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<document id="B48B7D39D842974C968D67593E04A766" ID="10.11646/zootaxa.4226.4.1" ID-CLB-Dataset="37251" ID-DOI="10.11646/zootaxa.4226.4.1" ID-GBIF-Dataset="8903776b-ace3-4ff6-aa28-635145d78260" ID-ISSN="1175-5326" ID-Zenodo-Dep="265060" ID-ZooBank="64B19A58-BBB5-4858-833F-F7937C3A351F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485846904537" checkinUser="plazi" docAuthor="Min, Bum Sik, Seo, Ji Eun, Grischenko, Andrei V. &amp; Gordon, Dennis P." docDate="2017" docId="03C287EA56254707FF26E4D8FDB1CC53" docLanguage="en" docName="zootaxa.4226.4.1.pdf" docOrigin="Zootaxa 4226 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Celleporella hyalina Linnaeus 1767" docType="treatment" docVersion="8" lastPageNumber="459" masterDocId="FFFBFF925623470FFFB1E025FFCCC82E" masterDocTitle="Intertidal Bryozoa from Korea — new additions to the fauna and a new genus of Bitectiporidae (Cheilostomata) from Baengnyeong Island, Yellow Sea" masterLastPageNumber="470" masterPageNumber="451" pageNumber="457" updateTime="1698436510809" updateUser="ExternalLinkService">
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<mods:title id="47C8C0747D93CD69118D99B0A9E32279">Intertidal Bryozoa from Korea — new additions to the fauna and a new genus of Bitectiporidae (Cheilostomata) from Baengnyeong Island, Yellow Sea</mods:title>
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<mods:namePart id="121F24883F31E4C1AFC724C2ACC8DC93">Min, Bum Sik</mods:namePart>
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<mods:namePart id="1A363E17A252AE58C7686E3BD638F499">Seo, Ji Eun</mods:namePart>
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<mods:namePart id="A7033DA90A8E0AAADA8B9CA3EB381954">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="C8F6CC6CE9BC422E285ED07B9AFFAD2D">Gordon, Dennis P.</mods:namePart>
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<mods:title id="60339B7C3E70AEB3A4781E2423966B26">Zootaxa</mods:title>
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<mods:date id="7C118FEF28698295A0F7B7E944BF6705">2017</mods:date>
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<treatment id="03C287EA56254707FF26E4D8FDB1CC53" ID-DOI="http://doi.org/10.5281/zenodo.5220974" ID-GBIF-Taxon="127627970" ID-Zenodo-Dep="5220974" LSID="urn:lsid:plazi:treatment:03C287EA56254707FF26E4D8FDB1CC53" httpUri="http://treatment.plazi.org/id/03C287EA56254707FF26E4D8FDB1CC53" lastPageId="8" lastPageNumber="459" pageId="6" pageNumber="457">
<subSubSection id="C371657756254709FF26E4D8FEEBCD16" pageId="6" pageNumber="457" type="nomenclature">
<paragraph id="8BD436FC56254709FF26E4D8FDAFCD39" blockId="6.[151,611,1277,1336]" box="[151,611,1277,1303]" pageId="6" pageNumber="457">
<heading id="D09C819056254709FF26E4D8FDAFCD39" bold="true" box="[151,611,1277,1303]" fontSize="11" level="1" pageId="6" pageNumber="457" reason="1">
<taxonomicName id="4C6B4D7F56254709FF26E4D8FDAFCD39" ID-CoL="S3W8" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[151,611,1277,1303]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="457" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE56254709FF26E4D8FDAFCD39" bold="true" box="[151,611,1277,1303]" pageId="6" pageNumber="457">
<emphasis id="B91FEAEE56254709FF26E4D8FE40CD39" bold="true" box="[151,396,1277,1303]" italics="true" pageId="6" pageNumber="457">Celleporella hyalina</emphasis>
(
<bibRefCitation id="EFFA4B0D56254709FE2AE4D8FD97CD39" author="Linnaeus" box="[411,603,1277,1303]" pageId="6" pageNumber="457" refString="Linnaeus, C. (1767) Systemae Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Regnum Animale. Edition XII. Holmiae, Laurentii Salvii, 1327 pp." type="book" year="1767">Linnaeus, 1767</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD436FC56254709FF26E53AFEEBCD16" blockId="6.[151,611,1277,1336]" box="[151,295,1311,1336]" pageId="6" pageNumber="457">
(
<figureCitation id="13502A7956254709FF2EE53AFED3CD16" box="[159,287,1311,1336]" captionStart-0="FIGURES 8 – 13" captionStart-1="FIGURES 14 – 18. 14" captionStartId-0="7.[151,264,1692,1714]" captionStartId-1="9.[151,264,1643,1665]" captionTargetBox-0="[257,1329,193,1669]" captionTargetBox-1="[271,1315,376,1621]" captionTargetId-0="figure@7.[257,1329,193,1670]" captionTargetId-1="figure@9.[271,1315,375,1621]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 8 – 13. Hippothoa imperforata Liu in Liu, Yin &amp; Ma, 2001, Dumujin. 8, Part of a ramifying runner-like colony showing three caudate autozooids; scale bar, 200 µm. 9, Autozooidal orifice; scale bar, 20 µm. 10, Frontolateral view of an autozooid, showing the elevated orificial rim; scale bar, 100 µm. 11, Autozooid viewed in transmitted light, showing the triangular pore-chambers in the lateral walls; scale bar, 100 µm. 12, 13, Celleporella hyalina (Linnaeus, 1767). 12, Young colony showing early astogeny (Namae port); scale bar, 500 µm. 13, Autozooidal (upper right), female (middle) and male (lower right) orifices (Jinchon-ri); scale bar, 100 µm." captionText-1="FIGURES 14 – 18. 14. Celleporella hyalina (Linnaeus, 1767), Jinchon-ri, showing autozooidal and male orifices; scale bar, 50 µm. 15 – 18, Celleporella nodasakae Dick, Grischenko &amp; Mawatari, 2005, Dumujin (except 15, from Ui Island). 15, Frontal view of a female zooid and ooecium (note the characteristic shape of the proximal orificial rim) surrounded by autozooids; scale bar, 200 µm. 16, Orifices of autozooids and female zooids (one with a broken ooecium); scale bar, 200 µm. 17, Autozooidal orifice; scale bar, 25 µm. 18, female orifice, with a possibly damaged left-hand sinus ‘ shoulder’; scale bar, 50 µm." httpUri-0="https://zenodo.org/record/265064/files/figure.png" httpUri-1="https://zenodo.org/record/265065/files/figure.png" pageId="6" pageNumber="457">Figs 12–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371657756254709FF26E54DFB03CDF4" pageId="6" pageNumber="457" type="reference_group">
<paragraph id="8BD436FC56254709FF26E54DFF2BCDB2" blockId="6.[151,1436,1384,1498]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="AB7B11D256254709FF26E54DFF2BCDB2" pageId="6" pageNumber="457">
<taxonomicName id="4C6B4D7F56254709FF26E54DFDC5CD50" ID-CoL="6M8M6" authority="Okada 1929: 17" authorityName="Okada" authorityPageNumber="17" authorityYear="1929" box="[151,521,1384,1406]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="457" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE56254709FF26E54DFE9ACD50" box="[151,342,1384,1406]" italics="true" pageId="6" pageNumber="457">Hippothoa hyalina</emphasis>
:
<treatmentCitation id="0ACA10ED56254709FED5E54DFDC5CD50" author="Okada" box="[356,521,1384,1406]" page="17" pageId="6" pageNumber="457" year="1929">
<bibRefCitation id="EFFA4B0D56254709FED5E54DFE2FCD50" author="Okada" box="[356,483,1384,1406]" pageId="6" pageNumber="457" refString="Okada, Y. (1929) Report of the Biological Survey of Mutsu Bay. 12. Cheilostomatous Bryozoa of Mutsu Bay. Science Reports of the Tohoku Imperial University, 4, 11 - 35, 5 pls." type="journal article" year="1929">Okada 1929</bibRefCitation>
: 17
</treatmentCitation>
</taxonomicName>
, pl. 1, fig. 4, pl. 4, fig. 3;
<treatmentCitation id="0ACA10ED56254709FCA2E54DFC3ECD50" author="Androsova" box="[787,1010,1384,1406]" page="130" pageId="6" pageNumber="457" year="1958">
<bibRefCitation id="EFFA4B0D56254709FCA2E54DFC73CD50" author="Androsova" box="[787,959,1384,1406]" pageId="6" pageNumber="457" refString="Androsova, E. I. (1958) Bryozoa of the order Cheilostomata of the northern part of the Sea of Japan. Issledovaniya dal'nevostochnikh Morei SSSR, 5, 90 - 204. [in Russian]" type="journal article" year="1958">Androsova 1958</bibRefCitation>
: 130
</treatmentCitation>
, fig. 48;
<treatmentCitation id="0ACA10ED56254709FBFFE54DFAC5CD50" author="Kubanin" box="[1102,1289,1384,1406]" page="33" pageId="6" pageNumber="457" year="1976">
<bibRefCitation id="EFFA4B0D56254709FBFFE54DFB2FCD50" author="Kubanin" box="[1102,1251,1384,1406]" pageId="6" pageNumber="457" refString="Kubanin, A. (1976) Bryozoa of Zav'yalov Island. Biologiya Morya, 1, 30 - 35. [in Russian]" type="journal article" year="1976">Kubanin 1976</bibRefCitation>
: 33
</treatmentCitation>
;
<treatmentCitation id="0ACA10ED56254709FAA4E54DFF2FCDB2" author="Gontar" page="14" pageId="6" pageNumber="457" year="1978">
<bibRefCitation id="EFFA4B0D56254709FAA4E54DFA54CD50" author="Gontar" box="[1301,1432,1384,1406]" pageId="6" pageNumber="457" refString="Gontar, V. I. (1978) On the fauna of bryozoans of the coastal waters of Iturup Island (southern Kurile Islands). Biologiya Morya, 1, 10 - 16. [in Russian]" type="book chapter" year="1978">Gontar 1978</bibRefCitation>
: 14
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FC56254709FF26E580FB03CDF4" blockId="6.[151,1436,1384,1498]" pageId="6" pageNumber="457">
<treatmentCitationGroup id="AB7B11D256254709FF26E580FB03CDF4" pageId="6" pageNumber="457">
<taxonomicName id="4C6B4D7F56254709FF26E580FD99CD95" ID-CoL="S3W8" authority="Soule et al. 1995: 183" authorityName="Soule et al." authorityPageNumber="183" authorityYear="1995" box="[151,597,1445,1467]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="457" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE56254709FF26E580FEABCD95" box="[151,359,1445,1467]" italics="true" pageId="6" pageNumber="457">Celleporella hyalina</emphasis>
:
<treatmentCitation id="0ACA10ED56254709FEC5E580FD99CD95" author="Soule" box="[372,597,1445,1467]" page="183" pageId="6" pageNumber="457" year="1995">
<bibRefCitation id="EFFA4B0D56254709FEC5E580FDE8CD95" author="Soule" box="[372,548,1445,1467]" pageId="6" pageNumber="457" refString="Soule, D. F., Soule, J. D. &amp; Chaney, H. W. (1995) Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western Santa Barbara Channel. Irene McCulloch Foundation Monograph Series, 2, 1 - 344." type="journal article" year="1995">
Soule
<emphasis id="B91FEAEE56254709FE05E583FE2CCD95" box="[436,480,1445,1467]" italics="true" pageId="6" pageNumber="457">et al</emphasis>
. 1995
</bibRefCitation>
: 183
</treatmentCitation>
</taxonomicName>
, pl. 66 (
<emphasis id="B91FEAEE56254709FD19E583FD36CD95" box="[680,762,1446,1467]" italics="true" pageId="6" pageNumber="457">cum syn</emphasis>
.);
<treatmentCitation id="0ACA10ED56254709FCA7E580FC2ECD95" author="Liu" box="[790,994,1445,1467]" page="536" pageId="6" pageNumber="457" year="2001">
<bibRefCitation id="EFFA4B0D56254709FCA7E580FC7CCD95" author="Liu" box="[790,944,1445,1467]" pageId="6" pageNumber="457" refString="Liu, X., Yin, X. &amp; Ma, J. (2001) Biology of Marine-Fouling Bryozoans in the Coastal Waters of China. Science Press, Beijing, 860 pp." type="book" year="2001">
Liu
<emphasis id="B91FEAEE56254709FCF1E583FCA0CD95" box="[832,876,1445,1467]" italics="true" pageId="6" pageNumber="457">et al</emphasis>
. 2001
</bibRefCitation>
: 536
</treatmentCitation>
, pl. 36, figs 1–2.
<taxonomicName id="4C6B4D7F56254709FF26E5E1FEABCDF4" ID-CoL="S3W8" box="[151,359,1476,1498]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="457" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE56254709FF26E5E1FEABCDF4" box="[151,359,1476,1498]" italics="true" pageId="6" pageNumber="457">Celleporella hyalina</emphasis>
</taxonomicName>
species complex:
<treatmentCitation id="0ACA10ED56254709FD90E5E1FCCACDF4" author="Dick" box="[545,774,1476,1498]" page="3724" pageId="6" pageNumber="457" year="2005">
<bibRefCitation id="EFFA4B0D56254709FD90E5E1FD04CDF4" author="Dick" box="[545,712,1476,1498]" pageId="6" pageNumber="457" refString="Dick, M. H., Grischenko, A. V. &amp; Mawatari, S. F. (2005) Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska. Journal of Natural History, 39, 3687 - 3784." type="journal article" year="2005">
Dick
<emphasis id="B91FEAEE56254709FDE9E5E0FD49CDF4" box="[600,645,1476,1498]" italics="true" pageId="6" pageNumber="457">et al</emphasis>
. 2005
</bibRefCitation>
: 3724
</treatmentCitation>
, fig. 9;
<treatmentCitation id="0ACA10ED56254709FCE2E5E1FBB0CDF4" author="Grischenko" box="[851,1148,1476,1498]" page="1097" pageId="6" pageNumber="457" year="2007">
<bibRefCitation id="EFFA4B0D56254709FCE2E5E1FBF1CDF4" author="Grischenko" box="[851,1085,1476,1498]" pageId="6" pageNumber="457" refString="Grischenko, A. V., Dick, M. H. &amp; Mawatari, S. F. (2007) Diversity and taxonomy of intertidal Bryozoa (Cheilostomata) at Akkeshi Bay, Hokkaido, Japan. Journal of Natural History, 41, 1047 - 1161." type="journal article" year="2007">
Grischenko
<emphasis id="B91FEAEE56254709FC7FE5E0FC37CDF4" box="[974,1019,1476,1498]" italics="true" pageId="6" pageNumber="457">et al</emphasis>
. 2007
</bibRefCitation>
: 1097
</treatmentCitation>
, fig. 19.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C371657756254709FF26E623FB54CEA5" pageId="6" pageNumber="457" type="materials_examined">
<paragraph id="8BD436FC56254709FF26E623FB54CEA5" blockId="6.[151,1437,1542,2035]" pageId="6" pageNumber="457">
<emphasis id="B91FEAEE56254709FF26E623FE48CE31" bold="true" box="[151,388,1542,1567]" pageId="6" pageNumber="457">Material examined.</emphasis>
NIBRIV0000325933, Jinchon-ri, Baengnyeong Island.
<materialsCitation id="3B033CA156254709FBB9E623FB54CEA5" ID-GBIF-Occurrence="1426520742" country="Iceland" location="East Sea" pageId="6" pageNumber="457" specimenCount="1">
Other material: Woosuk University collection—Baengnyeong
<collectingCountry id="F37C766C56254709FE79E60FFDDDCE6D" box="[456,529,1578,1603]" name="Iceland" pageId="6" pageNumber="457">Island</collectingCountry>
: Hwadong (11 colonies), Junghwadong (3 colonies), Dumujin (24 colonies), Jinchon-ri (51 colonies), Yeonhwa-ri (21 colonies); mostly on rocky substrata, plus shell and crustose coralline algae (
<taxonomicName id="4C6B4D7F56254709FF53E656FE6FCEA5" box="[226,419,1651,1675]" class="Florideophyceae" family="Hapalidiaceae" genus="Clathromorphum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="6" pageNumber="457" phylum="Rhodophyta" rank="genus">
<emphasis id="B91FEAEE56254709FF53E656FE6FCEA5" box="[226,419,1651,1675]" italics="true" pageId="6" pageNumber="457">Clathromorphum</emphasis>
</taxonomicName>
).
<location id="8EB4602756254709FE0AE656FDD2CEA5" LSID="urn:lsid:plazi:treatment:03C287EA56254707FF26E4D8FDB1CC53:8EB4602756254709FE0AE656FDD2CEA5" box="[443,542,1651,1675]" country="Iceland" name="East Sea" pageId="6" pageNumber="457">East Sea</location>
: Namae port (1 colony), on the basal side of a bryozoan.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C371657756254707FF76E6B3FD2AC9A6" lastPageId="8" lastPageNumber="459" pageId="6" pageNumber="457" type="description">
<paragraph id="8BD436FC56254709FF76E6B3FF3DCFDD" blockId="6.[151,1437,1542,2035]" pageId="6" pageNumber="457">
<emphasis id="B91FEAEE56254709FF76E6B3FE95CE81" bold="true" box="[199,345,1686,1711]" pageId="6" pageNumber="457">Description.</emphasis>
Colony encrusting, initially unilaminar, up to
<quantity id="4C939B1956254709FC2DE6B3FC3FCE80" box="[924,1011,1686,1711]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="457" unit="mm" value="15.0">15 mm</quantity>
in diameter, developing additional autozooids and reproductive zooids at a later stage, which increases the thickness of the crust; these frontally budded zooids are orientated irregularly. Autozooids more or less elongate-oval, the gymnocystal frontal shield highly convex, highest suborally where there may be an umbo, typically with weaker transverse ridges across other parts of the shield. Orifice evenly rounded, the near-circular shape interrupted by a stout, slightly upturned pair of condyles that mark the entrance to the sinus; distal rim of orifice somewhat cowl-like, rising up behind the operculum, which is more or less flat; sinus occupying c. 60% of the proximal rim at the level of the condyles regardless of variability in orifice width. Typically 1–3 holes in the interzooidal furrows between zooids, these representing the frontal expressions of pore-chambers from which additional zooids, mainly reproductive, will be budded.
</paragraph>
<caption id="DF14667456244708FF26E6B9FD63CF65" httpUri="https://zenodo.org/record/265064/files/figure.png" pageId="7" pageNumber="458" targetBox="[257,1329,193,1669]" targetPageId="7">
<paragraph id="8BD436FC56244708FF26E6B9FD63CF65" blockId="7.[151,1436,1692,1867]" pageId="7" pageNumber="458">
<emphasis id="B91FEAEE56244708FF26E6B9FE86CE9C" bold="true" box="[151,330,1692,1714]" pageId="7" pageNumber="458">FIGURES 8–13.</emphasis>
<taxonomicName id="4C6B4D7F56244708FEE2E6B9FCB4CE9C" authority="Liu in Liu, Yin &amp; Ma, 2001" authorityName="Liu in Liu, Yin &amp; Ma" authorityYear="2001" box="[339,888,1692,1714]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="7" pageNumber="458" phylum="Bryozoa" rank="species" species="imperforata">
<emphasis id="B91FEAEE56244708FEE2E6B9FD8DCE9C" box="[339,577,1692,1714]" italics="true" pageId="7" pageNumber="458">Hippothoa imperforata</emphasis>
Liu
<emphasis id="B91FEAEE56244708FDC6E6B8FD47CE9C" box="[631,651,1693,1714]" italics="true" pageId="7" pageNumber="458">in</emphasis>
Liu, Yin &amp; Ma, 2001
</taxonomicName>
, Dumujin.
<emphasis id="B91FEAEE56244708FC5EE6B9FBCDCE9C" bold="true" box="[1007,1025,1692,1714]" pageId="7" pageNumber="458">8,</emphasis>
Part of a ramifying runner-like colony showing three caudate autozooids; scale bar, 200 µm.
<emphasis id="B91FEAEE56244708FD7AE69FFD13CEFE" bold="true" box="[715,735,1722,1744]" pageId="7" pageNumber="458">9,</emphasis>
Autozooidal orifice; scale bar, 20 µm.
<emphasis id="B91FEAEE56244708FBCBE69FFB55CEFE" bold="true" box="[1146,1177,1722,1744]" pageId="7" pageNumber="458">10,</emphasis>
Frontolateral view of an autozooid, showing the elevated orificial rim; scale bar, 100 µm.
<emphasis id="B91FEAEE56244708FCE3E6FCFCA3CEC1" bold="true" box="[850,879,1753,1775]" pageId="7" pageNumber="458">11,</emphasis>
Autozooid viewed in transmitted light, showing the triangular pore-chambers in the lateral walls; scale bar, 100 µm.
<emphasis id="B91FEAEE56244708FCF2E6D2FC47CF23" bold="true" box="[835,907,1783,1805]" pageId="7" pageNumber="458">12, 13,</emphasis>
<taxonomicName id="4C6B4D7F56244708FC25E6D2FAE8CF23" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[916,1316,1783,1805]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="7" pageNumber="458" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE56244708FC25E6D2FBA4CF23" box="[916,1128,1783,1805]" italics="true" pageId="7" pageNumber="458">Celleporella hyalina</emphasis>
(Linnaeus, 1767)
</taxonomicName>
.
<emphasis id="B91FEAEE56244708FA81E6D2FA83CF23" bold="true" box="[1328,1359,1783,1805]" pageId="7" pageNumber="458">12,</emphasis>
Young colony showing early astogeny (Namae port); scale bar, 500 µm.
<emphasis id="B91FEAEE56244708FCFAE733FCA6CF02" bold="true" box="[843,874,1814,1836]" pageId="7" pageNumber="458">13,</emphasis>
Autozooidal (upper right), female (middle) and male (lower right) orifices (Jinchon-ri); scale bar, 100 µm.
</paragraph>
</caption>
<paragraph id="8BD436FC56244707FF76E757FE33C8FB" blockId="7.[151,1436,1906,2003]" lastBlockId="8.[151,1436,151,1149]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">Male zooids produced adventitiously, shorter and narrower than autozooids; orifice about half the width of autozooidal orifices but otherwise similar. Female zooids also budded adventitiously, these short, more or less triangular, the frontal shield rising to the suboral region; orifice mostly concealed by ooecium, but much wider than in autozooids, with a broad shallow poster; ectooecium more or less smoothly calcified, with 13–19 funnel-like pseudopores, sometimes ringed.</paragraph>
<paragraph id="8BD436FC562B4707FF76E0FAFBACC933" blockId="8.[151,1436,151,1149]" pageId="8" pageNumber="459">Ancestrula oval, smooth, with a sinusoid orifice; early astogeny asymmetrical, with a daughter zooid arising on one side distolaterally, in turn giving rise to a daughter from a proximolateral position.</paragraph>
<paragraph id="8BD436FC562B4707FF76E102FD2AC9A6" blockId="8.[151,1436,151,1149]" pageId="8" pageNumber="459">
<emphasis id="B91FEAEE562B4707FF76E102FEB1C96E" bold="true" box="[199,381,295,320]" pageId="8" pageNumber="459">Measurements.</emphasis>
ZL, 357–442 (392) µm; ZW, 195–241 (215) µm; OrL, 93–109 (99) µm; OrW 84–106, (93) µm;
<specimenCount id="9D6DFD75562B4707FF7DE169FF13C94A" box="[204,223,332,356]" pageId="8" pageNumber="459" type="male">♂</specimenCount>
OrL, 49–76 (62) µm;
<specimenCount id="9D6DFD75562B4707FE64E169FE24C94A" box="[469,488,332,356]" pageId="8" pageNumber="459" type="male">♂</specimenCount>
OrW, 52–66 (58) µm; OoL, 130–157 (141) µm; OoW, 202–234 (216) µm; AnL, 280 µm; AnW, 220 µm; AnOrL, 87 µm; AnOrW, 77 µm.
</paragraph>
</subSubSection>
<subSubSection id="C3716577562B4707FF76E1B1FE66CAE2" pageId="8" pageNumber="459" type="discussion">
<paragraph id="8BD436FC562B4707FF76E1B1FE66CAE2" blockId="8.[151,1436,151,1149]" pageId="8" pageNumber="459">
<materialsCitation id="3B033CA1562B4707FF76E1B1FE66CAE2" ID-GBIF-Occurrence="1426520750" country="South Korea" location="North Pacific" municipality="North Pacific" pageId="8" pageNumber="459" specimenCount="1">
<emphasis id="B91FEAEE562B4707FF76E1B1FEF7C983" bold="true" box="[199,315,404,429]" pageId="8" pageNumber="459">Remarks.</emphasis>
The synonymy given above is restricted to the
<location id="8EB46027562B4707FCC0E1B1FBC7C983" LSID="urn:lsid:plazi:treatment:03C287EA56254707FF26E4D8FDB1CC53:8EB46027562B4707FCC0E1B1FBC7C983" box="[881,1035,404,429]" country="South Korea" municipality="North Pacific" name="North Pacific" pageId="8" pageNumber="459">North Pacific</location>
and is not exhaustive.
<bibRefCitation id="EFFA4B0D562B4707FAAEE1B1FF2DC9FE" pageId="8" pageNumber="459" refString="Dick, M. H., Grischenko, A. V. &amp; Mawatari, S. F. (2005) Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska. Journal of Natural History, 39, 3687 - 3784." type="journal article">
Dick
<emphasis id="B91FEAEE562B4707FAD1E1B0FA59C982" box="[1376,1429,404,428]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. (2005)
</bibRefCitation>
and
<bibRefCitation id="EFFA4B0D562B4707FEACE192FDFBC9FE" box="[285,567,439,464]" pageId="8" pageNumber="459" refString="Grischenko, A. V., Dick, M. H. &amp; Mawatari, S. F. (2007) Diversity and taxonomy of intertidal Bryozoa (Cheilostomata) at Akkeshi Bay, Hokkaido, Japan. Journal of Natural History, 41, 1047 - 1161." type="journal article">
Grischenko
<emphasis id="B91FEAEE562B4707FE19E19CFE10C9FE" box="[424,476,440,464]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. (2007)
</bibRefCitation>
have suggested that
<taxonomicName id="4C6B4D7F562B4707FC9DE19DFC6FC9FE" box="[812,931,440,464]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE562B4707FC9DE19DFC6FC9FE" box="[812,931,440,464]" italics="true" pageId="8" pageNumber="459">C. hyalina</emphasis>
</taxonomicName>
in the
<collectingMunicipality id="6BB0AC86562B4707FC48E192FB5EC9FE" box="[1017,1170,439,464]" pageId="8" pageNumber="459">North Pacific</collectingMunicipality>
may both be a species complex inasmuch as there is some morphological variation (see
<bibRefCitation id="EFFA4B0D562B4707FC3EE1F9FBD3C9DB" author="Morris" box="[911,1055,476,501]" pageId="8" pageNumber="459" refString="Morris, P. A. (1976) Middle Pliocene temperature implications based on the bryozoa Hippothoa. Journal of Paleontology, 50, 1143 - 1149." type="journal article" year="1976">Morris 1976</bibRefCitation>
,
<bibRefCitation id="EFFA4B0D562B4707FB9FE1F9FBA6C9DA" author="Morris" box="[1070,1130,476,500]" pageId="8" pageNumber="459" refString="Morris, P. A. (1980) The bryozoan family Hippothoidae (Cheilostomata-Ascophora) with emphasis on the genus Hippothoa. Allan Hancock Monographs in Marine Biology, 10, 1 - 115." type="journal article" year="1980">1980</bibRefCitation>
). The ancestrula and early astogeny in the present material match what
<bibRefCitation id="EFFA4B0D562B4707FD3BE1DAFC7ECA36" author="Cancino" box="[650,946,511,536]" pageId="8" pageNumber="459" refString="Cancino, J. M. &amp; Hughes, R. H. (1988) The zooidal polymorphism and astogeny of Celleporella hyalina (Bryozoa: Cheilostomata). Journal of Zoology, 215, 167 - 181." type="journal article" year="1988">Cancino &amp; Hughes (1988)</bibRefCitation>
have described for
<taxonomicName id="4C6B4D7F562B4707FB3FE225FACFCA36" box="[1166,1283,512,536]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE562B4707FB3FE225FACFCA36" box="[1166,1283,512,536]" italics="true" pageId="8" pageNumber="459">C. hyalina</emphasis>
</taxonomicName>
from Britain, which is generally consistently unilateral (although occasional deviations may occur from the standard pattern at the same locality). This concordance in early astogeny between the material from Britain and
<collectingCountry id="F37C766C562B4707FB06E26DFB37CA71" box="[1207,1275,584,607]" name="South Korea" pageId="8" pageNumber="459">Korea</collectingCountry>
could support conspecificity inasmuch as early astogeny of multiserial hippothoids has been regarded as species-specific (e.g.
<bibRefCitation id="EFFA4B0D562B4707FF26E2AAFE68CA86" author="Ryland" box="[151,420,655,680]" pageId="8" pageNumber="459" refString="Ryland, J. S. &amp; Gordon, D. P. (1977) Some British and New Zealand species of Hippothoa (Bryozoa: Cheilostomata). Journal of the Royal Society of New Zealand, 7, 17 - 49." type="journal article" year="1977">Ryland &amp; Gordon 1977</bibRefCitation>
); as noted above, however, molecular evidence points to a complex of cryptic species (see also
<bibRefCitation id="EFFA4B0D562B4707FF7AE291FE52CAE2" author="Gomez" box="[203,414,692,716]" pageId="8" pageNumber="459" refString="Gomez, A., Wright, P. J., Lunt, D. H., Cancino, J. M., Carvalho, G. R. &amp; Hughes, R. N. (2007) Mating trials validate the use of DNA barcoding to reveal cryptic speciation of a marine bryozoan taxon. Proceedings of the Royal Society, B, 274, 199 - 207." type="journal article" year="2007">
Gómez
<emphasis id="B91FEAEE562B4707FE92E290FE9FCAE2" box="[291,339,692,716]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. 2007
</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716577562B4707FF76E2F2FDB1CC53" pageId="8" pageNumber="459" type="distribution">
<paragraph id="8BD436FC562B4707FF76E2F2FDB1CC53" blockId="8.[151,1436,151,1149]" pageId="8" pageNumber="459">
<emphasis id="B91FEAEE562B4707FF76E2F2FEACCADE" bold="true" box="[199,352,727,752]" pageId="8" pageNumber="459">Distribution.</emphasis>
The nominal species was first described from
<collectingRegion id="49AFF81E562B4707FCC7E2FDFC1DCAC1" box="[886,977,728,751]" country="American Samoa" name="Western" pageId="8" pageNumber="459">Western</collectingRegion>
European seas (
<bibRefCitation id="EFFA4B0D562B4707FB3AE2F2FAF9CADE" author="Linnaeus" box="[1163,1333,727,752]" pageId="8" pageNumber="459" refString="Linnaeus, C. (1767) Systemae Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Regnum Animale. Edition XII. Holmiae, Laurentii Salvii, 1327 pp." type="book" year="1767">Linnaeus 1767</bibRefCitation>
) and has subsequently been accorded a near-cosmopolitan distribution. In the northeastern Pacific, material attributed to
<taxonomicName id="4C6B4D7F562B4707FA33E2D9FF27CB16" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE562B4707FA33E2D9FF27CB16" italics="true" pageId="8" pageNumber="459">C. hyalina</emphasis>
</taxonomicName>
has been recorded from
<collectingRegion id="49AFF81E562B4707FDA0E33AFD93CB16" box="[529,607,799,824]" country="United States of America" name="Alaska" pageId="8" pageNumber="459">Alaska</collectingRegion>
to
<collectingRegion id="49AFF81E562B4707FD38E33AFD36CB16" box="[649,762,799,824]" country="United States of America" name="California" pageId="8" pageNumber="459">California</collectingRegion>
(
<bibRefCitation id="EFFA4B0D562B4707FCBDE33AFC14CB16" author="Soule" box="[780,984,799,824]" pageId="8" pageNumber="459" refString="Soule, D. F., Soule, J. D. &amp; Chaney, H. W. (1995) Taxonomic atlas of the benthic fauna of the Santa Maria Basin and western Santa Barbara Channel. Irene McCulloch Foundation Monograph Series, 2, 1 - 344." type="journal article" year="1995">
Soule
<emphasis id="B91FEAEE562B4707FCE9E304FC40CB16" box="[856,908,800,824]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. 1995
</bibRefCitation>
), while in the northwestern Pacific the species has been accorded a distribution from the Arctic (
<bibRefCitation id="EFFA4B0D562B4707FCABE361FC6ECB73" author="Kluge" box="[794,930,836,861]" pageId="8" pageNumber="459" refString="Kluge, G. A. (1962) Bryozoa of the northern seas of the USSR. Opredeliteli po Faune SSSR, Izdavaemye Zoologischeskim Muzeem Akademii Nauk, 76, 1 - 584. [In Russian]" type="journal article" year="1962">Kluge 1962</bibRefCitation>
;
<bibRefCitation id="EFFA4B0D562B4707FC1FE361FBA7CB72" author="Androsova" box="[942,1131,836,861]" pageId="8" pageNumber="459" refString="Androsova, E. I. (1977) Bryozoa in biocenoses of the Arctic Ocean (Franz Josef Land region). Akademiya Nauk SSSR, Zoologicheskii Institut, Issledovaniya Fauny Morei, 14, 194 - 204. [in Russian]" type="journal article" year="1977">Androsova 1977</bibRefCitation>
), Sea of Okhotsk (
<bibRefCitation id="EFFA4B0D562B4707FA8BE361FF1FCBAE" author="Kubanin" pageId="8" pageNumber="459" refString="Kubanin, A. (1976) Bryozoa of Zav'yalov Island. Biologiya Morya, 1, 30 - 35. [in Russian]" type="journal article" year="1976">Kubanin 1976</bibRefCitation>
), Kurile Islands (
<bibRefCitation id="EFFA4B0D562B4707FE13E34DFDF4CBAE" author="Gontar" box="[418,568,872,896]" pageId="8" pageNumber="459" refString="Gontar, V. I. (1978) On the fauna of bryozoans of the coastal waters of Iturup Island (southern Kurile Islands). Biologiya Morya, 1, 10 - 16. [in Russian]" type="book chapter" year="1978">Gontar 1978</bibRefCitation>
),
<collectingRegion id="49AFF81E562B4707FDE3E342FD78CBAE" box="[594,692,871,896]" country="Russia" name="Sakhalin" pageId="8" pageNumber="459">Sakhalin</collectingRegion>
and the
<collectingCountry id="F37C766C562B4707FC92E34DFCAFCB51" box="[803,867,872,895]" name="Japan" pageId="8" pageNumber="459">Japan</collectingCountry>
Sea (
<bibRefCitation id="EFFA4B0D562B4707FC1BE342FBA3CBAE" author="Androsova" box="[938,1135,871,896]" pageId="8" pageNumber="459" refString="Androsova, E. I. (1958) Bryozoa of the order Cheilostomata of the northern part of the Sea of Japan. Issledovaniya dal'nevostochnikh Morei SSSR, 5, 90 - 204. [in Russian]" type="journal article" year="1958">Androsova 1958</bibRefCitation>
). More recently, with the integration of micromorphological (SEM) and molecular characters, additional species have been recognized from different parts of the Northern Hemisphere that were (or would have been) once attributed to
<taxonomicName id="4C6B4D7F562B4707FB58E395FAADCBE6" box="[1257,1377,944,968]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="hyalina">
<emphasis id="B91FEAEE562B4707FB58E395FAADCBE6" box="[1257,1377,944,968]" italics="true" pageId="8" pageNumber="459">C. hyalina</emphasis>
</taxonomicName>
, e.g.
<taxonomicName id="4C6B4D7F562B4707FF26E3F1FDFDCBC2" authority="Alvarez, 1991" authorityName="Alvarez" authorityYear="1991" box="[151,561,980,1005]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="angusta">
<emphasis id="B91FEAEE562B4707FF26E3F1FE48CBC2" box="[151,388,980,1004]" italics="true" pageId="8" pageNumber="459">Celleporella angusta</emphasis>
<bibRefCitation id="EFFA4B0D562B4707FE3FE3F1FDFDCBC2" author="Alvarez" box="[398,561,980,1005]" pageId="8" pageNumber="459" refString="Alvarez, J. A. (1991) La Coleccion de Briozoos del Museo Nacional de Ciencias Naturales. Museo Nacional de Ciencias Naturales, Madrid, 188 pp." type="book" year="1991">Álvarez, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6B4D7F562B4707FD8EE3F1FBC5CBC2" authority="Dick &amp; Ross, 1988" authorityName="Dick &amp; Ross" authorityYear="1988" box="[575,1033,980,1005]" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="reflexa">
<emphasis id="B91FEAEE562B4707FD8EE3F1FCEDCBC2" box="[575,801,980,1004]" italics="true" pageId="8" pageNumber="459">Celleporella reflexa</emphasis>
<bibRefCitation id="EFFA4B0D562B4707FC9BE3F1FBC5CBC2" author="Dick" box="[810,1033,980,1005]" pageId="8" pageNumber="459" refString="Dick, M. H. &amp; Ross, J. P. (1988) Intertidal Bryozoa (Cheilostomata) of the Kodiak vicinity, Alaska. Center for Pacific Northwest Studies, Western Washington University Occasional Paper, 23, 1 - 133." type="journal article" year="1988">Dick &amp; Ross, 1988</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6B4D7F562B4707FBF5E3F1FEFCCC3E" authority="Hughes &amp; Wright, 2014" authorityName="Hughes &amp; Wright" authorityYear="2014" class="Gymnolaemata" family="Hippothoidae" genus="Celleporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="459" phylum="Bryozoa" rank="species" species="osiani">
<emphasis id="B91FEAEE562B4707FBF5E3F1FAD1CBC2" box="[1092,1309,980,1004]" italics="true" pageId="8" pageNumber="459">Celleporella osiani</emphasis>
<bibRefCitation id="EFFA4B0D562B4707FA96E3F1FEFCCC3E" author="Hughes" pageId="8" pageNumber="459" refString="Hughes, R. N. &amp; Wright, P. J. (2014) Self-fertilisation in the Celleporella angusta clade and a description of Celleporella osiani sp. nov. Studi Trenti di Scienze Naturali, 94, 119 - 124." type="journal article" year="2014">Hughes &amp; Wright, 2014</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFFA4B0D562B4707FE8AE3D2FD83CC3E" box="[315,591,1015,1040]" pageId="8" pageNumber="459" refString="Grischenko, A. V., Dick, M. H. &amp; Mawatari, S. F. (2007) Diversity and taxonomy of intertidal Bryozoa (Cheilostomata) at Akkeshi Bay, Hokkaido, Japan. Journal of Natural History, 41, 1047 - 1161." type="journal article">
Grischenko
<emphasis id="B91FEAEE562B4707FE75E3DCFE3ACC3E" box="[452,502,1016,1040]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. (2007)
</bibRefCitation>
noted differences in the numbers of ooecial pseudopores in populations in Akkeshi Bay,
<collectingRegion id="49AFF81E562B4707FE88E439FE67CC1B" box="[313,427,1052,1077]" country="Japan" name="Hokkaido" pageId="8" pageNumber="459">Hokkaido</collectingRegion>
. Further studies are needed on northern Pacific populations to determine the extent and geographical distribution of morphological variation. Published depth ranges are from
<quantity id="4C939B19562B4707FBCBE41AFB1ECC79" box="[1146,1234,1087,1112]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="5.5" metricValueMin="0.0" pageId="8" pageNumber="459" unit="m" value="27.5" valueMax="55.0" valueMin="0.0">0–55 m</quantity>
(e.g.
<bibRefCitation id="EFFA4B0D562B4707FAA7E41AFEE5CC52" author="Hayward" pageId="8" pageNumber="459" refString="Hayward, P. J. &amp; Ryland, J. S. (1999) Cheilostomatous Bryozoa. Part 2. Hippothooidea - Celleporoidea. Synopses of the British Fauna, New Series, 14, 1 - 416." type="journal article" year="1999">Hayward &amp; Ryland 1999</bibRefCitation>
).
<collectingCountry id="F37C766C562B4707FE8CE440FE4ACC52" box="[317,390,1125,1148]" name="South Korea" pageId="8" pageNumber="459">Korea</collectingCountry>
: Baengnyeong
<collectingCountry id="F37C766C562B4707FD80E441FDB5CC53" box="[561,633,1124,1149]" name="Iceland" pageId="8" pageNumber="459">Island</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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