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<document id="D9459382BB509BB8CC799EA584C8FF1E" ID-DOI="10.5281/zenodo.179222" ID-GBIF-Dataset="9584df9a-a4a2-49bb-a256-cb99776f4dc8" ID-ISSN="1175-5326" ID-Zenodo-Dep="179222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1459942420123" checkinUser="plazi" docAuthor="Wild, Alexander L." docDate="2007" docId="03C2878BFFEAFFEFF3D4D12B3338FB0B" docLanguage="en" docName="zt01622p055.pdf" docOrigin="Zootaxa 1622" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Camponotus pellitus Mayr 1862" docType="treatment" docVersion="5" lastPageNumber="46" masterDocId="FFFBFFF3FFC6FFC2F343D6263625FFB2" masterDocTitle="A catalogue of the ants of Paraguay (Hymenoptera: Formicidae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="45" updateTime="1698230627342" updateUser="plazi">
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<mods:title id="23A90D45873B57D32C586AB1251BDF3E">A catalogue of the ants of Paraguay (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="66DE44F2AE3E1433859EBBE3AD982A63">Wild, Alexander L.</mods:namePart>
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<mods:date id="4CEF3CBA4CBD943AD6D378CFB218976B">2007</mods:date>
<mods:detail id="576833C3E7EE50267DFE19FF9AAFFCBD" type="volume">
<mods:number id="C1D079FBB1B047BFF497B4FAFB8706CF">1622</mods:number>
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<mods:identifier id="512FE1926C6E6BCEEA7A2319B9A35788" type="DOI">10.5281/zenodo.179222</mods:identifier>
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<treatment id="03C2878BFFEAFFEFF3D4D12B3338FB0B" ID-DOI="http://doi.org/10.5281/zenodo.6247327" ID-GBIF-Taxon="168420214" ID-Zenodo-Dep="6247327" LSID="urn:lsid:plazi:treatment:03C2878BFFEAFFEFF3D4D12B3338FB0B" httpUri="http://treatment.plazi.org/id/03C2878BFFEAFFEFF3D4D12B3338FB0B" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C3716516FFEAFFEEF3D4D12B34CEF896" box="[151,747,1804,1829]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BD4369DFFEAFFEEF3D4D12B34CEF896" blockId="44.[151,1435,1804,2020]" box="[151,747,1804,1829]" pageId="44" pageNumber="45">
<heading id="D09C81F1FFEAFFEEF3D4D12B34C0F896" box="[151,741,1804,1829]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF3D4D12B37DCF896" authority="Mayr 1862" authorityName="Mayr" authorityYear="1862" box="[151,505,1805,1829]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEAFFEEF3D4D12B375CF897" bold="true" box="[151,377,1805,1829]" italics="true" pageId="44" pageNumber="45">Camponotus pellitus</emphasis>
Mayr 1862
</taxonomicName>
.
<emphasis id="B91FEA8FFFEAFFEEF141D12A34C0F896" bold="true" box="[514,741,1804,1828]" pageId="44" pageNumber="45">REVISED STATUS</emphasis>
</heading>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716516FFEAFFEFF384D10B35F8FEBD" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="8BD4369DFFEAFFEEF384D10B3223F8F6" blockId="44.[151,1435,1804,2020]" box="[199,1030,1837,1860]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF384D10B3474F8F6" authority="Mayr 1862:668" authorityName="Mayr" authorityPageNumber="668" authorityYear="1862" box="[199,593,1837,1860]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEAFFEEF384D10B3786F8F6" box="[199,419,1837,1860]" italics="true" pageId="44" pageNumber="45">Camponotus pellitus</emphasis>
Mayr 1862:668
</taxonomicName>
. [
<typeStatus id="54D0883FFFEAFFEEF121D10B34E7F8F6" box="[610,706,1837,1860]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
w examined, NHMW;
<collectingCountry id="F37C760DFFEAFFEEF0FFD10B3225F8F6" box="[956,1024,1837,1860]" name="Brazil" pageId="44" pageNumber="45">Brazil</collectingCountry>
]
</paragraph>
<paragraph id="8BD4369DFFEAFFEEF384D16B378FF836" blockId="44.[151,1435,1804,2020]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF384D16B3781F8D6" box="[199,420,1869,1892]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEAFFEEF384D16B3781F8D6" box="[199,420,1869,1892]" italics="true" pageId="44" pageNumber="45">Camponotus pellitus</emphasis>
</taxonomicName>
<taxonomicName id="4C6B4D1EFFEAFFEEF2EED16B3526F8D6" ID-CoL="5G5YZ" authority="Forel 1901e: 72" authorityName="Forel" authorityPageNumber="72" authorityYear="1901" box="[429,771,1869,1892]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="variety" species="blandus" variety="scintillans">
var.
<emphasis id="B91FEA8FFFEAFFEEF29ED16B346EF8D6" box="[477,587,1869,1892]" italics="true" pageId="44" pageNumber="45">scintillans</emphasis>
Forel 1901e: 72
</taxonomicName>
.
<emphasis id="B91FEA8FFFEAFFEEF04CD16A35DDF8D6" bold="true" box="[783,1016,1868,1892]" pageId="44" pageNumber="45">NEW SYNONYMY.</emphasis>
[w
<typeStatus id="54D0883FFFEAFFEEF766D16B32A7F8D6" box="[1061,1154,1869,1892]" pageId="44" pageNumber="45" type="syntype">syntypes</typeStatus>
examined, MHNG;
<collectingCountry id="F37C760DFFEAFFEEF620D16B3703F836" name="Paraguay" pageId="44" pageNumber="45">Paraguay</collectingCountry>
(Jerrmann)]
</paragraph>
<paragraph id="8BD4369DFFEAFFEEF384D1AB3222F816" blockId="44.[151,1435,1804,2020]" box="[199,1031,1933,1956]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF384D1AB37D3F816" ID-CoL="QDJY" authority="Emery" authorityName="Emery" box="[199,502,1933,1956]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEAFFEEF384D1AB378CF816" box="[199,425,1933,1956]" italics="true" pageId="44" pageNumber="45">Camponotus blandus</emphasis>
Emery
</taxonomicName>
(nec F. Smith) 1903: 67 (part). Misidentification.
</paragraph>
<paragraph id="8BD4369DFFEAFFEEF384D18B356CF876" blockId="44.[151,1435,1804,2020]" box="[199,841,1965,1988]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF384D18B3561F876" authority="Mayr. Bruch 1914: 229" authorityName="Mayr. Bruch" authorityPageNumber="229" authorityYear="1914" box="[199,836,1965,1988]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pellitus" subGenus="Myrmamblis">
<emphasis id="B91FEA8FFFEAFFEEF384D18B376BF876" box="[199,334,1965,1988]" italics="true" pageId="44" pageNumber="45">Camponotus</emphasis>
(
<emphasis id="B91FEA8FFFEAFFEEF21ED18B37C1F876" box="[349,484,1965,1988]" italics="true" pageId="44" pageNumber="45">Myrmamblis</emphasis>
)
<emphasis id="B91FEA8FFFEAFFEEF2B0D18B3464F876" box="[499,577,1965,1988]" italics="true" pageId="44" pageNumber="45">pellitus</emphasis>
Mayr. Bruch 1914: 229
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4369DFFEAFFEEF384D1EB350CF856" blockId="44.[151,1435,1804,2020]" box="[199,809,1997,2020]" pageId="44" pageNumber="45">
<taxonomicName id="4C6B4D1EFFEAFFEEF384D1EB3500F856" ID-CoL="5G5YW" authority="Mayr. Santschi 1922b: 111" authorityName="Mayr. Santschi" authorityPageNumber="111" authorityYear="1922" box="[199,805,1997,2020]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="blandus" subSpecies="pellitus">
<emphasis id="B91FEA8FFFEAFFEEF384D1EB37DBF856" box="[199,510,1997,2020]" italics="true" pageId="44" pageNumber="45">Camponotus blandus pellitus</emphasis>
Mayr. Santschi 1922b: 111
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF384D6BE3542FF1D" blockId="45.[199,1074,152,271]" box="[199,871,152,175]" pageId="45" pageNumber="46">
<taxonomicName id="4C6B4D1EFFEBFFEFF384D6BE378DFF1D" ID-CoL="QDJY" box="[199,424,152,175]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEBFFEFF384D6BE378DFF1D" box="[199,424,152,175]" italics="true" pageId="45" pageNumber="46">Camponotus blandus</emphasis>
</taxonomicName>
st.
<taxonomicName id="4C6B4D1EFFEBFFEFF28DD6BE3546FF1D" authority="Forel. Santschi 1922b: 111" authorityName="Forel. Santschi" authorityPageNumber="111" authorityYear="1922" box="[462,867,152,175]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="scintillans">
<emphasis id="B91FEA8FFFEBFFEFF28DD6BE3419FF1D" box="[462,572,152,175]" italics="true" pageId="45" pageNumber="46">scintillans</emphasis>
Forel. Santschi 1922b: 111
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF384D69E3217FF7D" blockId="45.[199,1074,152,271]" box="[199,1074,183,207]" pageId="45" pageNumber="46">
<taxonomicName id="4C6B4D1EFFEBFFEFF384D69E3443FF7D" ID-CoL="QDJY" box="[199,614,184,207]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus" subGenus="Myrmaphaenus">
<emphasis id="B91FEA8FFFEBFFEFF384D69E376BFF7D" box="[199,334,184,207]" italics="true" pageId="45" pageNumber="46">Camponotus</emphasis>
(
<emphasis id="B91FEA8FFFEBFFEFF21ED69E3426FF7D" box="[349,515,184,207]" italics="true" pageId="45" pageNumber="46">Myrmaphaenus</emphasis>
)
<emphasis id="B91FEA8FFFEBFFEFF151D69E3443FF7D" box="[530,614,184,207]" italics="true" pageId="45" pageNumber="46">blandus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="4C6B4D1EFFEBFFEFF1FBD69E320EFF7D" authority="Forel. Emery 1925: 154" authorityName="Forel. Emery" authorityPageNumber="154" authorityYear="1925" box="[696,1067,184,207]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="scintillans">
<emphasis id="B91FEA8FFFEBFFEFF1FBD69E3503FF7D" box="[696,806,184,207]" italics="true" pageId="45" pageNumber="46">scintillans</emphasis>
Forel. Emery 1925: 154
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF384D6FE35C9FF5D" blockId="45.[199,1074,152,271]" box="[199,1004,216,239]" pageId="45" pageNumber="46">
<taxonomicName id="4C6B4D1EFFEBFFEFF384D6FE3443FF5D" ID-CoL="QDJY" box="[199,614,216,239]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus" subGenus="Myrmaphaenus">
<emphasis id="B91FEA8FFFEBFFEFF384D6FE376BFF5D" box="[199,334,216,239]" italics="true" pageId="45" pageNumber="46">Camponotus</emphasis>
(
<emphasis id="B91FEA8FFFEBFFEFF21ED6FE3426FF5D" box="[349,515,216,239]" italics="true" pageId="45" pageNumber="46">Myrmaphaenus</emphasis>
)
<emphasis id="B91FEA8FFFEBFFEFF151D6FE3443FF5D" box="[530,614,216,239]" italics="true" pageId="45" pageNumber="46">blandus</emphasis>
</taxonomicName>
<taxonomicName id="4C6B4D1EFFEBFFEFF12ED6FE35CDFF5D" ID-CoL="5G5YW" authority="Mayr. Kempf 1972: 44" authorityName="Mayr. Kempf" authorityPageNumber="44" authorityYear="1972" box="[621,1000,216,239]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="variety" species="blandus" variety="pellitus">
var.
<emphasis id="B91FEA8FFFEBFFEFF1DFD6FE34CEFF5D" box="[668,747,216,239]" italics="true" pageId="45" pageNumber="46">pellitus</emphasis>
Mayr. Kempf 1972: 44
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF384D6DE35F8FEBD" blockId="45.[199,1074,152,271]" box="[199,989,248,271]" pageId="45" pageNumber="46">
<taxonomicName id="4C6B4D1EFFEBFFEFF384D6DE35FCFEBD" ID-CoL="5G5YZ" authority="Forel. Kempf 1972: 45" authorityName="Forel. Kempf" authorityPageNumber="45" authorityYear="1972" box="[199,985,248,271]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="blandus" subGenus="Myrmaphaenus" subSpecies="scintillans">
<emphasis id="B91FEA8FFFEBFFEFF384D6DE376BFEBD" box="[199,334,248,271]" italics="true" pageId="45" pageNumber="46">Camponotus</emphasis>
(
<emphasis id="B91FEA8FFFEBFFEFF21ED6DE3426FEBD" box="[349,515,248,271]" italics="true" pageId="45" pageNumber="46">Myrmaphaenus</emphasis>
)
<emphasis id="B91FEA8FFFEBFFEFF151D6DE34FEFEBD" box="[530,731,248,271]" italics="true" pageId="45" pageNumber="46">blandus scintillans</emphasis>
Forel. Kempf 1972: 45
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716516FFEBFFEFF3D4D7663338FB0B" pageId="45" pageNumber="46" type="discussion">
<paragraph id="8BD4369DFFEBFFEFF3D4D76636D4FE53" blockId="45.[151,1437,320,1210]" pageId="45" pageNumber="46">
The names associated with
<taxonomicName id="4C6B4D1EFFEBFFEFF2F8D76634B8FEE5" box="[443,669,320,343]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEBFFEFF2F8D76634B8FEE5" box="[443,669,320,343]" italics="true" pageId="45" pageNumber="46">Camponotus blandus</emphasis>
</taxonomicName>
present one of the most difficult taxonomic problems in the Paraguayan fauna, and one that is particularly frustrating given the abundance of these ants in open habitats throughout the region. Multiple closely-related species certainly occur in
<collectingCountry id="F37C760DFFEBFFEFF1F7D7A3353DFE2E" box="[692,792,389,412]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
, but aside from Santschi’s
<taxonomicName id="4C6B4D1EFFEBFFEFF77BD7A3329CFE2E" box="[1080,1209,389,412]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="crispulus">
<emphasis id="B91FEA8FFFEBFFEFF77BD7A3329CFE2E" box="[1080,1209,389,412]" italics="true" pageId="45" pageNumber="46">C. crispulus</emphasis>
</taxonomicName>
(see separate discussion under that species), the taxonomic status of these forms is ambiguous and the arrangement in the present study is tenuous.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF385D7CB322BFCAB" blockId="45.[151,1437,320,1210]" pageId="45" pageNumber="46">
In eastern
<collectingCountry id="F37C760DFFEBFFEFF276D7CB37BCFDB6" box="[309,409,493,516]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
, most collections are of dark-colored ants closely matching Mayr’s
<typeStatus id="54D0883FFFEBFFEFF72FD7CB32BFFDB6" box="[1132,1178,493,516]" pageId="45" pageNumber="46">type</typeStatus>
of
<taxonomicName id="4C6B4D1EFFEBFFEFF7FDD7CB3309FDB6" box="[1214,1324,493,516]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF7FDD7CB3309FDB6" box="[1214,1324,493,516]" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
from
<collectingCountry id="F37C760DFFEBFFEFF62FD7CB3697FD95" name="Brazil" pageId="45" pageNumber="46">Brazil</collectingCountry>
and Forel’s
<typeStatus id="54D0883FFFEBFFEFF279D4363757FD95" box="[314,370,528,551]" pageId="45" pageNumber="46">types</typeStatus>
of
<taxonomicName id="4C6B4D1EFFEBFFEFF2DAD4363402FD95" box="[409,551,528,551]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="scintillans">
<emphasis id="B91FEA8FFFEBFFEFF2DAD4363402FD95" box="[409,551,528,551]" italics="true" pageId="45" pageNumber="46">C. scintillans</emphasis>
</taxonomicName>
from
<collectingCountry id="F37C760DFFEBFFEFF128D43634F5FD95" box="[619,720,528,551]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
. Workers from western
<collectingCountry id="F37C760DFFEBFFEFF096D436321DFD95" box="[981,1080,528,551]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
are almost universally bicolored with a red head and mesosoma and dark legs and gaster. The
<taxonomicName id="4C6B4D1EFFEBFFEFF078D414355CFDFB" box="[827,889,562,585]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
workers of these chaco forms have finer suberect hairs on the sides of the head, spaced more densely, than do workers of
<taxonomicName id="4C6B4D1EFFEBFFEFF0D4D4733220FDDE" box="[919,1029,597,620]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF0D4D4733220FDDE" box="[919,1029,597,620]" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
, and the postero-lateral corners of the head of the largest workers are often more produced.
<taxonomicName id="4C6B4D1EFFEBFFEFF19DD45E3504FD3D" box="[734,801,632,655]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="50" phylum="Arthropoda" rank="species" species="minor">Minor</taxonomicName>
workers of the bicolored forms have smaller eyes than do minors of
<taxonomicName id="4C6B4D1EFFEBFFEFF245D4BC3750FD03" box="[262,373,666,689]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF245D4BC3750FD03" box="[262,373,666,689]" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
, and they have a slightly lower, less convex propodeum. Both forms co-occur in eastern
<collectingCountry id="F37C760DFFEBFFEFF67AD4BC33B9FD03" box="[1337,1436,666,689]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
near the
<collectingCountry id="F37C760DFFEBFFEFF3B1D49B3770FD66" box="[242,341,701,724]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
River with little evidence of interbreeding. Forel’s
<typeStatus id="54D0883FFFEBFFEFF03AD49B3594FD66" box="[889,945,701,724]" pageId="45" pageNumber="46">types</typeStatus>
of
<taxonomicName id="4C6B4D1EFFEBFFEFF097D49B324CFD66" box="[980,1129,701,724]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">
<emphasis id="B91FEA8FFFEBFFEFF097D49B324CFD66" box="[980,1129,701,724]" italics="true" pageId="45" pageNumber="46">C. rosariensis</emphasis>
</taxonomicName>
fall within the range of variation of these bicolored chacoan forms, so I adopt here a two-species scheme with
<taxonomicName id="4C6B4D1EFFEBFFEFF747D4C632BCFD45" box="[1028,1177,736,759]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">
<emphasis id="B91FEA8FFFEBFFEFF747D4C632BCFD45" box="[1028,1177,736,759]" italics="true" pageId="45" pageNumber="46">C. rosariensis</emphasis>
</taxonomicName>
representing the bulk of the chacoan collections and
<taxonomicName id="4C6B4D1EFFEBFFEFF280D5243414FCAB" box="[451,561,770,793]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF280D5243414FCAB" box="[451,561,770,793]" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
representing most of the eastern collections.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF385D5033314FC75" blockId="45.[151,1437,320,1210]" pageId="45" pageNumber="46">
The chacoan forms, while difficult to subdivide further on the basis of external worker morphology, show two distinct nest phenotypes in reproductively mature colonies. I have observed these nest forms within a few meters of each other. One form constructs large, conspicuous crater mounds while the other has cryptic nest entrances in the soil consisting of a single small hole with a slight scattering of excavated earth placed up to half a meter away, a pattern similar to the nests of
<taxonomicName id="4C6B4D1EFFEBFFEFF254D59637A1FC75" box="[279,388,944,967]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF254D59637A1FC75" box="[279,388,944,967]" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
. This sympatric difference in nesting behavior suggests the existence of cryptic species.
</paragraph>
<paragraph id="8BD4369DFFEBFFEFF385D5F43338FB0B" blockId="45.[151,1437,320,1210]" pageId="45" pageNumber="46">
F. Smith’s
<typeStatus id="54D0883FFFEBFFEFF27AD5F437B3FC5B" box="[313,406,978,1001]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6B4D1EFFEBFFEFF2F8D5F4340AFC5B" box="[443,559,978,1001]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEBFFEFF2F8D5F4340AFC5B" box="[443,559,978,1001]" italics="true" pageId="45" pageNumber="46">C. blandus</emphasis>
</taxonomicName>
, a
<taxonomicName id="4C6B4D1EFFEBFFEFF113D5F434B5FC5B" box="[592,656,978,1001]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
worker from Pará (BMNH, examined), is not a close match to any Paraguayan material. This ant has the smaller eyes and the lower propodeum more similar to
<taxonomicName id="4C6B4D1EFFEBFFEFF716D5D332CFFBBE" box="[1109,1258,1013,1036]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">
<emphasis id="B91FEA8FFFEBFFEFF716D5D332CFFBBE" box="[1109,1258,1013,1036]" italics="true" pageId="45" pageNumber="46">C. rosariensis</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C6B4D1EFFEBFFEFF607D5D33693FB9D" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF607D5D33693FB9D" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
, but with denser pubescence on gastric tergites 3 and 4 (= abd. tergites 5 and 6) than in most specimens of
<taxonomicName id="4C6B4D1EFFEBFFEFF6C7D23E3729FBE3" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">
<emphasis id="B91FEA8FFFEBFFEFF6C7D23E3729FBE3" italics="true" pageId="45" pageNumber="46">C. rosariensis</emphasis>
</taxonomicName>
. The
<taxonomicName id="4C6B4D1EFFEBFFEFF20AD21C3799FBE3" box="[329,444,1082,1105]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEBFFEFF20AD21C3799FBE3" box="[329,444,1082,1105]" italics="true" pageId="45" pageNumber="46">C. blandus</emphasis>
</taxonomicName>
<typeStatus id="54D0883FFFEBFFEFF280D21C37D4FBE3" box="[451,497,1082,1105]" pageId="45" pageNumber="46">type</typeStatus>
is also colored differently, with the posterior portion of the head quite dark, as in
<taxonomicName id="4C6B4D1EFFEBFFEFF617D21C36E0FBC6" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="B91FEA8FFFEBFFEFF617D21C36E0FBC6" italics="true" pageId="45" pageNumber="46">C. pellitus</emphasis>
</taxonomicName>
, the mesosoma lighter, as in
<taxonomicName id="4C6B4D1EFFEBFFEFF140D27B34BDFBC6" box="[515,664,1117,1140]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rosariensis">
<emphasis id="B91FEA8FFFEBFFEFF140D27B34BDFBC6" box="[515,664,1117,1140]" italics="true" pageId="45" pageNumber="46">C. rosariensis</emphasis>
</taxonomicName>
, and the first gastric tergite concolorous with the mesosoma and infuscated posteriorly. However, I have not examined any material occurring between the
<typeStatus id="54D0883FFFEBFFEFF764D2A63270FB25" box="[1063,1109,1152,1175]" pageId="45" pageNumber="46">type</typeStatus>
locality in Pará and
<collectingCountry id="F37C760DFFEBFFEFF677D2A633BDFB25" box="[1332,1432,1152,1175]" name="Paraguay" pageId="45" pageNumber="46">Paraguay</collectingCountry>
, so I cannot dismiss the eventuality that the name
<taxonomicName id="4C6B4D1EFFEBFFEFF1E0D2853532FB08" box="[675,791,1187,1210]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="blandus">
<emphasis id="B91FEA8FFFEBFFEFF1E0D2853532FB08" box="[675,791,1187,1210]" italics="true" pageId="45" pageNumber="46">C. blandus</emphasis>
</taxonomicName>
may apply one of these subtropical populations.
</paragraph>
</subSubSection>
</treatment>
</document>
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