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<document id="9CDEA060616AD296C452DE23EBA18720" ID-DOI="10.5281/zenodo.283190" ID-GBIF-Dataset="fbc85300-bbf7-4ca8-86c6-87342096166d" ID-ISSN="1175-5326" ID-Zenodo-Dep="283190" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564767146" checkinUser="plazi" docAuthor="Vilkamaa, Pekka, Hippa, Heikki &amp; Mohrig, Werner" docDate="2012" docId="03D07408FFF8FFFCFF30FB8238FD53A0" docLanguage="en" docName="zt03591p074.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Scatopsiara (Xenopygina) exigua Vilkamaa, Hippa &amp; Mohrig, 2012, sp. n." docType="treatment" docVersion="8" lastPageNumber="70" masterDocId="FFE90C70FFF9FFFFFFA7FF94394E5107" masterDocTitle="The genus Scatopsciara Edwards (Diptera, Sciaridae) in New Caledonia, with the description of four new species" masterLastPageNumber="74" masterPageNumber="67" pageNumber="68" updateTime="1698320815747" updateUser="plazi">
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<mods:title id="84A01059FA0C61091199F2AE4CE7933A">The genus Scatopsciara Edwards (Diptera, Sciaridae) in New Caledonia, with the description of four new species</mods:title>
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<mods:namePart id="5372EDAA6859F4D21649AA00DA9E4F22">Vilkamaa, Pekka</mods:namePart>
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<mods:namePart id="76F095840730B424B09C675119204777">Hippa, Heikki</mods:namePart>
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<mods:namePart id="F282FB6B74206E2BA0CD1B62B2A34172">Mohrig, Werner</mods:namePart>
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<mods:date id="4D8C28712B08D94C3881817D0C408719">2012</mods:date>
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<treatment id="03D07408FFF8FFFCFF30FB8238FD53A0" ID-DOI="http://doi.org/10.5281/zenodo.6166086" ID-GBIF-Taxon="119557587" ID-Zenodo-Dep="6166086" LSID="urn:lsid:plazi:treatment:03D07408FFF8FFFCFF30FB8238FD53A0" httpUri="http://treatment.plazi.org/id/03D07408FFF8FFFCFF30FB8238FD53A0" lastPageId="3" lastPageNumber="70" pageId="1" pageNumber="68">
<subSubSection id="C3639695FFF8FFFEFF30FB8238565556" pageId="1" pageNumber="68" type="nomenclature">
<paragraph id="8BC6C51EFFF8FFFEFF30FB823B3C5537" blockId="1.[151,626,1046,1105]" box="[151,626,1046,1072]" pageId="1" pageNumber="68">
<heading id="D08E7272FFF8FFFEFF30FB823B3C5537" bold="true" box="[151,626,1046,1072]" fontSize="11" level="1" pageId="1" pageNumber="68" reason="1">
<emphasis id="B90D190CFFF8FFFEFF30FB823B3C5537" bold="true" box="[151,626,1046,1072]" pageId="1" pageNumber="68">
<taxonomicName id="4C79BE9DFFF8FFFEFF30FB823B675537" box="[151,553,1046,1072]" class="Insecta" family="Mycetophilidae" genus="Scatopsiara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="exigua" status="sp. nov." subGenus="Xenopygina">
<emphasis id="B90D190CFFF8FFFEFF30FB823B675537" bold="true" box="[151,553,1046,1072]" italics="true" pageId="1" pageNumber="68">Scatopsiara (Xenopygina) exigua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23EA477FFF8FFFEFD97FB823B3C5537" box="[560,626,1046,1072]" pageId="1" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC6C51EFFF8FFFEFF30FBAD38565556" blockId="1.[151,626,1046,1105]" box="[151,280,1081,1105]" pageId="1" pageNumber="68">
<figureCitation id="1342D99BFFF8FFFEFF30FBAD39925556" box="[151,220,1081,1105]" captionStart="FIGURE 1" captionStartId="2.[151,250,1310,1333]" captionTargetBox="[160,1419,312,1256]" captionTargetId="figure@2.[151,1436,303,1286]" captionTargetPageId="2" captionText="FIGURE 1. Scatopsciara exigua sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Part of hypopygium, ventral view. C. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283191/files/figure.png" pageId="1" pageNumber="68">Figs 1</figureCitation>
AC
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFF8FFFEFF30FB153BB055BA" pageId="1" pageNumber="68" type="materials_examined">
<paragraph id="8BC6C51EFFF8FFFEFF30FB153BB055BA" blockId="1.[151,1436,1153,2005]" pageId="1" pageNumber="68">
<emphasis id="B90D190CFFF8FFFEFF30FB153824559D" bold="true" box="[151,362,1153,1178]" pageId="1" pageNumber="68">Material studied.</emphasis>
<emphasis id="B90D190CFFF8FFFEFED1FB153B69559E" box="[374,551,1153,1177]" italics="true" pageId="1" pageNumber="68">
<typeStatus id="54C27BBCFFF8FFFEFED1FB153893559E" box="[374,477,1153,1177]" pageId="1" pageNumber="68" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCountry id="F36E858EFFF8FFFEFD94FB163A6B559E" box="[563,805,1153,1177]" name="New Caledonia" pageId="1" pageNumber="68">
<collectionCode id="ED685DDBFFF8FFFEFD94FB163B3D559E" box="[563,627,1154,1177]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="1" pageNumber="68">NEW</collectionCode>
<collectionCode id="ED685DDBFFF8FFFEFDD8FB153A6B559E" box="[639,805,1153,1177]" pageId="1" pageNumber="68">CALEDONIA</collectionCode>
</collectingCountry>
, Rivière Bleue N.P., (parc 6), rainforest,
<date id="FFC7E3DEFFF8FFFEFAB1FB153CD9559E" box="[1302,1431,1153,1177]" pageId="1" pageNumber="68" value="1992-07-16">16.vii.1992</date>
, Bonnet de Larbogne, Chazeau &amp; Guilbert (in
<collectionCode id="ED685DDBFFF8FFFEFD3FFB313BBD55BA" LSID="urn:lsid:biocol.org:col:34988" box="[664,755,1189,1213]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="68">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFF8FFFEFF60FB5D38E257FA" pageId="1" pageNumber="68" type="description">
<paragraph id="8BC6C51EFFF8FFFEFF60FB5D3C5557DE" blockId="1.[151,1436,1153,2005]" pageId="1" pageNumber="68">
<emphasis id="B90D190CFFF8FFFEFF60FB5D381755E5" bold="true" box="[199,345,1225,1250]" pageId="1" pageNumber="68">Description.</emphasis>
Male.
<emphasis id="B90D190CFFF8FFFEFE15FB5E38BC55E5" bold="true" box="[434,498,1226,1250]" pageId="1" pageNumber="68">Head</emphasis>
. Brown, antenna unicolorous, slightly paler brown, maxillary palpus very pale brown. Eye bridge 3 facets wide. Face with 12 scattered longer and shorter setae. Clypeus with 3 setae. Maxillary palpus with 3 palpomeres; palpomere 1 and 3 of equal length, palpomere 2 shortest; palpomere 1 with 1 seta, with a small dorsal patch of sensilla in a depression; surface of antennal flagellomeres rather smooth, body of flagellomere 4 (
<figureCitation id="1342D99BFFF8FFFEFEEEFACD38C55476" box="[329,395,1369,1393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1310,1333]" captionTargetBox="[160,1419,312,1256]" captionTargetId="figure@2.[151,1436,303,1286]" captionTargetPageId="2" captionText="FIGURE 1. Scatopsciara exigua sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Part of hypopygium, ventral view. C. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283191/files/figure.png" pageId="1" pageNumber="68">Fig. 1</figureCitation>
A) 1.7x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere.
<emphasis id="B90D190CFFF8FFFEFEF3FAE938E35492" bold="true" box="[340,429,1405,1429]" pageId="1" pageNumber="68">Thorax</emphasis>
. Brown, setae pale. Anterior pronotum with 2 setae. Episternum 1 with 5 setae. Scutum with pale and short dorsocentrals, with some longer and shorter laterals.
<emphasis id="B90D190CFFF8FFFEFC62FA353D4354BD" bold="true" box="[965,1037,1441,1466]" pageId="1" pageNumber="68">Wing.</emphasis>
Hyalinous. Length 1.0 mm. Width/ length 0.45. Veins indistinct. R1/R 0.35. M1 not well discernible, c/w not measurable in the specimen. r-m/bM 0.35. M, CuA, bM and r-m non-setose.
<emphasis id="B90D190CFFF8FFFEFDD7FA7E3BE65705" bold="true" box="[624,680,1514,1538]" pageId="1" pageNumber="68">Legs</emphasis>
. Yellow. Coxal setae pale. Front tibial organ with pale vestiture forming a short row. Front tibial spur slightly longer than the tibial width. Middle and hind tibiae with a normal and a short spur. Claws without teeth.
<emphasis id="B90D190CFFF8FFFEFD82F9A53BD7574D" bold="true" box="[549,665,1585,1610]" pageId="1" pageNumber="68">Abdomen</emphasis>
. Setae dark.
<emphasis id="B90D190CFFF8FFFEFC93F9A53A98574D" bold="true" box="[820,982,1585,1610]" pageId="1" pageNumber="68">Hypopygium,</emphasis>
<figureCitation id="1342D99BFFF8FFFEFC46F9A53D67574E" box="[993,1065,1585,1609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1310,1333]" captionTargetBox="[160,1419,312,1256]" captionTargetId="figure@2.[151,1436,303,1286]" captionTargetPageId="2" captionText="FIGURE 1. Scatopsciara exigua sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Part of hypopygium, ventral view. C. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283191/files/figure.png" pageId="1" pageNumber="68">Figs 1</figureCitation>
B, C. Brown, concolorous with abdomen.
<emphasis id="B90D190CFFF8FFFEFEAAF9C138CB5769" bold="true" box="[269,389,1621,1646]" pageId="1" pageNumber="68">Gonocoxa</emphasis>
longer than gonostylus, mesial margin with sparse setosity, basoventrally with some longer setae.
<emphasis id="B90D190CFFF8FFFEFF78F9ED382B5795" bold="true" box="[223,357,1657,1682]" pageId="1" pageNumber="68">Gonostylus</emphasis>
narrow, curved, slightly narrowed towards apex, with the mesial side weakly impressed; with sparse apical vestiture, without an apical tooth, with 2 subapical-apical megasetae, 1 megaseta in more lateral position. Tegmen subquadrangular, apically straight, weakly sclerotized, aedeagal teeth not discernible,
</paragraph>
<paragraph id="8BC6C51EFFF8FFFEFF30F97138E257FA" blockId="1.[151,1436,1153,2005]" box="[151,428,1765,1789]" pageId="1" pageNumber="68">aedeagal apodeme short.</paragraph>
</subSubSection>
<subSubSection id="C3639695FFF8FFFDFF60F89D382F51D3" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68" type="discussion">
<paragraph id="8BC6C51EFFF8FFFDFF60F89D382F51D3" blockId="1.[151,1436,1153,2005]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
<emphasis id="B90D190CFFF8FFFEFF60F89D38025625" bold="true" box="[199,332,1801,1826]" pageId="1" pageNumber="68">Discussion.</emphasis>
<taxonomicName id="4C79BE9DFFF8FFFEFEF3F89E3B745626" box="[340,570,1802,1825]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="exigua">
<emphasis id="B90D190CFFF8FFFEFEF3F89E3B745626" box="[340,570,1802,1825]" italics="true" pageId="1" pageNumber="68">Scatopsciara exigua</emphasis>
</taxonomicName>
is unique among New Caledonian species of the genus because it lacks the apical tooth of the gonostylus. The species can be placed in the subgenus
<taxonomicName id="4C79BE9DFFF8FFFEFC40F8BA3DE75641" authority="Frey" authorityName="Frey" box="[999,1193,1838,1862]" class="Insecta" family="Sciaridae" genus="Xenopygina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B90D190CFFF8FFFEFC40F8BA3D235642" box="[999,1133,1838,1861]" italics="true" pageId="1" pageNumber="68">Xenopygina</emphasis>
Frey
</taxonomicName>
according to Menzel and Mohrigs (2000) rather wide concept of the taxon, because of this lack of the apical tooth. It resembles the Palaearctic
<taxonomicName id="4C79BE9DFFF8FFFEFEBCF8E23A71568A" authority="Frey, 1936" authorityName="Frey" authorityYear="1936" box="[283,831,1909,1934]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="dentifera" subGenus="Xenopygina">
<emphasis id="B90D190CFFF8FFFEFEBCF8E23BF3568A" box="[283,701,1909,1933]" italics="true" pageId="1" pageNumber="68">Scatopsciara (Xenopygina) dentifera</emphasis>
Frey,
<quantity id="4C8168FBFFF8FFFEFCA0F8E13A71568A" box="[775,831,1909,1933]" metricMagnitude="1" metricUnit="m" metricValue="4.91744" pageId="1" pageNumber="68" unit="in" value="1936.0">1936</quantity>
</taxonomicName>
in having a rather tumid gonostylus with a recurved megaseta in a very lateral position on the gonostylus (
<quantity id="4C8168FBFFF8FFFEFCB1F80D3A0B56B6" box="[790,837,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="68" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C79BE9DFFF8FFFEFCF7F80E3A9B56B6" box="[848,981,1945,1969]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="B90D190CFFF8FFFEFCF7F80E3A9B56B6" box="[848,981,1945,1969]" italics="true" pageId="1" pageNumber="68">S. dentifera</emphasis>
</taxonomicName>
), but has its gonostylus not broadened towards apex and mesially far less impressed.
<taxonomicName id="4C79BE9DFFF8FFFEFD05F82A3A4756D2" box="[674,777,1982,2005]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="exigua">
<emphasis id="B90D190CFFF8FFFEFD05F82A3A4756D2" box="[674,777,1982,2005]" italics="true" pageId="1" pageNumber="68">S. exigua</emphasis>
</taxonomicName>
also resembles the Palaearctic
<taxonomicName id="4C79BE9DFFF8FFFDFBCDF82A38A351A8" authority="Rudzinski, 1993" authorityName="Rudzinski" authorityYear="1993" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" lastPageId="2" lastPageNumber="69" order="Diptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="subapicalis" subGenus="Xenopygina">
<emphasis id="B90D190CFFF8FFFEFBCDF82A3DB256D2" box="[1130,1276,1982,2005]" italics="true" pageId="1" pageNumber="68">Scatopsciara</emphasis>
(
<emphasis id="B90D190CFFF8FFFEFAAAF82A3CDD56D2" box="[1293,1427,1982,2005]" italics="true" pageId="1" pageNumber="68">Xenopygina</emphasis>
)
<emphasis id="B90D190CFFFBFFFDFF30FF0C385951B7" box="[151,279,152,176]" italics="true" pageId="2" pageNumber="69">subapicalis</emphasis>
(
<bibRefCitation id="EFE8B8EFFFFBFFFDFE80FF0338AB51A8" author="Rudzinski" box="[295,485,151,175]" pageId="2" pageNumber="69" refString="Rudzinski, H. - G. (1993) Mucken und Fliegen aus dem Schluifelder Moos, Ober-Bayern. Zweite Liste (Diptera Nematocera: Sciaridae). Entomofauna, 14, 281 - 304." type="journal article" year="1993">Rudzinski, 1993</bibRefCitation>
)
</taxonomicName>
, which has a more globular gonostylus and different arrangement of its megasetae (
<bibRefCitation id="EFE8B8EFFFFBFFFDFF39FF28381B51D3" author="Rudzinski" box="[158,341,188,212]" pageId="2" pageNumber="69" refString="Rudzinski, H. - G. (1993) Mucken und Fliegen aus dem Schluifelder Moos, Ober-Bayern. Zweite Liste (Diptera Nematocera: Sciaridae). Entomofauna, 14, 281 - 304." type="journal article" year="1993">Rudzinski 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFFBFFFDFF60FF74385654D2" pageId="2" pageNumber="69" type="etymology">
<paragraph id="8BC6C51EFFFBFFFDFF60FF743CD251F0" blockId="2.[151,1436,151,248]" box="[199,1436,223,248]" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF60FF74380051FF" bold="true" box="[199,334,224,248]" pageId="2" pageNumber="69">Etymology.</emphasis>
The name is derived from the Latin word
<emphasis id="B90D190CFFFBFFFDFC8FFF753A3351FF" box="[808,893,225,248]" italics="true" pageId="2" pageNumber="69">exiguus</emphasis>
, small, referring to the small size of the species.
</paragraph>
<caption id="DF069596FFFBFFFDFF30FA8A3BC25454" httpUri="https://zenodo.org/record/283191/files/figure.png" pageId="2" pageNumber="69" targetBox="[160,1419,312,1256]" targetPageId="2">
<paragraph id="8BC6C51EFFFBFFFDFF30FA8A3BC25454" blockId="2.[151,1435,1310,1364]" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF30FA8A385A5433" bold="true" box="[151,276,1310,1333]" pageId="2" pageNumber="69">FIGURE 1.</emphasis>
<taxonomicName id="4C79BE9DFFFBFFFDFEBBFA8B38A25432" box="[284,492,1311,1333]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="exigua" status="sp. nov.">
<emphasis id="B90D190CFFFBFFFDFEBBFA8B38A25432" box="[284,492,1311,1333]" italics="true" pageId="2" pageNumber="69">Scatopsciara exigua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23EA477FFFBFFFDFE53FA8B3B605433" box="[500,558,1311,1332]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B90D190CFFFBFFFDFE53FA8B3B605433" bold="true" box="[500,558,1311,1332]" pageId="2" pageNumber="69">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype).
<emphasis id="B90D190CFFFBFFFDFD09FA8B3B8E5433" bold="true" box="[686,704,1311,1332]" pageId="2" pageNumber="69">A</emphasis>
. Antennal flagellomere 4, lateral view.
<emphasis id="B90D190CFFFBFFFDFBFCFA8B3D225433" bold="true" box="[1115,1132,1311,1332]" pageId="2" pageNumber="69">B</emphasis>
. Part of hypopygium, ventral view.
<emphasis id="B90D190CFFFBFFFDFF74FAA939A55453" bold="true" box="[211,235,1341,1364]" pageId="2" pageNumber="69">C.</emphasis>
Gonostylus, ventral view. Scale 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC6C51EFFFBFFFDFF30FA0D385654D2" blockId="2.[151,565,1433,1494]" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF30FA0D3B6354B5" bold="true" box="[151,557,1433,1458]" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF30FA0D38A154B5" bold="true" box="[151,495,1433,1458]" italics="true" pageId="2" pageNumber="69">
<taxonomicName id="4C79BE9DFFFBFFFDFF30FA0D386554B5" box="[151,299,1433,1458]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">Scatopsciara</taxonomicName>
(s. str.)
<taxonomicName id="4C79BE9DFFFBFFFDFE20FA0D38A154B5" box="[391,495,1433,1458]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="spiculata" status="sp. nov.">spiculata</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23EA477FFFBFFFDFE51FA0E3B6354B5" box="[502,557,1434,1458]" pageId="2" pageNumber="69" rank="species">sp. n</taxonomicNameLabel>
</emphasis>
.
<figureCitation id="1342D99BFFFBFFFDFF30FA29399254D2" box="[151,220,1469,1493]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[174,1426,739,1942]" captionTargetId="figure@3.[151,1436,721,1958]" captionTargetPageId="3" captionText="FIGURE 2. Scatopsciara spiculata sp. n. (A, B paratype, C, D, holotype). A. Antennal flagellomere 4, lateral view. B. Apical part of front tibia, prolateral view. C. Part of hypopygium, ventral view. D. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283192/files/figure.png" pageId="2" pageNumber="69">Figs 2</figureCitation>
AD
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFFBFFFDFF30F9913A29578E" pageId="2" pageNumber="69" type="materials_examined">
<paragraph id="8BC6C51EFFFBFFFDFF30F9913A29578E" blockId="2.[151,1436,1541,2034]" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF30F99138245719" bold="true" box="[151,362,1541,1566]" pageId="2" pageNumber="69">Material studied.</emphasis>
<emphasis id="B90D190CFFFBFFFDFED1F9913B69571A" box="[374,551,1541,1565]" italics="true" pageId="2" pageNumber="69">
<typeStatus id="54C27BBCFFFBFFFDFED1F9913893571A" box="[374,477,1541,1565]" pageId="2" pageNumber="69" type="holotype">Holotype</typeStatus>
male.
</emphasis>
<collectingCountry id="F36E858EFFFBFFFDFD94F9923A6B571A" box="[563,805,1541,1565]" name="New Caledonia" pageId="2" pageNumber="69">
<collectionCode id="ED685DDBFFFBFFFDFD94F9923B3D571A" box="[563,627,1542,1565]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="2" pageNumber="69">NEW</collectionCode>
<collectionCode id="ED685DDBFFFBFFFDFDD8F9913A6B571A" box="[639,805,1541,1565]" pageId="2" pageNumber="69">CALEDONIA</collectionCode>
</collectingCountry>
, Rivière Bleue N.P., (parc 7), rainforest,
<date id="FFC7E3DEFFFBFFFDFAB1F9913CD9571A" box="[1302,1431,1541,1565]" pageId="2" pageNumber="69" value="1992-07-21">21.vii.1992</date>
, Bonnet de Larbogne, Chazeau &amp; Guilbert (in
<collectionCode id="ED685DDBFFFBFFFDFD01F9BD3A4F5746" LSID="urn:lsid:biocol.org:col:34988" box="[678,769,1577,1601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="69">MNHN</collectionCode>
).
<emphasis id="B90D190CFFFBFFFDFCB2F9BE3AC15746" box="[789,911,1578,1601]" italics="true" pageId="2" pageNumber="69">
<typeStatus id="54C27BBCFFFBFFFDFCB2F9BE3AC45746" box="[789,906,1578,1601]" pageId="2" pageNumber="69" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D7F0E97FFFBFFFDFC30F9BD3AA95746" box="[919,999,1577,1601]" pageId="2" pageNumber="69" type="male">1 male</specimenCount>
, same data as
<typeStatus id="54C27BBCFFFBFFFDFB37F9BD3DBC5746" box="[1168,1266,1577,1601]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
(in
<collectionCode id="ED685DDBFFFBFFFDFA84F9BD3C375746" box="[1315,1401,1577,1601]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="2" pageNumber="69">SMNH</collectionCode>
);
<specimenCount id="9D7F0E97FFFBFFFDFA29F9BD399E5762" pageId="2" pageNumber="69" type="male">1 male</specimenCount>
, same data as previous but parc 6,
<date id="FFC7E3DEFFFBFFFDFDCBF9D93BA55762" box="[620,747,1613,1637]" pageId="2" pageNumber="69" value="1992-07-16">16.vii.1992</date>
(in
<collectionCode id="ED685DDBFFFBFFFDFCB9F9D93A3B5762" box="[798,885,1613,1637]" pageId="2" pageNumber="69">PWMP</collectionCode>
);
<specimenCount id="9D7F0E97FFFBFFFDFC2CF9D93A925762" box="[907,988,1613,1637]" pageId="2" pageNumber="69" type="male">1 male</specimenCount>
, Mont Nondoué, sclerophyllous forest,
<date id="FFC7E3DEFFFBFFFDFF30F9E53844578E" box="[151,266,1649,1673]" pageId="2" pageNumber="69" value="1992-07-03">3.vii.1992</date>
, Bonnet de Larbogne, Chazeau &amp; Guilbert (in
<collectionCode id="ED685DDBFFFBFFFDFCB0F9E53A14578E" box="[791,858,1649,1673]" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="2" pageNumber="69">MZH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFFBFFFCFF60F9013A815063" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69" type="description">
<paragraph id="8BC6C51EFFFBFFFCFF60F9013A815063" blockId="2.[151,1436,1541,2034]" lastBlockId="3.[151,1436,151,680]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
<emphasis id="B90D190CFFFBFFFDFF60F901381757A9" bold="true" box="[199,345,1685,1710]" pageId="2" pageNumber="69">Description.</emphasis>
Male.
<emphasis id="B90D190CFFFBFFFDFE15F90238BC57A9" bold="true" box="[434,498,1686,1710]" pageId="2" pageNumber="69">Head</emphasis>
. Brown, antenna unicolorous, slightly paler brown, maxillary palpus very pale brown. Eye bridge 3 facets wide. Face with 10 scattered longer and shorter setae. Clypeus with 1 seta. Maxillary palpus with 3 palpomeres; palpomeres short, palpomere 1 longer than palpomere 3, palpomere 2 shortest; palpomere 1 with 13 setae, with a dorsal patch of sensilla in a depression; surface of antennal flagellomeres rough, body of flagellomere 4 (
<figureCitation id="1342D99BFFFBFFFDFE02F8B138A9563A" box="[421,487,1829,1853]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[174,1426,739,1942]" captionTargetId="figure@3.[151,1436,721,1958]" captionTargetPageId="3" captionText="FIGURE 2. Scatopsciara spiculata sp. n. (A, B paratype, C, D, holotype). A. Antennal flagellomere 4, lateral view. B. Apical part of front tibia, prolateral view. C. Part of hypopygium, ventral view. D. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283192/files/figure.png" pageId="2" pageNumber="69">Fig. 2</figureCitation>
A) 1.651.90x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere.
<emphasis id="B90D190CFFFBFFFDFE51F8DE3B005665" bold="true" box="[502,590,1866,1890]" pageId="2" pageNumber="69">Thorax</emphasis>
. Brown, setae pale. Anterior pronotum with 2 setae. Episternum 1 with 37 setae. Scutum with long dorsocentrals, with some longer and shorter laterals, scutellum with 2 longer and some short setae.
<emphasis id="B90D190CFFFBFFFDFE80F805382056AD" bold="true" box="[295,366,1937,1962]" pageId="2" pageNumber="69">Wing.</emphasis>
Hyalinous. Length
<quantity id="4C8168FBFFFBFFFDFDF9F8053BA356AE" box="[606,749,1937,1961]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="69" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
. Width/length 0.45. Veins indistinct. R1/R 0.400.50. c/
<geoCoordinate id="EE4DA3D9FFFBFFFDFA2FF805398456CA" direction="west" orientation="longitude" pageId="2" pageNumber="69" precision="555" value="-0.6">w 0.60</geoCoordinate>
0.65. r-m/bM 0.5. M, CuA, r-m and bM non-setose.
<emphasis id="B90D190CFFFBFFFDFCBAF8223A1B56C9" bold="true" box="[797,853,1974,1998]" pageId="2" pageNumber="69">Legs</emphasis>
. Yellow. Coxal setae pale. Apical part of front tibia,
<figureCitation id="1342D99BFFFBFFFDFF30F84D399756F6" box="[151,217,2009,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[174,1426,739,1942]" captionTargetId="figure@3.[151,1436,721,1958]" captionTargetPageId="3" captionText="FIGURE 2. Scatopsciara spiculata sp. n. (A, B paratype, C, D, holotype). A. Antennal flagellomere 4, lateral view. B. Apical part of front tibia, prolateral view. C. Part of hypopygium, ventral view. D. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283192/files/figure.png" pageId="2" pageNumber="69">Fig. 2</figureCitation>
B: tibial organ with pale vestiture forming a short row. Front tibial spur longer than the tibial width. Middle and hind tibiae with a normal and a shortened spur. Claws without teeth.
<emphasis id="B90D190CFFFAFFFCFC66FF033D7B51B7" bold="true" box="[961,1077,151,176]" pageId="3" pageNumber="70">Abdomen</emphasis>
. Setae pale.
<emphasis id="B90D190CFFFAFFFCFB64FF033C2B51B7" bold="true" box="[1219,1381,151,176]" pageId="3" pageNumber="70">Hypopygium,</emphasis>
<figureCitation id="1342D99BFFFAFFFCFACBFF0339EB51D3" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[174,1426,739,1942]" captionTargetId="figure@3.[151,1436,721,1958]" captionTargetPageId="3" captionText="FIGURE 2. Scatopsciara spiculata sp. n. (A, B paratype, C, D, holotype). A. Antennal flagellomere 4, lateral view. B. Apical part of front tibia, prolateral view. C. Part of hypopygium, ventral view. D. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283192/files/figure.png" pageId="3" pageNumber="70">Figs 2</figureCitation>
C, D. Brown, concolorous with abdomen.
<emphasis id="B90D190CFFFAFFFCFD2FFF2F3A4E51D3" bold="true" box="[648,768,187,212]" pageId="3" pageNumber="70">Gonocoxa</emphasis>
longer than gonostylus, mesial margin with sparse setosity.
<emphasis id="B90D190CFFFAFFFCFF30FF4B385351FF" bold="true" box="[151,285,223,248]" pageId="3" pageNumber="70">Gonostylus</emphasis>
narrow, slightly curved, narrowed towards apex, with the mesial side impressed; with dense apical vestiture, with a strong and sharp apical tooth, with 1 subapical megaseta and 3 mesial megasetae (one specimen asymmetrically with 3 and 4 mesial megasetae). Tegmen subquadrangular, weakly sclerotized, with sharp apicolateral corners, with area of aedeagal teeth, aedeagal apodeme short.
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFFAFFFCFF60FEFB38825358" pageId="3" pageNumber="70" type="discussion">
<paragraph id="8BC6C51EFFFAFFFCFF60FEFB38825358" blockId="3.[151,1436,151,680]" pageId="3" pageNumber="70">
<emphasis id="B90D190CFFFAFFFCFF60FEFB3802508F" bold="true" box="[199,332,367,392]" pageId="3" pageNumber="70">Discussion.</emphasis>
<emphasis id="B90D190CFFFAFFFCFEF4FEE43B08508F" box="[339,582,368,392]" italics="true" pageId="3" pageNumber="70">
Scatopsiara
<taxonomicName id="4C79BE9DFFFAFFFCFE78FEE43B08508F" box="[479,582,368,392]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="spiculata">spiculata</taxonomicName>
</emphasis>
resembles
<taxonomicName id="4C79BE9DFFFAFFFCFD64FEE43A2C508F" box="[707,866,368,392]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="tenuistylata">
<emphasis id="B90D190CFFFAFFFCFD64FEE43A2C508F" box="[707,866,368,392]" italics="true" pageId="3" pageNumber="70">S. tenuistylata</emphasis>
</taxonomicName>
in having a narrow gonostylus with a strong apical tooth, and megasetae present at approximately the middle of the gonostylus.
<taxonomicName id="4C79BE9DFFFAFFFCFBE9FE013D9450AB" box="[1102,1242,404,428]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="spiculata">
<emphasis id="B90D190CFFFAFFFCFBE9FE013D9450AB" box="[1102,1242,404,428]" italics="true" pageId="3" pageNumber="70">S. spiculata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C79BE9DFFFAFFFCFA20FE01385450D7" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="tenuistylata">
<emphasis id="B90D190CFFFAFFFCFA20FE01385450D7" italics="true" pageId="3" pageNumber="70">S. tenuistylata</emphasis>
</taxonomicName>
in having 3 or 4 mesial megasetae, whereas
<taxonomicName id="4C79BE9DFFFAFFFCFCBCFE2C3AF450D7" box="[795,954,440,464]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="tenuistylata">
<emphasis id="B90D190CFFFAFFFCFCBCFE2C3AF450D7" box="[795,954,440,464]" italics="true" pageId="3" pageNumber="70">S. tenuistylata</emphasis>
</taxonomicName>
only has one.
<taxonomicName id="4C79BE9DFFFAFFFCFBC4FE2C3DA850D7" box="[1123,1254,440,464]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="spiculata">
<emphasis id="B90D190CFFFAFFFCFBC4FE2C3DA850D7" box="[1123,1254,440,464]" italics="true" pageId="3" pageNumber="70">S. spiculata</emphasis>
</taxonomicName>
also has denser vestiture of the antennal flagellomeres and a shorter, comb-like vestiture at the front tibial organ (
<figureCitation id="1342D99BFFFAFFFCFB77FE483C5850F3" box="[1232,1302,476,500]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[174,1426,739,1942]" captionTargetId="figure@3.[151,1436,721,1958]" captionTargetPageId="3" captionText="FIGURE 2. Scatopsciara spiculata sp. n. (A, B paratype, C, D, holotype). A. Antennal flagellomere 4, lateral view. B. Apical part of front tibia, prolateral view. C. Part of hypopygium, ventral view. D. Gonostylus, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/283192/files/figure.png" pageId="3" pageNumber="70">Figs 2</figureCitation>
B and 3 B). Judging by their hypopygium, both new species can be best placed in the
<taxonomicName id="4C79BE9DFFFAFFFCFC46FD943D98531F" box="[993,1238,512,536]" class="Insecta" family="Mycetophilidae" genus="Scatopsiara" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="atomaria">
<emphasis id="B90D190CFFFAFFFCFC46FD943D98531F" box="[993,1238,512,536]" italics="true" pageId="3" pageNumber="70">Scatopsiara atomaria</emphasis>
</taxonomicName>
group of
<bibRefCitation id="EFE8B8EFFFFAFFFCFAEFFE6B383C533B" author="Menzel" pageId="3" pageNumber="70" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999), 1 - 761." type="journal article" year="2000">Menzel and Mohrig (2000)</bibRefCitation>
, although both species have a shortened, partly reduced, spur in the middle and hind tibia, in addition to the normal spur.
</paragraph>
</subSubSection>
<subSubSection id="C3639695FFFAFFFCFF60FDF838FD53A0" pageId="3" pageNumber="70" type="etymology">
<paragraph id="8BC6C51EFFFAFFFCFF60FDF838FD53A0" blockId="3.[151,1436,151,680]" pageId="3" pageNumber="70">
<emphasis id="B90D190CFFFAFFFCFF60FDF838005383" bold="true" box="[199,334,620,644]" pageId="3" pageNumber="70">Etymology.</emphasis>
The name is derived from the Latin word
<emphasis id="B90D190CFFFAFFFCFCC3FDF83AFB5383" box="[868,949,620,644]" italics="true" pageId="3" pageNumber="70">spicula</emphasis>
, point, referring to the sharp gonostylar megasetae of the species.
</paragraph>
</subSubSection>
</treatment>
</document>