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<mods:title id="2165BE8A8803211DA20DF94A4B44E764">Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
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<treatment id="03DA133F7518B13E4BC2FB848EF16833" ID-DOI="http://doi.org/10.5281/zenodo.6567170" ID-GBIF-Taxon="195537729" ID-Zenodo-Dep="6567170" LSID="urn:lsid:plazi:treatment:03DA133F7518B13E4BC2FB848EF16833" httpUri="http://treatment.plazi.org/id/03DA133F7518B13E4BC2FB848EF16833" lastPageNumber="2" pageId="3" pageNumber="2">
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<heading id="D08415457518B13E4BC2FB848BFF6A17" bold="true" box="[192,500,1063,1088]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<taxonomicName id="4C73D9AA7518B13E4BC2FB848BFF6A17" ID-CoL="4M4Y" authority="Benesh, 1955" authorityName="Benesh" authorityYear="1955" box="[192,500,1063,1088]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4BC2FB848BFF6A17" bold="true" box="[192,500,1063,1088]" pageId="3" pageNumber="2">
<emphasis id="B9077E3B7518B13E4BC2FB848B306A68" bold="true" box="[192,315,1063,1087]" italics="true" pageId="3" pageNumber="2">Ganelius</emphasis>
Benesh, 1955
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<treatmentCitation id="0AD284387518B13E4BC2FBCB880C6AD7" author="Laporte, F." box="[192,519,1128,1152]" page="174" pageId="3" pageNumber="2" year="1840">
<taxonomicName id="4C73D9AA7518B13E4BC2FBCB880C6AD7" ID-CoL="4FHS" authority="Laporte, 1840: 174" authorityName="Laporte" authorityPageNumber="174" authorityYear="1840" box="[192,519,1128,1152]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4BC2FBCB8B176AD7" box="[192,284,1128,1152]" italics="true" pageId="3" pageNumber="2">Eudora</emphasis>
<bibRefCitation id="EFE2DFD87518B13E4A21FBCA880C6AD7" author="Laporte, F." box="[291,519,1128,1152]" pageId="3" pageNumber="2" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte, 1840: 174</bibRefCitation>
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(not
<taxonomicName id="4C73D9AA7518B13E4946FBCB89C66AD7" ID-CoL="4FHS" authority="Peron and Leseur, 1810" authorityName="Peron and Leseur" authorityYear="1810" box="[580,973,1128,1152]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4946FBCB88AB6AD7" box="[580,672,1128,1152]" italics="true" pageId="3" pageNumber="2">Eudora</emphasis>
Péron and Leseur, 1810
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;
<taxonomicName id="4C73D9AA7518B13E48D5FBCB8F1B6AD7" ID-CoL="4FHS" authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" box="[983,1296,1128,1152]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E48D5FBCB8E386AD7" box="[983,1075,1128,1152]" italics="true" pageId="3" pageNumber="2">Eudora</emphasis>
Rafinesque, 1815
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;
<taxonomicName id="4C73D9AA7518B13E4E19FBCB8B896AF6" ID-CoL="4FHS" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4E19FBCB8F7C6AD7" box="[1307,1399,1128,1152]" italics="true" pageId="3" pageNumber="2">Eudora</emphasis>
De Haan, 1833
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)
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<paragraph id="8BCCA2297518B13E4BC2FB0A896F6A96" blockId="3.[192,1440,1128,1249]" box="[192,868,1193,1217]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB6385077518B13E4BC2FB0A896F6A96" box="[192,868,1193,1217]" pageId="3" pageNumber="2">
<treatmentCitation id="0AD284387518B13E4BC2FB0A881E6A96" author="Benesh, B." box="[192,533,1193,1217]" page="62" pageId="3" pageNumber="2" year="1955">
<taxonomicName id="4C73D9AA7518B13E4BC2FB0A881E6A96" ID-CoL="5X3V" authority="Benesh, 1955 a: 62" authorityName="Benesh" authorityPageNumber="62" authorityYear="1955" box="[192,533,1193,1217]" class="Insecta" family="Histeridae" genus="Nagelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4BC2FB0A8B266A96" box="[192,301,1193,1217]" italics="true" pageId="3" pageNumber="2">Nagelius</emphasis>
<bibRefCitation id="EFE2DFD87518B13E4A37FB0A881E6A96" author="Benesh, B." box="[309,533,1193,1217]" pageId="3" pageNumber="2" pagination="57 - 63" refId="ref7347" refString="Benesh, B. 1955 a. On the genus Macrocrates, with notes on other stagbeetles (Coleoptera: Lucanidae). Entomological News 66 (3): 57 - 63." type="journal article" year="1955">Benesh, 1955a: 62</bibRefCitation>
</taxonomicName>
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(not
<taxonomicName id="4C73D9AA7518B13E4956FB0A89576A96" ID-CoL="5X3V" authority="Lewis, 1909" authorityName="Lewis" authorityYear="1909" box="[596,860,1193,1217]" class="Insecta" family="Histeridae" genus="Nagelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4956FB0A88CA6A96" box="[596,705,1193,1217]" italics="true" pageId="3" pageNumber="2">Nagelius</emphasis>
Lewis, 1909
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)
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</paragraph>
<paragraph id="8BCCA2297518B13E4BC2FB6A89A66AB6" blockId="3.[192,1440,1128,1249]" box="[192,941,1225,1249]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB6385077518B13E4BC2FB6A89A66AB6" box="[192,941,1225,1249]" pageId="3" pageNumber="2">
<treatmentCitation id="0AD284387518B13E4BC2FB6A881D6AB6" author="Benesh, B." box="[192,534,1225,1249]" page="70" pageId="3" pageNumber="2" year="1955">
<taxonomicName id="4C73D9AA7518B13E4BC2FB6A881D6AB6" ID-CoL="4M4Y" authority="Benesh, 1955 b: 70" authorityName="Benesh" authorityPageNumber="70" authorityYear="1955" box="[192,534,1225,1249]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4BC2FB6A8B256AB6" box="[192,302,1225,1249]" italics="true" pageId="3" pageNumber="2">Ganelius</emphasis>
<bibRefCitation id="EFE2DFD87518B13E4A37FB6A881D6AB6" author="Benesh, B." box="[309,534,1225,1249]" pageId="3" pageNumber="2" pagination="59 - 76" refId="ref7381" refString="Benesh, B. 1955 b. Some further notes on the stagbeetles, with especial reference to Figulinae (Coleoptera: Lucanidae). Transactions of the American Entomological Society 81 (2): 59 - 76." type="journal article" year="1955">Benesh, 1955b: 70</bibRefCitation>
</taxonomicName>
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(new name for
<taxonomicName id="4C73D9AA7518B13E49D7FB6A89AF6AB6" ID-CoL="5X3V" authority="Benesh" authorityName="Benesh" authorityYear="1955" box="[725,932,1225,1249]" class="Insecta" family="Histeridae" genus="Nagelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E49D7FB6A89496AB6" box="[725,834,1225,1249]" italics="true" pageId="3" pageNumber="2">Nagelius</emphasis>
Benesh
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)
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</paragraph>
</subSubSection>
<subSubSection id="C369F1A27518B13E4BC2FAAA8EF16833" pageId="3" pageNumber="2" type="description">
<paragraph id="8BCCA2297518B13E4BC2FAAA8EF16833" blockId="3.[190,1440,1289,1636]" pageId="3" pageNumber="2">
<emphasis id="B9077E3B7518B13E4BC2FAAA8B606B76" bold="true" box="[192,363,1289,1313]" pageId="3" pageNumber="2">Description.</emphasis>
<taxonomicName id="4C73D9AA7518B13E4A73FAAA8BD46B76" authorityName="Benesh" authorityYear="1955" box="[369,479,1289,1313]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4A73FAAA8BD46B76" box="[369,479,1289,1313]" italics="true" pageId="3" pageNumber="2">Ganelius</emphasis>
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are moderately sized (
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) figuline species that display distinctly sexually dimorphic mandibles in which males have a short mandible with a long, vertically produced dorsal ramus or branch, while females lack the dorsal ramus and have a simple, more-or-less tricuspid mandible. The pronotum appears smooth, bearing only fine punctures and always lacking a strongly indicated midline. In both sexes the pronotum has an anterior tubercle that varies from distinct to almost obsolete (especially in larger males that have the anterior margin produced on either side), and is generally elevated. The male genitalia of
<taxonomicName id="4C73D9AA7518B13E4945FA6889186BB4" box="[583,787,1483,1507]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9077E3B7518B13E4945FA6888BE6BB4" box="[583,693,1483,1507]" italics="true" pageId="3" pageNumber="2">Ganelius</emphasis>
species
</taxonomicName>
have an exceptionally long, thin, and gradually narrowing flagellum that varies from
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in length. The genus only occurs in Madagascar. Species within the genus vary most importantly in the form of the mandibles, shape of the ocular canthus and anterior margin of the head, and extent of punctation of the elytra. The male genitalia between species is fairly uniform but with apparent interspecific variation in the length of the flagellum.
</paragraph>
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