadlo" box="[665,855,187,212]" pageId="3" pageNumber="64" refString="Podsiadlo, E. (2012) Morphology of second instar nymphs of Kermes quercus (Linnaeus) (Hemiptera: Kermesidae). Polskie Pismo Entomologiczne, 81, 35 - 42. http: // dx. doi. org / 10.2478 / v 10200 - 011 - 0061 - 9" type="journal article" year="2012">Podsiadlo, 2012</bibRefCitation>
; Spodek &amp; Ben-Dov, 2014) which have 5 or 6- segmented antennae, and short, 3-segmented legs; these characters are shared with the second-instar females of the Nearctic
<taxonomicName id="4C77DE8DFFEB4305FF74FEA1719BB868" authority="Ehrhorn" authorityName="Ehrhorn" box="[252,497,259,284]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="B903791CFFEB4305FF74FEA171E0B86F" box="[252,394,259,283]" italics="true" pageId="3" pageNumber="64">K. cockerelli</emphasis>
Ehrhorn
</taxonomicName>
,
<taxonomicName id="4C77DE8DFFEB4305FE73FEA1737DB868" authority="Cockerell" authorityName="Cockerell" box="[507,791,259,284]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="concinnulus">
<emphasis id="B903791CFFEB4305FE73FEA172C9B86F" box="[507,675,259,283]" italics="true" pageId="3" pageNumber="64">K. concinnulus</emphasis>
Cockerell
</taxonomicName>
and
<taxonomicName id="4C77DE8DFFEB4305FCC5FEA1752FB868" authority="Ferris (Baer &amp; Kosztarab, 1985)" authorityName="Ferris (Baer &amp; Kosztarab" authorityYear="1985" box="[845,1349,259,284]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="rimarum">
<emphasis id="B903791CFFEB4305FCC5FEA173BAB86F" box="[845,976,260,283]" italics="true" pageId="3" pageNumber="64">K. rimarum</emphasis>
Ferris (
<bibRefCitation id="EFE6D8FFFFEB4305FBAFFEA67557B868" author="Baer" box="[1063,1341,259,284]" pageId="3" pageNumber="64" refString="Baer, R. G. &amp; Kosztarab, M. (1985) Studies on the morphology and systematics of scale insects - No. 12. II. A morphological and systematic study of the first and second instars of the family Kermesidae in the Nearctic region (Homoptera: Coccoidea). Bulletin of the Virginia Polytechnic Institute and University Research Division, 85, 119 - 261." type="journal article" year="1985">Baer &amp; Kosztarab, 1985</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFE837450B817" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFE8372E4B84B" bold="true" box="[199,654,294,319]" pageId="3" pageNumber="64">THIRD-INSTAR FEMALE NYMPH</emphasis>
(
<figureCitation id="134CB98BFFEB4305FD2AFE837281B84B" box="[674,747,294,319]" captionStart="FIGURE 3" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[162,1426,203,1846]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 3. Kermes echinatus Balachowsky: third instar nymph, female." httpUri="https://zenodo.org/record/287346/files/figure.png" pageId="3" pageNumber="64">Fig. 3</figureCitation>
). Described from
<specimenCount id="9D716E87FFEB4305FC4EFE827409B84B" box="[966,1123,294,319]" pageId="3" pageNumber="64" type="generic">13 specimens</specimenCount>
in good condition; details checked on another
<specimenCount id="9D716E87FFEB4305FEFFFEEE7268B817" box="[375,514,330,355]" pageId="3" pageNumber="64" type="generic">7 specimens</specimenCount>
(3 from Crete, 6 from Athens, 11 from Kalamata).
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFECB7328B8DE" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFECB71FAB8F3" bold="true" box="[199,400,366,391]" pageId="3" pageNumber="64">Living specimen</emphasis>
: body rounded, convex, bright yellow just after moult, otherwise orange- brown, with numerous protruding white waxy tufts on dorsum (plate 1: a)
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFE1071F3B886" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFE1071C5B8BA" bold="true" box="[199,431,437,462]" pageId="3" pageNumber="64">Mounted specimen</emphasis>
: body rounded, 1.8 (1.62) mm long and 1.5 (1.41.6) mm wide, slightly tapering posteriorly (plate 1: h).
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFE587287BBF5" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFE587144BB62" bold="true" box="[199,302,509,534]" pageId="3" pageNumber="64">Dorsum.</emphasis>
Marginal setae spine-like, conical, of one
<typeStatus id="54CC1BACFFEB4305FCA4FE5B7335BB62" box="[812,863,510,534]" pageId="3" pageNumber="64">type</typeStatus>
, with 2632 setae on each margin, each seta 17 (1419) Μm long and about 9.6 Μm wide at base. Dorsal setae either small conical or trichoid, irregularly present throughout. Microtubular ducts scattered throughout. Tubular ducts, each 1516 Μm long, about 5 Μm wide, with inner ductule 14 long, rare, mostly near body margin.
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFD29740EBD94" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFD297173BBD1" bold="true" box="[199,281,652,677]" pageId="3" pageNumber="64">Venter.</emphasis>
Cuticle medially crenulated on head, thorax and abdomen. Eyes located antero-laterally to each antenna, misshapen in mounted specimens. Antennae short, monomerous, tubercle-like, about 30 Μm long and 22 Μm wide, with 2 short setae at base and a group of 58 chaetic setae, each 2227 Μm long, on apex. Frontal lobes present, not very pronounced, about as long as antennae. Labium sub-triangular, 140 Μm long and 90 wide at base, 3-segmented, basal and second segments each with one pair of setae, third segment with 4 pairs. Anterior legs each reduced to a tubercle, with 35 short setae; other pairs of legs entirely absent, or indistinct and represented only by groups 2 or 3 short setae. Spiracles well developed, each anterior peritreme about 22 Μm wide, usually with 2 (14) associated spiracular disc-pores, each about 4 Μm wide with 5 or 6 loculi; each posterior spiracle with peritreme 30 Μm wide, with 2 (03) associated spiracular disc-pores; bilocular (seldom trilocular) pores, each about 3 Μm wide, present on abdominal submargin. A few specimens with 13 isolated multilocular pores on last abdominal segments. Tubular ducts of two sizes: large ducts, each 1516 Μm long, 5 Μm wide, with inner ductule 14 Μm long, distributed in a wide marginal band, and sparsely medially on thorax and head and each abdominal segment; and thin tubular ducts, each 1416 Μm long and 23 Μm wide, with the opening of outer ductule narrower than its inner end, present medially on head and thorax (plate 2, p). Trichoid setae 618 Μm long, with basal socket 2 Μm wide, irregularly distributed on abdominal segments and sparse on head and thorax. Anal lobes fused; anal lobe setae each 44 Μm long; auxiliary setae 21 (1530) Μm long. Anal ring almost circular 3236 Μm wide, with pores and 6 setae, each 2232 Μm long; with three pairs of suranal setae, each 16 (1023) Μm long.
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFB4E7411BC1B" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFB4E7125BC70" bold="true" box="[199,335,1259,1284]" pageId="3" pageNumber="64">Comments.</emphasis>
Third-instar female
<taxonomicName id="4C77DE8DFFEB4305FDBFFB4972A9BC70" box="[567,707,1260,1284]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEB4305FDBFFB4972A9BC70" box="[567,707,1260,1284]" italics="true" pageId="3" pageNumber="64">K. echinatus</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4C77DE8DFFEB4305FC27FB497444BC70" box="[943,1070,1260,1284]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEB4305FC27FB497444BC70" box="[943,1070,1260,1284]" italics="true" pageId="3" pageNumber="64">K. vermilio</emphasis>
</taxonomicName>
mostly as follows (characters of
<taxonomicName id="4C77DE8DFFEB4305FF1FFAB57172BC53" box="[151,280,1295,1319]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEB4305FF1FFAB57172BC53" box="[151,280,1295,1319]" italics="true" pageId="3" pageNumber="64">K. vermilio</emphasis>
</taxonomicName>
in brackets): monomerous antennae (2- or 3-segmented); and with about 5060 marginal spine-like setae (132222). The third instar females of
<taxonomicName id="4C77DE8DFFEB4305FD01FA91735DBC3F" box="[649,823,1331,1355]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B903791CFFEB4305FD01FA91735DBC3F" box="[649,823,1331,1355]" italics="true" pageId="3" pageNumber="64">K. hermonensis</emphasis>
</taxonomicName>
differ from those of both
<taxonomicName id="4C77DE8DFFEB4305FBD3FA9174B3BC3F" box="[1115,1241,1331,1355]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="vermilio">
<emphasis id="B903791CFFEB4305FBD3FA9174B3BC3F" box="[1115,1241,1331,1355]" italics="true" pageId="3" pageNumber="64">K. vermilio</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77DE8DFFEB4305FA98FA9175F7BC3F" box="[1296,1437,1331,1355]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B903791CFFEB4305FA98FA9175F7BC3F" box="[1296,1437,1331,1355]" italics="true" pageId="3" pageNumber="64">K. echinatus</emphasis>
</taxonomicName>
in having 3-segmented legs and 5- or 6-segmented antennae (Spodek &amp; Ben-Dov, 2014).
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFAD87235BCCD" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
Comparison and comments on a few other non-European
<taxonomicName id="4C77DE8DFFEB4305FCECFADB73D3BCE1" box="[868,953,1406,1429]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B903791CFFEB4305FCECFADB73D3BCE1" box="[868,953,1406,1429]" italics="true" pageId="3" pageNumber="64">Kermes</emphasis>
</taxonomicName>
3rd -instars are reported in
<bibRefCitation id="EFE6D8FFFFEB4305FB65FAD8708BBCCD" pageId="3" pageNumber="64" refString="Pellizzari, G., Porcelli, F., Convertini, S. &amp; Marotta, S. (2012) Description of nymphal instars and adult female of Kermes vermilio Planchon (Hemiptera, Coccoidea, Kermesidae), with a synopsis of the European and Mediterranean species Zootaxa, 3336, 36 - 50." type="journal article">
Pellizzari
<emphasis id="B903791CFFEB4305FAE9FADB75FFBCE1" box="[1377,1429,1405,1429]" italics="true" pageId="3" pageNumber="64">et al</emphasis>
. (2012)
</bibRefCitation>
and in Spodek &amp; Ben-Dov, 2014.
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FFA61732FBF75" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FFA6171C0BCA9" bold="true" box="[199,426,1476,1501]" pageId="3" pageNumber="64">ADULT FEMALE</emphasis>
(
<figureCitation id="134CB98BFFEB4305FE34FA617269BCA9" box="[444,515,1476,1501]" captionStart="FIGURE 4" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[180,1406,193,1929]" captionTargetId="figure@8.[180,1406,193,1932]" captionTargetPageId="8" captionText="FIGURE 4. Kermes echinatus Balachowsky: adult female." httpUri="https://zenodo.org/record/287347/files/figure.png" pageId="3" pageNumber="64">Fig. 4</figureCitation>
). Described from
<specimenCount id="9D716E87FFEB4305FD5DFA61730BBCA9" box="[725,865,1476,1501]" pageId="3" pageNumber="64" type="generic">5 specimens</specimenCount>
in good condition, details checked on another
<specimenCount id="9D716E87FFEB4305FA08FA617166BF75" pageId="3" pageNumber="64" type="generic">16 specimens</specimenCount>
(9 from Crete, 2 from Athens, 10 from Kalamata).
</paragraph>
</subSubSection>
<subSubSection id="C36DF685FFEB4302FF4FF9A97115BB4E" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64" type="materials_examined">
<paragraph id="8BC8A50EFFEB4305FF4FF9A97100BF19" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FF9A971B3BF51" bold="true" box="[199,473,1548,1573]" pageId="3" pageNumber="64">Unmounted specimens</emphasis>
: pre-reproductive females, found dead on twigs, brown-yellowish, with transverse wrinkles on dorsum. Reproductive female sub-spherical, dark, with transverse wrinkles on dorsum and red eggs inside body cavity.
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FF9D27293BFC0" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FF9D271DABFE4" bold="true" box="[199,432,1655,1680]" pageId="3" pageNumber="64">Mounted specimen</emphasis>
: body of pre-reproductive female largely oval or rounded, tapering posteriorly, 2.8 (2.23.2) mm long and 2.6 (23) mm wide (plate 2, r).
</paragraph>
<paragraph id="8BC8A50EFFEB4305FF4FF91A746BBE37" blockId="3.[151,1437,151,2038]" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FF91A714DBFAC" bold="true" box="[199,295,1727,1752]" pageId="3" pageNumber="64">Dorsum</emphasis>
. Marginal setae spine-like, conical of one
<typeStatus id="54CC1BACFFEB4305FC85F965732ABFAC" box="[781,832,1728,1752]" pageId="3" pageNumber="64">type</typeStatus>
, each 16 (1420) Μm long, 68 Μm wide at base, with 2834 on each margin. Dorsal setae all conical, spine-like setae, each 1314 µm long and 56 µm wide at base, unevenly distributed over dorsum. Large tubular ducts, each about 1922 Μm long and 45 Μm wide, inner ductule 12 (816) Μm, numerous and scattered. Microtubular ducts sparse throughout.
</paragraph>
<paragraph id="8BC8A50EFFEB4302FF4FF8E87115BB4E" blockId="3.[151,1437,151,2038]" lastBlockId="4.[151,1437,151,570]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
<emphasis id="B903791CFFEB4305FF4FF8E87173BE12" bold="true" box="[199,281,1869,1894]" pageId="3" pageNumber="64">Venter.</emphasis>
Cuticle crenulated medially on abdomen and thorax. Antennae short, tubercle-like, one segmented, each 28 (2234) Μm long, 23 (2026) Μm wide, with a group of 58 chaetic setae, about 2024 Μm long, at apex. Eyes located antero-laterally to each antenna, misshapen in mounted specimens. Clypeolabral shield 234 (213250) Μm long. Labium subtriangular, 166 (160170) Μm long and 102 (90110) Μm wide, 3-segmented, basal and second segments each with one pair of setae, third segment with 4 pairs. Legs absent. Spiracles well developed and sclerotized, each anterior spiracle 60 Μm wide, each posterior spiracle 75 Μm wide. Multilocular pores (plate 2, n) each about 8 Μm wide with 1012 loculi, forming a large group near each antenna and with 1418 near each spiracle; multilocular pores also in transverse bands on all abdominal segments and around vulva. Bilocular pores each about 3 Μm long, sparse throughout. Tubular ducts of two/three sizes: larger ducts each about 1220 Μm long and about 45 Μm wide, inner ductule 816 Μm long, distributed in a wide marginal band, sparse medially on thorax and head and medially on each abdominal segment; and a thinner duct, with opening of outer ductule narrower than inner end, each about 16 Μm long and 2.43 Μm wide, more frequent than larger
<typeStatus id="54CC1BACFFEC4302FB55FECA7567B8F3" box="[1245,1293,367,391]" pageId="4" pageNumber="65">type</typeStatus>
medially on frons, around labium and medially on thorax and abdomen (plate 2, o, q). Ventral trichoid setae each 624 Μm long, irregularly distributed across abdominal segments, plus a few medially and submedially on head and thorax, these usually shorter than on abdominal segments. Anal lobes not found. Apical seta each 47 (4056) Μm long. Anal ring circular and sclerotized, 48 (4056) Μm wide, with 6 setae, each 40 (3644) long plus 3 pairs of suranal setae, each 19 (1325) Μm long.
</paragraph>
</subSubSection>
</treatment>
</document>