treatments-xml/data/03/C2/4E/03C24E22F639562DFF01C594C7A1F32E.xml
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<mods:title id="EB4788560792AC8F422E56FA1F81AD81">A review of the late Cenozoic genus Bohra (Diprotodontia: Macropodidae) and the evolution of tree-kangaroos</mods:title>
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<treatment id="03C24E22F639562DFF01C594C7A1F32E" ID-DOI="http://doi.org/10.5281/zenodo.8017936" ID-GBIF-Taxon="209675092" ID-Zenodo-Dep="8017936" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E" httpUri="http://treatment.plazi.org/id/03C24E22F639562DFF01C594C7A1F32E" lastPageId="37" lastPageNumber="38" pageId="32" pageNumber="33">
<subSubSection id="C371ACBFF6395628FF01C594C639F6C5" box="[151,612,1720,1747]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BD4FF34F6395628FF01C594C639F6C5" blockId="32.[151,612,1720,1747]" box="[151,612,1720,1747]" pageId="32" pageNumber="33">
<heading id="D09C4858F6395628FF01C594C639F6C5" bold="true" box="[151,612,1720,1747]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
<taxonomicName id="4C6B84B7F6395628FF01C594C639F6C5" authority="Dawson, 2004 a" authorityName="Dawson" authorityYear="2004" box="[151,612,1720,1747]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F6395628FF01C594C639F6C5" bold="true" box="[151,612,1720,1747]" pageId="32" pageNumber="33">
<emphasis id="B91F2326F6395628FF01C594C5FFF6C4" bold="true" box="[151,418,1720,1747]" italics="true" pageId="32" pageNumber="33">Bohra wilkinsonorum</emphasis>
<bibRefCitation id="EFFA82C5F6395628FE3FC594C639F6C5" author="Dawson, L." box="[425,612,1720,1746]" pageId="32" pageNumber="33" pagination="267 - 273" refId="ref49751" refString="Dawson, L. (2004 a) A new Pliocene tree kangaroo species (Marsupialia, Macropodinae) from the Chinchilla Local Fauna, southeastern Queensland. Alcheringa, 28 (1), 267 - 273. https: // doi. org / 10.1080 / 03115510408619284" type="journal article" year="2004">Dawson, 2004a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BD4FF34F6395628FF01C42CC10CF70D" blockId="32.[151,1436,1792,1999]" box="[151,1361,1792,1818]" pageId="32" pageNumber="33">
<materialsCitation id="3B03F569F6395628FF01C42CC10CF70D" ID-GBIF-Occurrence="4127780323" box="[151,1361,1792,1818]" collectionCode="QM" pageId="32" pageNumber="33" specimenCode="F43277" specimenCount="1" typeStatus="holotype">
<emphasis id="B91F2326F6395628FF01C42CC556F70D" bold="true" box="[151,267,1792,1818]" pageId="32" pageNumber="33">
<typeStatus id="54D04196F6395628FF01C42CC55BF70D" box="[151,262,1792,1818]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED7A67F1F6395628FE84C42CC562F70D" box="[274,319,1792,1818]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="32" pageNumber="33" type="Museum">QM</collectionCode>
<specimenCode id="DBCD574FF6395628FED0C42CC5C2F70D" box="[326,415,1792,1818]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="32" pageNumber="33" type="Museum">F43277</specimenCode>
, right juvenile maxilla fragment (preserving dP23, M12; P3 removed from crypt).
</materialsCitation>
</paragraph>
<paragraph id="8BD4FF34F6395628FF51C408C10FF775" blockId="32.[151,1436,1792,1999]" pageId="32" pageNumber="33">
<emphasis id="B91F2326F6395628FF51C408C538F729" bold="true" box="[199,357,1828,1854]" pageId="32" pageNumber="33">Type locality.</emphasis>
<materialsCitation id="3B03F569F6395628FEFDC408C10FF775" ID-GBIF-Occurrence="4127780301" collectorName="Doris &amp; Cecil Wilkinson." county="Chinchilla Sand" location="Condamine River" municipality="Chinchilla Rifle Range" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingCounty id="62B587B8F6395628FEFDC408C67DF729" box="[363,544,1828,1854]" pageId="32" pageNumber="33">Chinchilla Sand</collectingCounty>
,
<collectingMunicipality id="6BB0654EF6395628FDBDC408C771F728" box="[555,812,1828,1855]" pageId="32" pageNumber="33">Chinchilla Rifle Range</collectingMunicipality>
(site not specified), north bank of the
<location id="8EB4A9EFF6395628FB47C408C1C5F729" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E:8EB4A9EFF6395628FB47C408C1C5F729" box="[1233,1432,1828,1854]" county="Chinchilla Sand" municipality="Chinchilla Rifle Range" name="Condamine River" pageId="32" pageNumber="33" stateProvince="Queensland">Condamine River</location>
,
<location id="8EB4A9EFF6395628FF01C464C552F775" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E:8EB4A9EFF6395628FF01C464C552F775" box="[151,271,1864,1890]" county="Chinchilla Sand" municipality="Chinchilla Rifle Range" name="Chinchilla" pageId="32" pageNumber="33" stateProvince="Queensland">Chinchilla</location>
,
<location id="8EB4A9EFF6395628FE8CC465C67BF774" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E:8EB4A9EFF6395628FE8CC465C67BF774" box="[282,550,1864,1891]" county="Chinchilla Sand" municipality="Chinchilla Rifle Range" name="western Darling Downs" pageId="32" pageNumber="33" stateProvince="Queensland">western Darling Downs</location>
,
<collectingRegion id="49AF31D6F6395628FDA4C464C6E7F775" box="[562,698,1864,1890]" country="Australia" name="Queensland" pageId="32" pageNumber="33">Queensland</collectingRegion>
.
<location id="8EB4A9EFF6395628FD55C464C6B2F775" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E:8EB4A9EFF6395628FD55C464C6B2F775" box="[707,751,1864,1890]" county="Chinchilla Sand" municipality="Chinchilla Rifle Range" name="The" pageId="32" pageNumber="33" stateProvince="Queensland">The</location>
<typeStatus id="54D04196F6395628FD60C464C705F775" box="[758,856,1864,1890]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
was collected by
<collectorName id="269E9AE2F6395628FBB7C464C002F775" box="[1057,1119,1864,1890]" pageId="32" pageNumber="33">Doris</collectorName>
and
<collectorName id="269E9AE2F6395628FB03C464C10FF775" box="[1173,1362,1864,1890]" pageId="32" pageNumber="33">Cecil Wilkinson.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD4FF34F6395628FF51C440C1CDF7D9" blockId="32.[151,1436,1792,1999]" pageId="32" pageNumber="33">
<emphasis id="B91F2326F6395628FF51C440C5E5F791" bold="true" box="[199,440,1900,1926]" pageId="32" pageNumber="33">Referred specimens.</emphasis>
Probably Chinchilla.
<collectionCode id="ED7A67F1F6395628FD3FC440C68BF791" box="[681,726,1900,1926]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="32" pageNumber="33" type="Museum">QM</collectionCode>
F675, right adult calcaneus. This specimen retains no associated locality or collection information, although it was likely collected in the late 19
<superScript id="7C1E527CF6395628FC64C4BCC7A2F789" attach="left" box="[1010,1023,1936,1950]" fontSize="6" pageId="32" pageNumber="33">th</superScript>
century. Chinchilla Sand is suggested by preservation. See earlier comments on specimens referred to
<emphasis id="B91F2326F6395628FCF0C499C723F7D9" box="[870,894,1973,1998]" italics="true" pageId="32" pageNumber="33">B.</emphasis>
sp. cf.
<taxonomicName id="4C6B84B7F6395628FC59C499C005F7D9" box="[975,1112,1972,1998]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F6395628FC59C499C005F7D9" box="[975,1112,1972,1998]" italics="true" pageId="32" pageNumber="33">B. bandharr</emphasis>
</taxonomicName>
putatively from Chinchilla.
</paragraph>
<caption id="DF14AFBCF6385629FF01C1F2C59EF375" ID-DOI="http://doi.org/10.5281/zenodo.8012710" ID-Zenodo-Dep="8012710" httpUri="https://zenodo.org/record/8012710/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,734,758]" targetBox="[153,1434,183,708]" targetPageId="33" targetType="figure">
<paragraph id="8BD4FF34F6385629FF01C1F2C59EF375" blockId="33.[151,1437,734,866]" pageId="33" pageNumber="34">
<emphasis id="B91F2326F6385629FF01C1F2C57CF2E1" bold="true" box="[151,289,734,758]" pageId="33" pageNumber="34">FIGURE 18.</emphasis>
Cheek teeth of
<taxonomicName id="4C6B84B7F6385629FE5EC1F2C670F2E1" box="[456,557,734,758]" class="Mammalia" family="Macropodidae" genus="Bohra" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B91F2326F6385629FE5EC1F2C65AF2E1" box="[456,519,734,758]" italics="true" pageId="33" pageNumber="34">Bohra</emphasis>
sp.
</taxonomicName>
cf.
<taxonomicName id="4C6B84B7F6385629FDC1C1F3C689F2E1" box="[599,724,734,758]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F6385629FDC1C1F3C689F2E1" box="[599,724,734,758]" italics="true" pageId="33" pageNumber="34">B. bandharr</emphasis>
</taxonomicName>
. NMV P257274, left M3 (AE). A: stereo occlusal view. B: anterior view. C: posterior view. D: lingual view. E: buccal view. NMV P38267, left M2 (EG). F: stereo occlusal view. G: anterior view. H: anterior view. I: NMV P257273, partial left P3, stereo occlusal view. NMV P38325, right p3 (JL). J: buccal view. K: lingual view. L: stereo occlusal view.
</paragraph>
</caption>
<paragraph id="8BD4FF34F6385629FF51C0BCC16CF3D8" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">
<materialsCitation id="3B03F569F6385629FF51C0BCC48CF3D8" ID-GBIF-Occurrence="4127780326" collectingDate="2011-11-24" collectionCode="NMV" location="Childers Cove" pageId="33" pageNumber="34" specimenCode="P221950" specimenCount="1" stateProvince="Victoria">
<location id="8EB4A9EFF6385629FF51C0BCC530F3BC" LSID="urn:lsid:plazi:treatment:03C24E22F639562DFF01C594C7A1F32E:8EB4A9EFF6385629FF51C0BCC530F3BC" box="[199,365,912,939]" name="Childers Cove" pageId="33" pageNumber="34" stateProvince="Victoria">Childers Cove</location>
,
<collectingRegion id="49AF31D6F6385629FEECC0BCC585F3BD" box="[378,472,912,938]" country="Australia" name="Victoria" pageId="33" pageNumber="34">Victoria</collectingRegion>
.
<collectionCode id="ED7A67F1F6385629FE73C0BDC67BF3BC" box="[485,550,913,939]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="34" type="Museum">NMV</collectionCode>
<specimenCode id="DBCD574FF6385629FDB9C0BCC6C8F3BC" box="[559,661,912,939]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="34" type="Museum">P221950</specimenCode>
, left P3. Collected by David Pickering and others on
<date id="FFD5D9F4F6385629FA97C0BDC48CF3D8" pageId="33" pageNumber="34" value="2011-11-24">
<collectingDate id="EF91201CF6385629FA97C0BDC48CF3D8" pageId="33" pageNumber="34" value="2011-11-24">24 November 2011</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B03F569F6385629FF4AC099C170F3D8" ID-GBIF-Occurrence="4127780312" box="[220,1325,948,975]" collectingDate="2004-08-04" collectionCode="NMV" pageId="33" pageNumber="34" specimenCode="P222378" specimenCount="1" stateProvince="Victoria">
<collectionCode id="ED7A67F1F6385629FF4AC099C540F3D8" box="[220,285,949,975]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="34" type="Museum">NMV</collectionCode>
<specimenCode id="DBCD574FF6385629FEB2C099C5D6F3D8" box="[292,395,949,975]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="33" pageNumber="34" type="Museum">P222378</specimenCode>
, right M1. Collected by David Pickering and Wayne Gerdtz on 3 or
<date id="FFD5D9F4F6385629FB10C099C170F3D8" box="[1158,1325,949,975]" pageId="33" pageNumber="34" value="2004-08-04">
<collectingDate id="EF91201CF6385629FB10C099C170F3D8" box="[1158,1325,949,975]" pageId="33" pageNumber="34" value="2004-08-04">4 August 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C371ACBFF6385629FF51C0F5C65BF5B5" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BD4FF34F6385629FF51C0F5C522F401" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">
<emphasis id="B91F2326F6385629FF51C0F5C513F3E4" bold="true" box="[199,334,985,1011]" pageId="33" pageNumber="34">Etymology.</emphasis>
Named in honour of Doris and Cecil Wilkinson for their fossil collecting efforts at the Chinchilla Rifle Range locality.
</paragraph>
<paragraph id="8BD4FF34F6385629FF51C70CC640F4DC" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">
<emphasis id="B91F2326F6385629FF51C70CC5FFF42D" bold="true" box="[199,418,1056,1082]" pageId="33" pageNumber="34">Revised diagnosis.</emphasis>
The largest-known species of
<taxonomicName id="4C6B84B7F6385629FD68C70CC719F42D" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[766,836,1056,1082]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F6385629FD68C70CC719F42D" box="[766,836,1056,1082]" italics="true" pageId="33" pageNumber="34">Bohra</emphasis>
</taxonomicName>
based on molar and calcaneal dimensions. The P3 is similar in size and morphology to that of
<taxonomicName id="4C6B84B7F6385629FDFFC769C6AFF449" box="[617,754,1092,1118]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F6385629FDFFC769C6AFF449" box="[617,754,1092,1118]" italics="true" pageId="33" pageNumber="34">B. bandharr</emphasis>
</taxonomicName>
, but is less markedly narrowed anteriorly. The postpara- and premeta-cristae on M2 are strongly developed and incurved, more so than in any other species of
<taxonomicName id="4C6B84B7F6385629FB4EC744C143F495" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1240,1310,1128,1154]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F6385629FB4EC744C143F495" box="[1240,1310,1128,1154]" italics="true" pageId="33" pageNumber="34">Bohra</emphasis>
</taxonomicName>
. The slight convexity of the buccal side and distinct interloph concavity of the lingual side of the M12 crown resembles that of the much smaller
<taxonomicName id="4C6B84B7F6385629FEEBC79DC648F4DD" authorityName="Prideaux &amp; Warburton" authorityYear="2009" box="[381,533,1200,1226]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nullarbora">
<emphasis id="B91F2326F6385629FEEBC79DC648F4DD" box="[381,533,1200,1226]" italics="true" pageId="33" pageNumber="34">B. nullarbora</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4FF34F6385629FF51C7F8C65BF5B5" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">The species is also distinguished by the following combination of calcaneal features: massive, dorsoventrally compressed tuber calcanei; plantar aspect very broad and strongly flared posteriorly, particularly on the lateral aspect of the epiphysis; medial and lateral talar facets of the calcaneus discontinuous; pronounced and strongly demarked crest from the caudomesial border of the medial talar facet for the posterior talocalcaneal ligament; calcaneocuboid facets transversely broad, with oblique, moderately deep step; ventromedial facet separated from dorsomedial facet by deep fossa.</paragraph>
</subSubSection>
<subSubSection id="C371ACBFF638562DFF51C680C5C1F1BB" lastPageId="37" lastPageNumber="38" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD4FF34F6385629FF51C680C0DDF624" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">
<emphasis id="B91F2326F6385629FF51C680C677F5D1" bold="true" box="[199,554,1452,1479]" pageId="33" pageNumber="34">Description and comparisons.</emphasis>
Maxilla. Essentially all that is preserved of the
<typeStatus id="54D04196F6385629FBA8C680C0FDF5D1" box="[1086,1184,1452,1478]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
maxilla is that portion bearing the erupted dP23 and M12, the unerupted P3, and the masseteric process (
<figureCitation id="1350E3B1F6385629FBE8C6FCC177F5FC" box="[1150,1322,1488,1515]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="33" pageNumber="34">Figure 19AC</figureCitation>
). With its extremity positioned adjacent to the M2 protoloph in this juvenile specimen, the masseteric process is posteroventrally orientated and bears a low, slightly rugose crest along its anterolateral edge (
<figureCitation id="1350E3B1F6385629FC79C534C02EF624" box="[1007,1139,1560,1587]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="33" pageNumber="34">Figure 19C</figureCitation>
).
</paragraph>
<paragraph id="8BD4FF34F6385629FF51C510C799F738" blockId="33.[151,1437,912,2019]" pageId="33" pageNumber="34">
The juvenile maxilla is known for no other species of
<taxonomicName id="4C6B84B7F6385629FCBCC510C72DF641" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[810,880,1596,1622]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F6385629FCBCC510C72DF641" box="[810,880,1596,1622]" italics="true" pageId="33" pageNumber="34">Bohra</emphasis>
</taxonomicName>
, and so no direct morphological comparisons are possible due to the changes that can occur from the ontogenetic stage where M2 has just come into occlusion versus a full adult where M4 is in occlusion. Nevertheless, it is worth noting that in
<taxonomicName id="4C6B84B7F6385629FC69C5A9C0ECF689" authorityName="DeVis" authorityYear="1887" box="[1023,1201,1668,1694]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bennettianus">
<emphasis id="B91F2326F6385629FC69C5A9C0ECF689" box="[1023,1201,1668,1694]" italics="true" pageId="33" pageNumber="34">D. bennettianus</emphasis>
</taxonomicName>
, the molars progress anteriorly relative to the position of the masseteric process between these two ontogenetic stages. When M2 first comes into occlusion, its protoloph is positioned adjacent to the masseteric process, but in adults the abutment of M23 is adjacent to the masseteric process. The same may well have been the case for
<taxonomicName id="4C6B84B7F6385629FBF2C5DDC174F71D" authorityName="Dawson" authorityYear="2004" box="[1124,1321,1776,1802]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F6385629FBF2C5DDC174F71D" box="[1124,1321,1776,1802]" italics="true" pageId="33" pageNumber="34">B. wilkinsonorum</emphasis>
</taxonomicName>
. In shape, the masseteric process is more rounded than in
<taxonomicName id="4C6B84B7F6385629FD3FC439C761F739" authorityName=", Prideaux &amp; Warburton" authorityYear="2008" box="[681,828,1812,1838]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="illuminata">
<emphasis id="B91F2326F6385629FD3FC439C761F739" box="[681,828,1812,1838]" italics="true" pageId="33" pageNumber="34">B. illuminata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B84B7F6385629FCE4C439C7E0F739" authorityName="Dawson" authorityYear="2004" box="[882,957,1812,1838]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bila">
<emphasis id="B91F2326F6385629FCE4C439C7E0F739" box="[882,957,1812,1838]" italics="true" pageId="33" pageNumber="34">B. bila</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4FF34F6385629FF51C415C7C7F744" blockId="33.[151,1437,912,2019]" box="[199,922,1848,1875]" pageId="33" pageNumber="34">
Upper dentition. Previously described in detail (
<bibRefCitation id="EFFA82C5F6385629FD76C415C7D2F744" author="Dawson, L." box="[736,911,1849,1875]" pageId="33" pageNumber="34" pagination="267 - 273" refId="ref49751" refString="Dawson, L. (2004 a) A new Pliocene tree kangaroo species (Marsupialia, Macropodinae) from the Chinchilla Local Fauna, southeastern Queensland. Alcheringa, 28 (1), 267 - 273. https: // doi. org / 10.1080 / 03115510408619284" type="journal article" year="2004">Dawson 2004a</bibRefCitation>
).
</paragraph>
<paragraph id="8BD4FF34F638562AFF51C470C5A0F1C6" blockId="33.[151,1437,912,2019]" lastBlockId="34.[151,1436,151,465]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
The dP23 remain unknown for other species of
<taxonomicName id="4C6B84B7F6385629FD77C470C77AF761" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[737,807,1884,1910]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F6385629FD77C470C77AF761" box="[737,807,1884,1910]" italics="true" pageId="33" pageNumber="34">Bohra</emphasis>
</taxonomicName>
, but comparisons with the much smaller
<taxonomicName id="4C6B84B7F6385629FB7AC471C1C1F761" authorityName="DeVis" authorityYear="1887" box="[1260,1436,1884,1910]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bennettianus">
<emphasis id="B91F2326F6385629FB7AC471C1C1F761" box="[1260,1436,1884,1910]" italics="true" pageId="33" pageNumber="34">D. bennettianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B84B7F6385629FF51C4ADC565F78D" authorityName="Muller" authorityYear="1840" box="[199,312,1921,1946]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="inustus">
<emphasis id="B91F2326F6385629FF51C4ADC565F78D" box="[199,312,1921,1946]" italics="true" pageId="33" pageNumber="34">D. inustus</emphasis>
</taxonomicName>
reveal very similar general crown outlines, although those of
<taxonomicName id="4C6B84B7F6385629FC64C4ADC0FEF78D" authorityName="DeVis" authorityYear="1887" box="[1010,1187,1920,1946]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bennettianus">
<emphasis id="B91F2326F6385629FC64C4ADC0FEF78D" box="[1010,1187,1920,1946]" italics="true" pageId="33" pageNumber="34">D. bennettianus</emphasis>
</taxonomicName>
are the closest match. Their dP2 crowns are relatively flat-sided oval shapes, and they share a low weak buccal cingulum. However,
<taxonomicName id="4C6B84B7F6385629FA12C489C560F7F5" authorityName="Dawson" authorityYear="2004" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F6385629FA12C489C560F7F5" italics="true" pageId="33" pageNumber="34">B. wilkinsonorum</emphasis>
</taxonomicName>
differs by lacking the large anterior cusp of the main crest bordered posteriorly by a deep cleft (
<figureCitation id="1350E3B1F63B562AFF09C3BBC51BF0A6" box="[159,326,151,177]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="34" pageNumber="35">Figure 19AC</figureCitation>
), which is characteristic of both the dP2 and P3 of species of
<taxonomicName id="4C6B84B7F63B562AFB94C3BBC0CEF0A6" box="[1026,1171,151,177]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63B562AFB94C3BBC0CEF0A6" box="[1026,1171,151,177]" italics="true" pageId="34" pageNumber="35">Dendrolagus</emphasis>
</taxonomicName>
. These species are also very similar in dP3 morphology, sharing: a strongly developed postparacrista, which abuts the end of the dP2 main crest; well-developed postpara- and premeta-cristae, which are anteroposteriorly orientated and confluent, rather than incurved as on M12; and a distinct preprotocrista that marks the lingual end of the transversely narrow precingulum. No preprotocrista is evident on the
<typeStatus id="54D04196F63B562AFD5BC20BC772F156" box="[717,815,295,321]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
M1, but there is a slight preprotocrista present on the unworn (unerupted) referred specimen, NMV P222378 (
<figureCitation id="1350E3B1F63B562AFC93C267C7DBF172" box="[773,902,331,357]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="34" pageNumber="35">Figure 19L</figureCitation>
). There is a stylar eminence in the region of cusp C on M1, but not M2 (
<figureCitation id="1350E3B1F63B562AFE04C243C64BF19E" box="[402,534,367,393]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="34" pageNumber="35">Figure 19C</figureCitation>
). Whether a urocrista (of which there is a slight manifestation in
<taxonomicName id="4C6B84B7F63B562AFB7DC243C1C1F19E" authorityName="DeVis" authorityYear="1887" box="[1259,1436,367,393]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="bennettianus">
<emphasis id="B91F2326F63B562AFB7DC243C1C1F19E" box="[1259,1436,367,393]" italics="true" pageId="34" pageNumber="35">D. bennettianus</emphasis>
</taxonomicName>
and a very distinct expression in
<taxonomicName id="4C6B84B7F63B562AFD81C2BFC6D6F1BB" authorityName="Muller" authorityYear="1840" box="[535,651,403,428]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="inustus">
<emphasis id="B91F2326F63B562AFD81C2BFC6D6F1BB" box="[535,651,403,428]" italics="true" pageId="34" pageNumber="35">D. inustus</emphasis>
</taxonomicName>
) was also present in
<taxonomicName id="4C6B84B7F63B562AFC15C2BFC016F1BA" authorityName="Dawson" authorityYear="2004" box="[899,1099,403,429]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63B562AFC15C2BFC016F1BA" box="[899,1099,403,429]" italics="true" pageId="34" pageNumber="35">B. wilkinsonorum</emphasis>
</taxonomicName>
cannot be ascertained due to extensive wear of the metaloph.
</paragraph>
<caption id="DF14AFBCF63B562AFF01C420C775F7A3" ID-DOI="http://doi.org/10.5281/zenodo.8012712" ID-Zenodo-Dep="8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1804,1828]" targetBox="[153,1433,526,1774]" targetPageId="34" targetType="figure">
<paragraph id="8BD4FF34F63B562AFF01C420C775F7A3" blockId="34.[151,1437,1804,1972]" pageId="34" pageNumber="35">
<emphasis id="B91F2326F63B562AFF01C420C57FF733" bold="true" box="[151,290,1804,1828]" pageId="34" pageNumber="35">FIGURE 19.</emphasis>
Maxillary and upper dental specimens of
<taxonomicName id="4C6B84B7F63B562AFD4FC420C7E5F733" authorityName="Dawson" authorityYear="2004" box="[729,952,1804,1828]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63B562AFD4FC420C7E5F733" box="[729,952,1804,1828]" italics="true" pageId="34" pageNumber="35">Bohra wilkinsonorum</emphasis>
</taxonomicName>
. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view.
</paragraph>
</caption>
<caption id="DF14AFBCF63A562BFF01C1B3C734F2AE" ID-Table-UUID="DF14AFBCF63A562BFF01C1B3C734F2AE" box="[151,873,671,697]" httpUri="http://table.plazi.org/id/DF14AFBCF63A562BFF01C1B3C734F2AE" pageId="35" pageNumber="36" startId="35.[151,239,671,697]" targetBox="[159,2025,712,922]" targetIsTable="true" targetPageId="35" targetType="table">
<paragraph id="8BD4FF34F63A562BFF01C1B3C734F2AE" blockId="35.[151,873,671,697]" box="[151,873,671,697]" pageId="35" pageNumber="36">
<emphasis id="B91F2326F63A562BFF01C1B3C557F2AE" bold="true" box="[151,266,671,697]" pageId="35" pageNumber="36">TABLE 3.</emphasis>
Cheek-tooth dimensions (mm) of
<taxonomicName id="4C6B84B7F63A562BFDECC1B3C73EF2AE" authorityName="Dawson" authorityYear="2004" box="[634,867,671,697]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63A562BFDECC1B3C73EF2AE" box="[634,867,671,697]" italics="true" pageId="35" pageNumber="36">Bohra wilkinsonorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD4FF34F63A562BFE5DC1E4C2DAF38D" pageId="35" pageNumber="36">
<table id="F96B0D94F63AA9F7FF09C1E4C3B4F38D" box="[159,2025,712,922]" gridcols="26" gridrows="5" pageId="35" pageNumber="36">
<tr id="355BFD76F63AA9F7FF09C1E4C3B4F2C8" box="[159,2025,712,735]" gridrow="0" pageId="35" pageNumber="36" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-24="1" rowspan-25="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="768A940AF63AA9F7FE5DC1E4C5A8F2C8" box="[459,501,712,735]" gridcol="3" gridrow="0" pageId="35" pageNumber="36">dP2</th>
<th id="768A940AF63AA9F7FC89C1E4C714F2C8" box="[799,841,712,735]" gridcol="8" gridrow="0" pageId="35" pageNumber="36">dP3</th>
<th id="768A940AF63AA9F7FBECC1E4C0F9F2C8" box="[1146,1188,712,735]" gridcol="13" gridrow="0" pageId="35" pageNumber="36">P3</th>
<th id="768A940AF63AA9F7FA45C1E4C1A0F2C8" box="[1491,1533,712,735]" gridcol="18" gridrow="0" pageId="35" pageNumber="36">M1</th>
<th id="768A940AF63AA9F7F8A3C1E4C302F2C8" box="[1845,1887,712,735]" gridcol="23" gridrow="0" pageId="35" pageNumber="36">M2</th>
</tr>
<tr id="355BFD76F63AA9F7FF09C1D9C3B4F31B" box="[159,2025,757,780]" gridrow="1" pageId="35" pageNumber="36" rowspan-0="1">
<td id="768A940AF63AA9F7FEDDC1D9C528F31B" box="[331,373,757,780]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FE10C1D9C5EDF31B" box="[390,432,757,780]" gridcol="2" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FE5DC1D9C5A8F31B" box="[459,501,757,780]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FD86C1D9C66EF31B" box="[528,563,757,780]" gridcol="4" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FDC7C1D9C62DF31B" box="[593,624,757,780]" gridcol="5" gridrow="1" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7FD03C1D9C6E2F31B" box="[661,703,757,780]" gridcol="6" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FD4CC1D9C759F31B" box="[730,772,757,780]" gridcol="7" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FC89C1D9C714F31B" box="[799,841,757,780]" gridcol="8" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FCF2C1D9C7DAF31B" box="[868,903,757,780]" gridcol="9" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FC3FC1D9C795F31B" box="[937,968,757,780]" gridcol="10" gridrow="1" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7FC75C1D9C044F31B" box="[995,1049,757,780]" gridcol="11" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FBA3C1D9C002F31B" box="[1077,1119,757,780]" gridcol="12" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FBECC1D9C0F9F31B" box="[1146,1188,757,780]" gridcol="13" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FB29C1D9C0B4F31B" box="[1215,1257,757,780]" gridcol="14" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FA92C1D9C173F31B" box="[1284,1326,757,780]" gridcol="15" gridrow="1" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7FADFC1D9C12EF31B" box="[1353,1395,757,780]" gridcol="16" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FA18C1D9C1E5F31B" box="[1422,1464,757,780]" gridcol="17" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FA45C1D9C1A0F31B" box="[1491,1533,757,780]" gridcol="18" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7F98EC1D9C21FF31B" box="[1560,1602,757,780]" gridcol="19" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7F9CBC1D9C2DAF31B" box="[1629,1671,757,780]" gridcol="20" gridrow="1" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7F934C1D9C285F31B" box="[1698,1752,757,780]" gridcol="21" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7F966C1D9C347F31B" box="[1776,1818,757,780]" gridcol="22" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7F8A3C1D9C302F31B" box="[1845,1887,757,780]" gridcol="23" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7F8ECC1D9C3F9F31B" box="[1914,1956,757,780]" gridcol="24" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7F829C1D9C3B4F31B" box="[1983,2025,757,780]" gridcol="25" gridrow="1" pageId="35" pageNumber="36">PH</td>
</tr>
<tr id="355BFD76F63AA9F7FF09C00EC3B4F328" box="[159,2025,802,831]" gridrow="2" pageId="35" pageNumber="36">
<th id="768A940AF63AA9F7FF09C00EC56CF328" box="[159,305,802,831]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">QM F43277</th>
<td id="768A940AF63AA9F7FEDDC00EC528F328" box="[331,373,802,831]" gridcol="1" gridrow="2" pageId="35" pageNumber="36">7.38</td>
<td id="768A940AF63AA9F7FE10C00EC5EDF328" box="[390,432,802,831]" gridcol="2" gridrow="2" pageId="35" pageNumber="36">5.19</td>
<td id="768A940AF63AA9F7FE5DC00EC5A8F328" box="[459,501,802,831]" gridcol="3" gridrow="2" pageId="35" pageNumber="36">5.30</td>
<td id="768A940AF63AA9F7FD86C00EC66EF328" box="[528,563,802,831]" gridcol="4" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FDC7C00EC62DF328" box="[593,624,802,831]" gridcol="5" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FD03C00EC6E2F328" box="[661,703,802,831]" gridcol="6" gridrow="2" pageId="35" pageNumber="36">7.27</td>
<td id="768A940AF63AA9F7FD4CC00EC759F328" box="[730,772,802,831]" gridcol="7" gridrow="2" pageId="35" pageNumber="36">5.93</td>
<td id="768A940AF63AA9F7FC89C00EC714F328" box="[799,841,802,831]" gridcol="8" gridrow="2" pageId="35" pageNumber="36">6.72</td>
<td id="768A940AF63AA9F7FCF2C00EC7DAF328" box="[868,903,802,831]" gridcol="9" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC3FC00EC795F328" box="[937,968,802,831]" gridcol="10" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC75C00EC044F328" box="[995,1049,802,831]" gridcol="11" gridrow="2" pageId="35" pageNumber="36">12.28</td>
<td id="768A940AF63AA9F7FBA3C00EC002F328" box="[1077,1119,802,831]" gridcol="12" gridrow="2" pageId="35" pageNumber="36">5.26</td>
<td id="768A940AF63AA9F7FBECC00EC0F9F328" box="[1146,1188,802,831]" gridcol="13" gridrow="2" pageId="35" pageNumber="36">6.43</td>
<td id="768A940AF63AA9F7FB29C00EC0B4F328" box="[1215,1257,802,831]" gridcol="14" gridrow="2" pageId="35" pageNumber="36">5.99</td>
<td id="768A940AF63AA9F7FA92C00EC173F328" box="[1284,1326,802,831]" gridcol="15" gridrow="2" pageId="35" pageNumber="36">6.08</td>
<td id="768A940AF63AA9F7FADFC00EC12EF328" box="[1353,1395,802,831]" gridcol="16" gridrow="2" pageId="35" pageNumber="36">8.93</td>
<td id="768A940AF63AA9F7FA18C00EC1E5F328" box="[1422,1464,802,831]" gridcol="17" gridrow="2" pageId="35" pageNumber="36">7.88</td>
<td id="768A940AF63AA9F7FA45C00EC1A0F328" box="[1491,1533,802,831]" gridcol="18" gridrow="2" pageId="35" pageNumber="36">8.42</td>
<td id="768A940AF63AA9F7F98EC00EC21FF328" box="[1560,1602,802,831]" gridcol="19" gridrow="2" pageId="35" pageNumber="36">4.35</td>
<td id="768A940AF63AA9F7F9CBC00EC2DAF328" box="[1629,1671,802,831]" gridcol="20" gridrow="2" pageId="35" pageNumber="36">4.74</td>
<td id="768A940AF63AA9F7F934C00EC285F328" box="[1698,1752,802,831]" gridcol="21" gridrow="2" pageId="35" pageNumber="36">10.20</td>
<td id="768A940AF63AA9F7F966C00EC347F328" box="[1776,1818,802,831]" gridcol="22" gridrow="2" pageId="35" pageNumber="36">8.72</td>
<td id="768A940AF63AA9F7F8A3C00EC302F328" box="[1845,1887,802,831]" gridcol="23" gridrow="2" pageId="35" pageNumber="36">8.31</td>
<td id="768A940AF63AA9F7F8ECC00EC3F9F328" box="[1914,1956,802,831]" gridcol="24" gridrow="2" pageId="35" pageNumber="36">4.56</td>
<td id="768A940AF63AA9F7F829C00EC3B4F328" box="[1983,2025,802,831]" gridcol="25" gridrow="2" pageId="35" pageNumber="36">5.32</td>
</tr>
<tr id="355BFD76F63AA9F7FF09C07CC3B4F37A" box="[159,2025,848,877]" gridrow="3" pageId="35" pageNumber="36" rowspan-1="1" rowspan-10="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="768A940AF63AA9F7FF09C07CC56CF37A" box="[159,305,848,877]" gridcol="0" gridrow="3" pageId="35" pageNumber="36">NMV P221950</th>
<td id="768A940AF63AA9F7FC75C07CC044F37A" box="[995,1049,848,877]" gridcol="11" gridrow="3" pageId="35" pageNumber="36">12.48</td>
<td id="768A940AF63AA9F7FBA3C07CC002F37A" box="[1077,1119,848,877]" gridcol="12" gridrow="3" pageId="35" pageNumber="36">5.40</td>
<td id="768A940AF63AA9F7FBECC07CC0F9F37A" box="[1146,1188,848,877]" gridcol="13" gridrow="3" pageId="35" pageNumber="36">6.73</td>
<td id="768A940AF63AA9F7FB29C07CC0B4F37A" box="[1215,1257,848,877]" gridcol="14" gridrow="3" pageId="35" pageNumber="36">7.25</td>
<td id="768A940AF63AA9F7FA92C07CC173F37A" box="[1284,1326,848,877]" gridcol="15" gridrow="3" pageId="35" pageNumber="36">5.15</td>
</tr>
<tr id="355BFD76F63AA9F7FF09C051C3B4F38D" box="[159,2025,893,922]" gridrow="4" pageId="35" pageNumber="36" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-2="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="768A940AF63AA9F7FF09C051C56CF38D" box="[159,305,893,922]" gridcol="0" gridrow="4" pageId="35" pageNumber="36">NMV P222378</th>
<td id="768A940AF63AA9F7FADFC051C12EF38D" box="[1353,1395,893,922]" gridcol="16" gridrow="4" pageId="35" pageNumber="36">9.17</td>
<td id="768A940AF63AA9F7FA18C051C1E5F38D" box="[1422,1464,893,922]" gridcol="17" gridrow="4" pageId="35" pageNumber="36">7.85</td>
<td id="768A940AF63AA9F7FA45C051C1A0F38D" box="[1491,1533,893,922]" gridcol="18" gridrow="4" pageId="35" pageNumber="36">7.99</td>
<td id="768A940AF63AA9F7F98EC051C21FF38D" box="[1560,1602,893,922]" gridcol="19" gridrow="4" pageId="35" pageNumber="36">5.16</td>
<td id="768A940AF63AA9F7F9CBC051C2DAF38D" box="[1629,1671,893,922]" gridcol="20" gridrow="4" pageId="35" pageNumber="36">5.14</td>
</tr>
</table>
</paragraph>
<caption id="DF14AFBCF63A562BFF01C0C3C152F41E" ID-Table-UUID="DF14AFBCF63A562BFF01C0C3C152F41E" box="[151,1295,1007,1033]" httpUri="http://table.plazi.org/id/DF14AFBCF63A562BFF01C0C3C152F41E" pageId="35" pageNumber="36" startId="35.[151,239,1007,1033]" targetBox="[151,2037,1048,1525]" targetIsTable="true" targetPageId="35" targetType="table">
<paragraph id="8BD4FF34F63A562BFF01C0C3C152F41E" blockId="35.[151,1295,1007,1033]" box="[151,1295,1007,1033]" pageId="35" pageNumber="36">
<emphasis id="B91F2326F63A562BFF01C0C3C557F41E" bold="true" box="[151,266,1007,1033]" pageId="35" pageNumber="36">TABLE 4.</emphasis>
Cheek-tooth dimensions (mm) of
<taxonomicName id="4C6B84B7F63A562BFDECC0C3C6B0F41E" authorityName="Prideaux &amp; Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[634,749,1007,1033]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bila">
<emphasis id="B91F2326F63A562BFDECC0C3C6B0F41E" box="[634,749,1007,1033]" italics="true" pageId="35" pageNumber="36">Bohra bila</emphasis>
</taxonomicName>
. Data for QM F4740 is from
<bibRefCitation id="EFFA82C5F63A562BFBBFC0C3C156F41E" author="Bartholomai, A." box="[1065,1291,1007,1033]" pageId="35" pageNumber="36" pagination="153 - 154" refId="ref48064" refString="Bartholomai, A. (1976 a) Notes on the fossiliferous Pleistocene fluviatile deposits of the eastern Darling Downs. Bureau of Mineral Resources, Geology and Geophysics, Bulletin, 166, 153 - 154." type="journal article" year="1976">Bartholomai (1976a)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BD4FF34F63A562BFE5CC734C280F5E2" pageId="35" pageNumber="36">
<table id="F96B0D94F63AA9F7FF01C734C3A8F5E2" box="[151,2037,1048,1525]" gridcols="26" gridrows="11" pageId="35" pageNumber="36">
<tr id="355BFD76F63AA9F7FF01C734C3A8F438" box="[151,2037,1048,1071]" gridrow="0" pageId="35" pageNumber="36" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="768A940AF63AA9F7FE50C734C5ADF438" box="[454,496,1048,1071]" gridcol="3" gridrow="0" pageId="35" pageNumber="36">M1</th>
<th id="768A940AF63AA9F7FCB6C734C717F438" box="[800,842,1048,1071]" gridcol="8" gridrow="0" pageId="35" pageNumber="36">M2</th>
<th id="768A940AF63AA9F7FBE5C734C0F0F438" box="[1139,1197,1048,1071]" gridcol="13" gridrow="0" pageId="35" pageNumber="36">M3</th>
<th id="768A940AF63AA9F7FA4EC734C257F438" box="[1496,1546,1048,1071]" gridcol="18" gridrow="0" pageId="35" pageNumber="36">M4</th>
</tr>
<tr id="355BFD76F63AA9F7FF01C76AC3A8F44A" box="[151,2037,1094,1117]" gridrow="1" pageId="35" pageNumber="36" rowspan-0="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1">
<td id="768A940AF63AA9F7FEAEC76AC507F44A" box="[312,346,1094,1117]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FEE2C76AC5F6F44A" box="[372,427,1094,1117]" gridcol="2" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FE50C76AC5ADF44A" box="[454,496,1094,1117]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FD99C76AC626F44A" box="[527,635,1094,1117]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="35" pageNumber="36">AH PH</td>
<td id="768A940AF63AA9F7FD0EC76AC69FF44A" box="[664,706,1094,1117]" gridcol="6" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FD42C76AC757F44A" box="[724,778,1094,1117]" gridcol="7" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FCB6C76AC7BAF44A" box="[800,999,1094,1117]" colspan="3" colspanRight="2" gridcol="8" gridrow="1" pageId="35" pageNumber="36">PW AH PH</td>
<td id="768A940AF63AA9F7FC60C76AC07CF44A" box="[1014,1057,1094,1117]" gridcol="11" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FBA4C76AC03BF44A" box="[1074,1126,1094,1117]" gridcol="12" gridrow="1" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FBE5C76AC0F0F44A" box="[1139,1197,1094,1117]" gridcol="13" gridrow="1" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FB5EC76AC0AFF44A" box="[1224,1266,1094,1117]" gridcol="14" gridrow="1" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FA9BC76AC119F44A" box="[1293,1348,1094,1117]" gridcol="15" gridrow="1" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7FAC2C76AC123F44A" box="[1364,1406,1094,1117]" gridcol="16" gridrow="1" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FA02C76AC257F44A" box="[1428,1546,1094,1117]" colspan="2" colspanRight="1" gridcol="17" gridrow="1" pageId="35" pageNumber="36">AW PW</td>
<td id="768A940AF63AA9F7F9B0C76AC2FFF44A" box="[1574,1698,1094,1117]" colspan="2" colspanRight="1" gridcol="19" gridrow="1" pageId="35" pageNumber="36">AH PH</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C75FC3A8F49D" box="[151,2037,1139,1162]" gridrow="2" pageId="35" pageNumber="36" rowspan-1="1" rowspan-2="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="768A940AF63AA9F7FF01C75FC550F49D" box="[151,269,1139,1162]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">QM F4740</th>
<td id="768A940AF63AA9F7FD0EC75FC69FF49D" box="[664,706,1139,1162]" gridcol="6" gridrow="2" pageId="35" pageNumber="36">8.6</td>
<td id="768A940AF63AA9F7FD42C75FC757F49D" box="[724,778,1139,1162]" gridcol="7" gridrow="2" pageId="35" pageNumber="36">7.4</td>
<td id="768A940AF63AA9F7FCB6C75FC717F49D" box="[800,842,1139,1162]" gridcol="8" gridrow="2" pageId="35" pageNumber="36">7.3</td>
<td id="768A940AF63AA9F7FCEDC75FC7FDF49D" box="[891,928,1139,1162]" gridcol="9" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC2BC75FC7BAF49D" box="[957,999,1139,1162]" gridcol="10" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC60C75FC07CF49D" box="[1014,1057,1139,1162]" gridcol="11" gridrow="2" pageId="35" pageNumber="36">9.6</td>
<td id="768A940AF63AA9F7FBA4C75FC03BF49D" box="[1074,1126,1139,1162]" gridcol="12" gridrow="2" pageId="35" pageNumber="36">7.7</td>
<td id="768A940AF63AA9F7FBE5C75FC0F0F49D" box="[1139,1197,1139,1162]" gridcol="13" gridrow="2" pageId="35" pageNumber="36">7.0</td>
<td id="768A940AF63AA9F7FB5EC75FC0AFF49D" box="[1224,1266,1139,1162]" gridcol="14" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FA9BC75FC119F49D" box="[1293,1348,1139,1162]" gridcol="15" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FAC2C75FC123F49D" box="[1364,1406,1139,1162]" gridcol="16" gridrow="2" pageId="35" pageNumber="36">9.7</td>
<td id="768A940AF63AA9F7FA02C75FC1E2F49D" box="[1428,1471,1139,1162]" gridcol="17" gridrow="2" pageId="35" pageNumber="36">7.2</td>
<td id="768A940AF63AA9F7FA4EC75FC257F49D" box="[1496,1546,1139,1162]" gridcol="18" gridrow="2" pageId="35" pageNumber="36">6.2</td>
<td id="768A940AF63AA9F7F9B0C75FC239F49D" box="[1574,1636,1139,1162]" gridcol="19" gridrow="2" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7F9EAC75FC2FFF49D" box="[1660,1698,1139,1162]" gridcol="20" gridrow="2" pageId="35" pageNumber="36">-</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C78DC3A8F4AF" box="[151,2037,1185,1208]" gridrow="3" pageId="35" pageNumber="36" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1">
<th id="768A940AF63AA9F7FF01C78DC550F4AF" box="[151,269,1185,1208]" gridcol="0" gridrow="3" pageId="35" pageNumber="36">QM F43276</th>
<td id="768A940AF63AA9F7FEAEC78DC5F6F4AF" box="[312,427,1185,1208]" colspan="2" colspanRight="1" gridcol="1" gridrow="3" pageId="35" pageNumber="36">7.36 7.21</td>
<td id="768A940AF63AA9F7FE50C78DC5ADF4AF" box="[454,496,1185,1208]" gridcol="3" gridrow="3" pageId="35" pageNumber="36">7.30</td>
<td id="768A940AF63AA9F7FD99C78DC61BF4AF" box="[527,582,1185,1208]" gridcol="4" gridrow="3" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FDCEC78DC626F4AF" box="[600,635,1185,1208]" gridcol="5" gridrow="3" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FD0EC78DC69FF4AF" box="[664,706,1185,1208]" gridcol="6" gridrow="3" pageId="35" pageNumber="36">8.66</td>
<td id="768A940AF63AA9F7FD42C78DC757F4AF" box="[724,778,1185,1208]" gridcol="7" gridrow="3" pageId="35" pageNumber="36">7.51</td>
<td id="768A940AF63AA9F7FCB6C78DC7FDF4AF" box="[800,928,1185,1208]" colspan="2" colspanRight="1" gridcol="8" gridrow="3" pageId="35" pageNumber="36">7.11 -</td>
<td id="768A940AF63AA9F7FC2BC78DC7BAF4AF" box="[957,999,1185,1208]" gridcol="10" gridrow="3" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC60C78DC07CF4AF" box="[1014,1057,1185,1208]" gridcol="11" gridrow="3" pageId="35" pageNumber="36">9.35</td>
<td id="768A940AF63AA9F7FBA4C78DC0F0F4AF" box="[1074,1197,1185,1208]" colspan="2" colspanRight="1" gridcol="12" gridrow="3" pageId="35" pageNumber="36">7.96 6.75</td>
<td id="768A940AF63AA9F7FB5EC78DC0AFF4AF" box="[1224,1266,1185,1208]" gridcol="14" gridrow="3" pageId="35" pageNumber="36">4.67</td>
<td id="768A940AF63AA9F7FA9BC78DC119F4AF" box="[1293,1348,1185,1208]" gridcol="15" gridrow="3" pageId="35" pageNumber="36">4.67</td>
<td id="768A940AF63AA9F7FAC2C78DC123F4AF" box="[1364,1406,1185,1208]" gridcol="16" gridrow="3" pageId="35" pageNumber="36">9.13</td>
<td id="768A940AF63AA9F7FA02C78DC257F4AF" box="[1428,1546,1185,1208]" colspan="2" colspanRight="1" gridcol="17" gridrow="3" pageId="35" pageNumber="36">7.33 5.49</td>
<td id="768A940AF63AA9F7F9B0C78DC2FFF4AF" box="[1574,1698,1185,1208]" colspan="2" colspanRight="1" gridcol="19" gridrow="3" pageId="35" pageNumber="36">4.07 4.17</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C7E2C3A8F4F2" box="[151,2037,1230,1253]" gridrow="4" pageId="35" pageNumber="36" rowspan-1="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="768A940AF63AA9F7FF01C7E2C550F4F2" box="[151,269,1230,1253]" gridcol="0" gridrow="4" pageId="35" pageNumber="36">QM F58667</th>
<td id="768A940AF63AA9F7FD0EC7E2C69FF4F2" box="[664,706,1230,1253]" gridcol="6" gridrow="4" pageId="35" pageNumber="36">8.42</td>
<td id="768A940AF63AA9F7FD42C7E2C757F4F2" box="[724,778,1230,1253]" gridcol="7" gridrow="4" pageId="35" pageNumber="36">7.53</td>
<td id="768A940AF63AA9F7FCB6C7E2C7BAF4F2" box="[800,999,1230,1253]" colspan="3" colspanRight="2" gridcol="8" gridrow="4" pageId="35" pageNumber="36">7.17 4.62 4.89</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C7D7C3A8F505" box="[151,2037,1275,1298]" gridrow="5" pageId="35" pageNumber="36" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-24="1" rowspan-25="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="768A940AF63AA9F7FEE2C7D7C5ADF505" box="[372,496,1275,1298]" colspan="2" colspanRight="1" gridcol="2" gridrow="5" pageId="35" pageNumber="36">dp3</td>
<td id="768A940AF63AA9F7FD42C7D7C757F505" box="[724,778,1275,1298]" gridcol="7" gridrow="5" pageId="35" pageNumber="36">p3</td>
<td id="768A940AF63AA9F7FC60C7D7C07CF505" box="[1014,1057,1275,1298]" gridcol="11" gridrow="5" pageId="35" pageNumber="36">m1</td>
<td id="768A940AF63AA9F7FA9BC7D7C119F505" box="[1293,1348,1275,1298]" gridcol="15" gridrow="5" pageId="35" pageNumber="36">m2</td>
<td id="768A940AF63AA9F7F9B0C7D7C239F505" box="[1574,1636,1275,1298]" gridcol="19" gridrow="5" pageId="35" pageNumber="36">m3</td>
<td id="768A940AF63AA9F7F8B9C7D7C3CEF505" box="[1839,1939,1275,1298]" gridcol="23" gridrow="5" pageId="35" pageNumber="36">m4</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C605C3A8F557" box="[151,2037,1321,1344]" gridrow="6" pageId="35" pageNumber="36" rowspan-0="1">
<td id="768A940AF63AA9F7FEAEC605C507F557" box="[312,346,1321,1344]" gridcol="1" gridrow="6" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FEE2C605C61BF557" box="[372,582,1321,1344]" colspan="3" colspanRight="2" gridcol="2" gridrow="6" pageId="35" pageNumber="36">AW PW AH PH</td>
<td id="768A940AF63AA9F7FDCEC605C626F557" box="[600,635,1321,1344]" gridcol="5" gridrow="6" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7FD0EC605C69FF557" box="[664,706,1321,1344]" gridcol="6" gridrow="6" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FD42C605C7FDF557" box="[724,928,1321,1344]" colspan="3" colspanRight="2" gridcol="7" gridrow="6" pageId="35" pageNumber="36">PW AH PH L</td>
<td id="768A940AF63AA9F7FC2BC605C7BAF557" box="[957,999,1321,1344]" gridcol="10" gridrow="6" pageId="35" pageNumber="36">AW</td>
<td id="768A940AF63AA9F7FC60C605C07CF557" box="[1014,1057,1321,1344]" gridcol="11" gridrow="6" pageId="35" pageNumber="36">PW</td>
<td id="768A940AF63AA9F7FBA4C605C03BF557" box="[1074,1126,1321,1344]" gridcol="12" gridrow="6" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FBE5C605C119F557" box="[1139,1348,1321,1344]" colspan="3" colspanRight="2" gridcol="13" gridrow="6" pageId="35" pageNumber="36">PH L AW PW</td>
<td id="768A940AF63AA9F7FAC2C605C123F557" box="[1364,1406,1321,1344]" gridcol="16" gridrow="6" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7FA02C605C1E2F557" box="[1428,1471,1321,1344]" gridcol="17" gridrow="6" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7FA4EC605C239F557" box="[1496,1636,1321,1344]" colspan="2" colspanRight="1" gridcol="18" gridrow="6" pageId="35" pageNumber="36">L AW PW</td>
<td id="768A940AF63AA9F7F9EAC605C2FFF557" box="[1660,1698,1321,1344]" gridcol="20" gridrow="6" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7F925C605C280F557" box="[1715,1757,1321,1344]" gridcol="21" gridrow="6" pageId="35" pageNumber="36">PH</td>
<td id="768A940AF63AA9F7F97AC605C37FF557" box="[1772,1826,1321,1344]" gridcol="22" gridrow="6" pageId="35" pageNumber="36">L</td>
<td id="768A940AF63AA9F7F8B9C605C3CEF557" box="[1839,1939,1321,1344]" gridcol="23" gridrow="6" pageId="35" pageNumber="36">AW PW</td>
<td id="768A940AF63AA9F7F833C605C395F557" box="[1957,1992,1321,1344]" gridcol="24" gridrow="6" pageId="35" pageNumber="36">AH</td>
<td id="768A940AF63AA9F7F840C605C3A8F557" box="[2006,2037,1321,1344]" gridcol="25" gridrow="6" pageId="35" pageNumber="36">PH</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C67AC3A8F57A" box="[151,2037,1366,1389]" gridrow="7" pageId="35" pageNumber="36" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="768A940AF63AA9F7FF01C67AC550F57A" box="[151,269,1366,1389]" gridcol="0" gridrow="7" pageId="35" pageNumber="36">QM F4749</th>
<td id="768A940AF63AA9F7FEAEC67AC626F57A" box="[312,635,1366,1389]" colspan="5" colspanRight="4" gridcol="1" gridrow="7" pageId="35" pageNumber="36">6.03 3.47 4.09 3.19 3.49</td>
<td id="768A940AF63AA9F7FCEDC67AC7FDF57A" box="[891,928,1366,1389]" gridcol="9" gridrow="7" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FC2BC67AC7BAF57A" box="[957,999,1366,1389]" gridcol="10" gridrow="7" pageId="35" pageNumber="36">5.02</td>
<td id="768A940AF63AA9F7FC60C67AC07CF57A" box="[1014,1057,1366,1389]" gridcol="11" gridrow="7" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FBA4C67AC03BF57A" box="[1074,1126,1366,1389]" gridcol="12" gridrow="7" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FBE5C67AC0F0F57A" box="[1139,1197,1366,1389]" gridcol="13" gridrow="7" pageId="35" pageNumber="36">-</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C6AFC3A8F58D" box="[151,2037,1411,1434]" gridrow="8" pageId="35" pageNumber="36" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="768A940AF63AA9F7FF01C6AFC550F58D" box="[151,269,1411,1434]" gridcol="0" gridrow="8" pageId="35" pageNumber="36">QM F43280</th>
<td id="768A940AF63AA9F7FDCEC6AFC69FF58D" box="[600,706,1411,1434]" colspan="2" colspanRight="1" gridcol="5" gridrow="8" pageId="35" pageNumber="36">10.61 4.17</td>
<td id="768A940AF63AA9F7FD42C6AFC757F58D" box="[724,778,1411,1434]" gridcol="7" gridrow="8" pageId="35" pageNumber="36">4.56</td>
<td id="768A940AF63AA9F7FCB6C6AFC7BAF58D" box="[800,999,1411,1434]" colspan="3" colspanRight="2" gridcol="8" gridrow="8" pageId="35" pageNumber="36">- - 7.43 -</td>
<td id="768A940AF63AA9F7FC60C6AFC07CF58D" box="[1014,1057,1411,1434]" gridcol="11" gridrow="8" pageId="35" pageNumber="36">5.40</td>
<td id="768A940AF63AA9F7FBA4C6AFC03BF58D" box="[1074,1126,1411,1434]" gridcol="12" gridrow="8" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FBE5C6AFC119F58D" box="[1139,1348,1411,1434]" colspan="3" colspanRight="2" gridcol="13" gridrow="8" pageId="35" pageNumber="36">- 8.30 6.01 6.04</td>
<td id="768A940AF63AA9F7FAC2C6AFC123F58D" box="[1364,1406,1411,1434]" gridcol="16" gridrow="8" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7FA02C6AFC2FFF58D" box="[1428,1698,1411,1434]" colspan="4" colspanRight="3" gridcol="17" gridrow="8" pageId="35" pageNumber="36">- 9.91 7.13 6.91 -</td>
<td id="768A940AF63AA9F7F925C6AFC280F58D" box="[1715,1757,1411,1434]" gridcol="21" gridrow="8" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7F97AC6AFC37FF58D" box="[1772,1826,1411,1434]" gridcol="22" gridrow="8" pageId="35" pageNumber="36">10.51</td>
<td id="768A940AF63AA9F7F8B9C6AFC3CEF58D" box="[1839,1939,1411,1434]" gridcol="23" gridrow="8" pageId="35" pageNumber="36">7.15 6.26</td>
<td id="768A940AF63AA9F7F833C6AFC395F58D" box="[1957,1992,1411,1434]" gridcol="24" gridrow="8" pageId="35" pageNumber="36">-</td>
<td id="768A940AF63AA9F7F840C6AFC3A8F58D" box="[2006,2037,1411,1434]" gridcol="25" gridrow="8" pageId="35" pageNumber="36">-</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C69DC3A8F5DF" box="[151,2037,1457,1480]" gridrow="9" pageId="35" pageNumber="36" rowspan-1="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="768A940AF63AA9F7FF01C69DC550F5DF" box="[151,269,1457,1480]" gridcol="0" gridrow="9" pageId="35" pageNumber="36">QM F58668</th>
<td id="768A940AF63AA9F7FCEDC69DC7BAF5DF" box="[891,999,1457,1480]" colspan="2" colspanRight="1" gridcol="9" gridrow="9" pageId="35" pageNumber="36">8.11 5.14</td>
<td id="768A940AF63AA9F7FC60C69DC07CF5DF" box="[1014,1057,1457,1480]" gridcol="11" gridrow="9" pageId="35" pageNumber="36">5.52</td>
<td id="768A940AF63AA9F7FBA4C69DC0F0F5DF" box="[1074,1197,1457,1480]" colspan="2" colspanRight="1" gridcol="12" gridrow="9" pageId="35" pageNumber="36">5.27 6.05</td>
</tr>
<tr id="355BFD76F63AA9F7FF01C6F2C3A8F5E2" box="[151,2037,1502,1525]" gridrow="10" pageId="35" pageNumber="36" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-22="1" rowspan-23="1" rowspan-24="1" rowspan-25="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="768A940AF63AA9F7FF01C6F2C550F5E2" box="[151,269,1502,1525]" gridcol="0" gridrow="10" pageId="35" pageNumber="36">QM F58669</th>
<td id="768A940AF63AA9F7FA02C6F2C2FFF5E2" box="[1428,1698,1502,1525]" colspan="4" colspanRight="3" gridcol="17" gridrow="10" pageId="35" pageNumber="36">9.60 6.72 6.68 6.02</td>
<td id="768A940AF63AA9F7F925C6F2C280F5E2" box="[1715,1757,1502,1525]" gridcol="21" gridrow="10" pageId="35" pageNumber="36">6.48</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD4FF34F63D562CFF51C3BBC710F10A" blockId="36.[151,1437,151,825]" pageId="36" pageNumber="37">
<taxonomicName id="4C6B84B7F63D562CFF51C3BBC5E0F0A6" authorityName="Dawson" authorityYear="2004" box="[199,445,151,177]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63D562CFF51C3BBC5E0F0A6" box="[199,445,151,177]" italics="true" pageId="36" pageNumber="37">Bohra wilkinsonorum</emphasis>
</taxonomicName>
<materialsCitation id="3B03F569F63D562CFE51C3BBC714F10A" ID-GBIF-Occurrence="4127780334" collectionCode="NMV" pageId="36" pageNumber="37" specimenCode="P221950" specimenCount="1" typeStatus="holotype">
is most similar in P3 size and morphology to
<taxonomicName id="4C6B84B7F63D562CFC4DC3BBC03AF0A6" box="[987,1127,151,177]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F63D562CFC4DC3BBC03AF0A6" box="[987,1127,151,177]" italics="true" pageId="36" pageNumber="37">B. bandharr</emphasis>
</taxonomicName>
(
<tableCitation id="C6E9CA8FF63D562CFBEEC3BBC087F0A6" box="[1144,1242,151,177]" captionStart="TABLE 1" captionStartId="28.[151,239,483,509]" captionTargetId="graphics-52@28.[151,2042,547,912]" captionTargetPageId="28" captionText="TABLE 1. Cheek-tooth dimensions (mm) of Bohra bandharr. *Anterior end missing enamel; length estimated by extrapolating from anterior edge of anterior root and contours of adjacent enamel." httpUri="http://table.plazi.org/id/DF14AFBCF6055614FF01C2CFC535F236" pageId="36" pageNumber="37" tableUuid="DF14AFBCF6055614FF01C2CFC535F236">Tables 1</tableCitation>
,
<tableCitation id="C6E9CA8FF63D562CFB7DC3BBC0A6F0A6" box="[1259,1275,151,177]" captionStart="TABLE 3" captionStartId="35.[151,239,671,697]" captionTargetId="graphics-24@35.[151,2040,699,927]" captionTargetPageId="35" captionText="TABLE 3. Cheek-tooth dimensions (mm) of Bohra wilkinsonorum." httpUri="http://table.plazi.org/id/DF14AFBCF63A562BFF01C1B3C734F2AE" pageId="36" pageNumber="37" tableUuid="DF14AFBCF63A562BFF01C1B3C734F2AE">3</tableCitation>
). It differs by narrowing less markedly anteriorly, possessing three buccal cingular cuspules and bearing a slightly less pronounced posterobuccal eminence. Although the
<typeStatus id="54D04196F63D562CFDC7C3F3C6EEF0EE" box="[593,691,223,249]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
P3 bears an anterior crest on the posterobuccal eminence (
<figureCitation id="1350E3B1F63D562CFAC7C3F3C4B6F10A" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="36" pageNumber="37">Figure 19DE</figureCitation>
), this is lacking from
<collectionCode id="ED7A67F1F63D562CFE48C22FC642F10A" box="[478,543,259,285]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="36" pageNumber="37" type="Museum">NMV</collectionCode>
<specimenCode id="DBCD574FF63D562CFDB0C22EC6D4F10A" box="[550,649,258,285]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="36" pageNumber="37" type="Museum">P221950</specimenCode>
(
<figureCitation id="1350E3B1F63D562CFD0EC22FC71CF10A" box="[664,833,259,285]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="36" pageNumber="37">Figure 19GK</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD4FF34F63D562CFF51C20BC59FF20E" blockId="36.[151,1437,151,825]" pageId="36" pageNumber="37">
In M12 occlusal outline (
<figureCitation id="1350E3B1F63D562CFE6DC20BC69EF156" box="[507,707,295,321]" captionStart="FIGURE 19" captionStartId="34.[151,250,1804,1828]" captionTargetBox="[153,1433,526,1774]" captionTargetId="figure-176@34.[151,1436,522,1778]" captionTargetPageId="34" captionText="FIGURE 19. Maxillary and upper dental specimens of Bohra wilkinsonorum. QM F43277 (holotype) right juvenile maxilla (AF). A: lateral view. B: mesial view. C: occlusal view. D: stereo occlusal view of unerupted P3 (cast). E: lingual view of unerupted P3 (cast). F: stereo occlusal view of dP23, M12 (cast). NMV P221950, left P3 (GK). G: stereo occlusal view. H: anterior view. I: lingual view. J: posterior view. K: buccal view. NMV P222378, right M1 (LP). L: stereo occlusal view. M: buccal view. N: lingual view. O: anterior view. P: posterior view." figureDoi="http://doi.org/10.5281/zenodo.8012712" httpUri="https://zenodo.org/record/8012712/files/figure.png" pageId="36" pageNumber="37">Figure 19C, F, L</figureCitation>
),
<taxonomicName id="4C6B84B7F63D562CFD4DC20BC7FEF156" authorityName="Dawson" authorityYear="2004" box="[731,931,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63D562CFD4DC20BC7FEF156" box="[731,931,295,321]" italics="true" pageId="36" pageNumber="37">B. wilkinsonorum</emphasis>
</taxonomicName>
resembles the much smaller
<taxonomicName id="4C6B84B7F63D562CFB6CC20BC1C8F156" authorityName="Prideaux &amp; Warburton" authorityYear="2009" box="[1274,1429,295,321]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="nullarbora">
<emphasis id="B91F2326F63D562CFB6CC20BC1C8F156" box="[1274,1429,295,321]" italics="true" pageId="36" pageNumber="37">B. nullarbora</emphasis>
</taxonomicName>
. They share a slightly convex, but essentially quite flat, buccal side of the crown, a trait that also characterises
<taxonomicName id="4C6B84B7F63D562CFA12C267C55CF19E" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F63D562CFA12C267C55CF19E" italics="true" pageId="36" pageNumber="37">B. bandharr</emphasis>
</taxonomicName>
. However, the lingual sides of M12 of
<taxonomicName id="4C6B84B7F63D562CFD47C243C701F19E" box="[721,860,367,393]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bandharr">
<emphasis id="B91F2326F63D562CFD47C243C701F19E" box="[721,860,367,393]" italics="true" pageId="36" pageNumber="37">B. bandharr</emphasis>
</taxonomicName>
are also flat lingually, rather than convex adjacent to the interloph valley. The strongly developed and incurved form of the postpara- and premeta-cristae on M2 distinguishes
<taxonomicName id="4C6B84B7F63D562CFEA4C29BC5A4F1C6" authorityName="Dawson" authorityYear="2004" box="[306,505,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63D562CFEA4C29BC5A4F1C6" box="[306,505,439,465]" italics="true" pageId="36" pageNumber="37">B. wilkinsonorum</emphasis>
</taxonomicName>
from all other species of
<taxonomicName id="4C6B84B7F63D562CFCB5C29BC734F1C6" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[803,873,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63D562CFCB5C29BC734F1C6" box="[803,873,439,465]" italics="true" pageId="36" pageNumber="37">Bohra</emphasis>
</taxonomicName>
. Absence from M2 of a stylar crest in the region of cusp C is a condition shared with
<emphasis id="B91F2326F63D562CFDA2C2F7C706F1E2" box="[564,859,475,501]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="4C6B84B7F63D562CFDA2C2F7C6E0F1E2" box="[564,701,475,501]" class="Mammalia" family="Macropodidae" genus="Silvaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bandharr">B. bandharr</taxonomicName>
,
<taxonomicName id="4C6B84B7F63D562CFD5EC2F7C706F1E2" authorityName=", Prideaux &amp; Warburton" authorityYear="2008" box="[712,859,475,501]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="illuminata">B. illuminata</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6B84B7F63D562CFC04C2F7C076F1E2" authorityName="Prideaux &amp; Warburton" authorityYear="2009" box="[914,1067,475,501]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="nullarbora">
<emphasis id="B91F2326F63D562CFC04C2F7C076F1E2" box="[914,1067,475,501]" italics="true" pageId="36" pageNumber="37">B. nullarbora</emphasis>
</taxonomicName>
, but not
<taxonomicName id="4C6B84B7F63D562CFB06C2F7C081F1E2" authorityName="Prideaux &amp; Warburton" authorityYear="2023" baseAuthorityName="Dawson" baseAuthorityYear="2004" box="[1168,1244,475,501]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bila">
<emphasis id="B91F2326F63D562CFB06C2F7C081F1E2" box="[1168,1244,475,501]" italics="true" pageId="36" pageNumber="37">B. bila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B84B7F63D562CFA84C2F7C125F1E2" authorityName="Prideaux &amp; Warburton" authorityYear="2023" box="[1298,1400,475,501]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="planei">
<emphasis id="B91F2326F63D562CFA84C2F7C125F1E2" box="[1298,1400,475,501]" italics="true" pageId="36" pageNumber="37">B. planei</emphasis>
</taxonomicName>
, in which this crest is distinct.
</paragraph>
<paragraph id="8BD4FF34F63D562CFF51C10FC16BF32E" blockId="36.[151,1437,151,825]" pageId="36" pageNumber="37">
Calcaneus. The referred calcaneus, QM F675, is well preserved and complete except for the caudolateral extremity of the tuber calcanei (
<figureCitation id="1350E3B1F63D562CFD86C16BC6E2F276" box="[528,703,583,609]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B, D</figureCitation>
). The calcaneus is large, stout, broad, stout and dorsoventrally flattened, with clearly defined articular surfaces and rugged ligamentous insertions. The tuber calcanei is strongly flared caudally (
<figureCitation id="1350E3B1F63D562CFEDEC1A3C597F2BE" box="[328,458,655,681]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B</figureCitation>
). From the plantar view, the medial margin diverges from the long axis more continuously along its length, in comparison to the lateral margin, which widens distinctly at the epiphysis. The rugose plantar surface is broad cranially, with a large anterolateral tubercle (
<figureCitation id="1350E3B1F63D562CFCDEC1FBC793F2E6" box="[840,974,727,753]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20A</figureCitation>
). The transverse sulcus that separates the cranial margin of the rugose surface from the cuboid articulation is slightly oblique from medial to lateral sides. The sustentaculum tali is deep (from medial view), medially expanded and very slightly convex (
<figureCitation id="1350E3B1F63D562CFB32C033C174F32E" box="[1188,1321,799,825]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20C</figureCitation>
).
</paragraph>
<caption id="DF14AFBCF63D562CFF01C66FC5E5F597" ID-DOI="http://doi.org/10.5281/zenodo.8012714" ID-Zenodo-Dep="8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1347,1371]" targetBox="[166,1418,846,1316]" targetPageId="36" targetType="figure">
<paragraph id="8BD4FF34F63D562CFF01C66FC5E5F597" blockId="36.[151,1437,1347,1408]" pageId="36" pageNumber="37">
<emphasis id="B91F2326F63D562CFF01C66FC57DF54C" bold="true" box="[151,288,1347,1371]" pageId="36" pageNumber="37">FIGURE 20.</emphasis>
Calcaneus (right, adult) of
<taxonomicName id="4C6B84B7F63D562CFDA3C668C74FF54B" authorityName="Dawson" authorityYear="2004" box="[565,786,1348,1372]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63D562CFDA3C668C74FF54B" box="[565,786,1348,1372]" italics="true" pageId="36" pageNumber="37">Bohra wilkinsonorum</emphasis>
</taxonomicName>
(QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view.
</paragraph>
</caption>
<paragraph id="8BD4FF34F63D562CFF51C682C67FF743" blockId="36.[151,1437,1454,2020]" pageId="36" pageNumber="37">
The medial and lateral talar facets are separated by a narrow but distinct, non-articular area (
<figureCitation id="1350E3B1F63D562CFB28C682C102F5DF" box="[1214,1375,1454,1480]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B, E</figureCitation>
). The lateral facet is barrel-shaped (
<figureCitation id="1350E3B1F63D562CFE75C6FEC635F5FB" box="[483,616,1490,1516]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B</figureCitation>
). The fibular facet is broad and rounded, and projects over the deep sulci for the posterior calcaneofibular ligament (
<figureCitation id="1350E3B1F63D562CFDE1C6DAC740F607" box="[631,797,1526,1552]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B, D</figureCitation>
). The tubercle and sulcus for the anterior calcaneofibular ligament are large, laterally expanded and rugose. The posterior margin of the oval medial talar facet projects dorsally and expands onto the dorsal surface toward the base of the calcaneal tubercle, with a marked projection for posterior talocalcaneal ligament (
<figureCitation id="1350E3B1F63D562CFDAFC54EC6E1F66B" box="[569,700,1634,1660]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B</figureCitation>
). Cranial to the talar facets, the mid-dorsal surface is has a deep scar for the ligamentum cervicis tali. The calcaneocuboid articulation is transversely very broad (laterally extending beyond the margin of the fibular facet;
<figureCitation id="1350E3B1F63D562CFDC3C586C685F6D3" box="[597,728,1706,1732]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20B</figureCitation>
); and dorsoventrally compressed in cranial view (
<figureCitation id="1350E3B1F63D562CFA9AC586C1CDF6D3" box="[1292,1424,1706,1732]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20E</figureCitation>
). The step between the rectangular dorsomedial and rounded dorsolateral facets is distinct, obliquely aligned and moderately smoothed (
<figureCitation id="1350E3B1F63D562CFE09C5DEC674F71B" box="[415,553,1778,1804]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20A</figureCitation>
). The ventromedial facet is transversely broad, but dorsoventrally relatively shallow (
<figureCitation id="1350E3B1F63D562CFE97C43AC5D5F727" box="[257,392,1814,1840]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="36" pageNumber="37">Figure 20E</figureCitation>
). It is smoothly continuous with the dorsolateral facet but separated by a deep transverse groove from the dorsomedial facet.
</paragraph>
<paragraph id="8BD4FF34F63D562DFF51C472C5C1F1BB" blockId="36.[151,1437,1454,2020]" lastBlockId="37.[151,1437,151,825]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
The lateral flaring of the tuber calcanei epiphysis is more pronounced than in all other species of
<taxonomicName id="4C6B84B7F63D562CFAD9C472C1C8F76F" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1359,1429,1886,1912]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63D562CFAD9C472C1C8F76F" box="[1359,1429,1886,1912]" italics="true" pageId="36" pageNumber="37">Bohra</emphasis>
</taxonomicName>
, especially the similarly large
<taxonomicName id="4C6B84B7F63D562CFE70C4AFC60EF78B" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[486,595,1922,1948]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="paulae">
<emphasis id="B91F2326F63D562CFE70C4AFC60EF78B" box="[486,595,1922,1948]" italics="true" pageId="36" pageNumber="37">B. paulae</emphasis>
</taxonomicName>
(
<figureCitation id="1350E3B1F63D562CFDF5C4AEC6BBF78B" box="[611,742,1922,1948]" captionStart="FIGURE 12" captionStartId="23.[151,250,1904,1928]" captionTargetBox="[217,1367,183,1876]" captionTargetId="figure-17@23.[216,1371,181,1880]" captionTargetPageId="23" captionText="FIGURE 12. Calcaneus and talus (right, adult) of the holotype of Bohra paulae (AM F62099F62100). Calcaneus (AE). A: dorsal view. B: plantar view. C: medial view. D: lateral view. E: cranial view. Calcaneus and talus in articulation (FI). F: dorsal view. G: medial view. H: cranial view. I: lateral view. Talus (JO). J: cranial view. K: dorsal view. L: medial view. M: caudal view. N: plantar view. O: lateral view." figureDoi="http://doi.org/10.5281/zenodo.8012698" httpUri="https://zenodo.org/record/8012698/files/figure.png" pageId="36" pageNumber="37">Figure 12B</figureCitation>
). The transverse plantar sulcus is relatively deep and long in comparison to all other species of
<emphasis id="B91F2326F63D562CFD81C48AC639F7D7" box="[535,612,1958,1984]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="4C6B84B7F63D562CFD81C48AC63DF7D7" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[535,608,1958,1984]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">Bohra</taxonomicName>
,
</emphasis>
in particular
<taxonomicName id="4C6B84B7F63D562CFD6CC48BC738F7D7" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[762,869,1958,1984]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="paulae">
<emphasis id="B91F2326F63D562CFD6CC48BC738F7D7" box="[762,869,1958,1984]" italics="true" pageId="36" pageNumber="37">B. paulae</emphasis>
</taxonomicName>
and QM F51762 (
<taxonomicName id="4C6B84B7F63D562CFBA0C48AC0FEF7D7" box="[1078,1187,1958,1984]" class="Mammalia" family="Macropodidae" genus="Bohra" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B91F2326F63D562CFBA0C48AC021F7D7" box="[1078,1148,1958,1984]" italics="true" pageId="36" pageNumber="37">Bohra</emphasis>
sp.
</taxonomicName>
indet. 1), and is much deeper than in the species of
<taxonomicName id="4C6B84B7F63D562CFE77C4E6C62FF7F3" box="[481,626,1994,2020]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63D562CFE77C4E6C62FF7F3" box="[481,626,1994,2020]" italics="true" pageId="36" pageNumber="37">Dendrolagus</emphasis>
</taxonomicName>
(
<figureCitation id="1350E3B1F63D562CFD17C4E6C6A5F7F3" box="[641,760,1994,2020]" captionStart="FIGURE 9" captionStartId="18.[151,250,1670,1694]" captionTargetBox="[297,1289,181,1633]" captionTargetId="figure-26@18.[296,1292,181,1645]" captionTargetPageId="18" captionText="FIGURE 9. Calcaneus and talus (right, adult) of Dendrolagus bennettianus (QM JM19863). Calcaneus (AE). A: dorsal view. B: plantar view. C: medial view. D: cranial view. E: lateral view. Talus (FK). F. plantar view. G: medial view. H: dorsal view. I: caudal view. J: lateral view. K: cranial view. AHN, head of talus (articular facet for navicular); APT, anterior plantar tuberosity; ATF, anteromedial facet for head of talus; CFA, articular facet for cuboid; CPS, calcaneal plantar surface; DMF, dorsomedial facet of calcaneal-cuboid articulation; DLF, dorsolateral facet of calcaneal-cuboid articulation; FGS, flexor groove of sustentaculum tali; FIB, fibular articular surface; FIL, tubercles for fibular ligaments; LCS, lateral calcaneal facet; LTF, lateral facet for talus; LTC, lateral trochlear crest; MCF, medial calcaneal facet; MFO, malleolar fossa; MMA, medial malleolus; MTC, medial trochlear crest; MTF, medial facet for talus; PVP, posteroventral process; STA, sustentaculum tali; TCA, tuber calcanei; TCE, epiphysis of tuber calcanei; TPS, transverse plantar sulcus; TRO, trochlea (talo-crural joint); VMF, ventromedial facet of calcaneal-cuboid articulation." figureDoi="http://doi.org/10.5281/zenodo.8012688" httpUri="https://zenodo.org/record/8012688/files/figure.png" pageId="36" pageNumber="37">Figure 9B</figureCitation>
). The expanded medial plantar margin of the sustentaculum tali accentuates the moderately deep flexor sulcus of the sustentaculum tali to a greater extent than in
<emphasis id="B91F2326F63C562DFABEC3BBC4EDF0C3" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="4C6B84B7F63C562DFABEC3BBC1C5F0A6" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1320,1432,151,177]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="paulae">B. paulae</taxonomicName>
, B.
</emphasis>
sp. indet. 1 and the species of
<taxonomicName id="4C6B84B7F63C562DFD85C397C6F9F0C2" box="[531,676,187,213]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFD85C397C6F9F0C2" box="[531,676,187,213]" italics="true" pageId="37" pageNumber="38">Dendrolagus</emphasis>
</taxonomicName>
, though it is not as deeply flared as in
<taxonomicName id="4C6B84B7F63C562DFBF2C397C096F0C2" authorityName="Prideaux &amp; Warburton" authorityYear="2023" box="[1124,1227,187,213]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="planei">
<emphasis id="B91F2326F63C562DFBF2C397C096F0C2" box="[1124,1227,187,213]" italics="true" pageId="37" pageNumber="38">B. planei</emphasis>
</taxonomicName>
. The gap between the medial and lateral talar facets resembles that observed in
<taxonomicName id="4C6B84B7F63C562DFCDBC3F3C7BBF0EE" authorityName="Prideaux &amp; Warburton" authorityYear="2009" box="[845,998,223,249]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="nullarbora">
<emphasis id="B91F2326F63C562DFCDBC3F3C7BBF0EE" box="[845,998,223,249]" italics="true" pageId="37" pageNumber="38">B. nullarbora</emphasis>
</taxonomicName>
, but differs from the condition seen in other species of
<taxonomicName id="4C6B84B7F63C562DFEDAC22FC5CFF10A" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[332,402,259,285]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFEDAC22FC5CFF10A" box="[332,402,259,285]" italics="true" pageId="37" pageNumber="38">Bohra</emphasis>
</taxonomicName>
and species
<taxonomicName id="4C6B84B7F63C562DFD88C22FC6F2F10A" box="[542,687,259,285]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFD88C22FC6F2F10A" box="[542,687,259,285]" italics="true" pageId="37" pageNumber="38">Dendrolagus</emphasis>
</taxonomicName>
(
<figureCitation id="1350E3B1F63C562DFD2AC22FC76EF10A" box="[700,819,259,285]" captionStart="FIGURE 9" captionStartId="18.[151,250,1670,1694]" captionTargetBox="[297,1289,181,1633]" captionTargetId="figure-26@18.[296,1292,181,1645]" captionTargetPageId="18" captionText="FIGURE 9. Calcaneus and talus (right, adult) of Dendrolagus bennettianus (QM JM19863). Calcaneus (AE). A: dorsal view. B: plantar view. C: medial view. D: cranial view. E: lateral view. Talus (FK). F. plantar view. G: medial view. H: dorsal view. I: caudal view. J: lateral view. K: cranial view. AHN, head of talus (articular facet for navicular); APT, anterior plantar tuberosity; ATF, anteromedial facet for head of talus; CFA, articular facet for cuboid; CPS, calcaneal plantar surface; DMF, dorsomedial facet of calcaneal-cuboid articulation; DLF, dorsolateral facet of calcaneal-cuboid articulation; FGS, flexor groove of sustentaculum tali; FIB, fibular articular surface; FIL, tubercles for fibular ligaments; LCS, lateral calcaneal facet; LTF, lateral facet for talus; LTC, lateral trochlear crest; MCF, medial calcaneal facet; MFO, malleolar fossa; MMA, medial malleolus; MTC, medial trochlear crest; MTF, medial facet for talus; PVP, posteroventral process; STA, sustentaculum tali; TCA, tuber calcanei; TCE, epiphysis of tuber calcanei; TPS, transverse plantar sulcus; TRO, trochlea (talo-crural joint); VMF, ventromedial facet of calcaneal-cuboid articulation." figureDoi="http://doi.org/10.5281/zenodo.8012688" httpUri="https://zenodo.org/record/8012688/files/figure.png" pageId="37" pageNumber="38">Figure 9A</figureCitation>
), in which the facets are confluent. The calcaneocuboid articulation is transversely much broader than in any other species (
<figureCitation id="1350E3B1F63C562DFC1AC20BC076F156" box="[908,1067,295,321]" captionStart="FIGURE 20" captionStartId="36.[151,250,1347,1371]" captionTargetBox="[166,1418,846,1316]" captionTargetId="figure-341@36.[164,1423,846,1321]" captionTargetPageId="36" captionText="FIGURE 20. Calcaneus (right, adult) of Bohra wilkinsonorum (QM F675). A: plantar view. B: dorsal view. C: medial view. D: lateral view. E: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012714" httpUri="https://zenodo.org/record/8012714/files/figure.png" pageId="37" pageNumber="38">Figure 20B, E</figureCitation>
cf.
<figureCitation id="1350E3B1F63C562DFBC0C20BC0B0F156" box="[1110,1261,295,321]" captionStart="FIGURE 9" captionStartId="18.[151,250,1670,1694]" captionTargetBox="[297,1289,181,1633]" captionTargetId="figure-26@18.[296,1292,181,1645]" captionTargetPageId="18" captionText="FIGURE 9. Calcaneus and talus (right, adult) of Dendrolagus bennettianus (QM JM19863). Calcaneus (AE). A: dorsal view. B: plantar view. C: medial view. D: cranial view. E: lateral view. Talus (FK). F. plantar view. G: medial view. H: dorsal view. I: caudal view. J: lateral view. K: cranial view. AHN, head of talus (articular facet for navicular); APT, anterior plantar tuberosity; ATF, anteromedial facet for head of talus; CFA, articular facet for cuboid; CPS, calcaneal plantar surface; DMF, dorsomedial facet of calcaneal-cuboid articulation; DLF, dorsolateral facet of calcaneal-cuboid articulation; FGS, flexor groove of sustentaculum tali; FIB, fibular articular surface; FIL, tubercles for fibular ligaments; LCS, lateral calcaneal facet; LTF, lateral facet for talus; LTC, lateral trochlear crest; MCF, medial calcaneal facet; MFO, malleolar fossa; MMA, medial malleolus; MTC, medial trochlear crest; MTF, medial facet for talus; PVP, posteroventral process; STA, sustentaculum tali; TCA, tuber calcanei; TCE, epiphysis of tuber calcanei; TPS, transverse plantar sulcus; TRO, trochlea (talo-crural joint); VMF, ventromedial facet of calcaneal-cuboid articulation." figureDoi="http://doi.org/10.5281/zenodo.8012688" httpUri="https://zenodo.org/record/8012688/files/figure.png" pageId="37" pageNumber="38">Figure 9B, D</figureCitation>
). The profile of the calcaneocuboid step and complete separation of the ventromedial and dorsomedial facets is most like
<emphasis id="B91F2326F63C562DFAA6C267C1C1F172" box="[1328,1436,331,357]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="4C6B84B7F63C562DFAA6C267C1C5F172" authorityName="Prideaux &amp; Warburton" authorityYear="2023" box="[1328,1432,331,357]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="planei">B. planei</taxonomicName>
,
</emphasis>
while the rounded and laterally expanded of the dorsolateral facet is more like other species of
<emphasis id="B91F2326F63C562DFB21C243C159F19E" box="[1207,1284,367,393]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="4C6B84B7F63C562DFB21C243C15DF19E" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1207,1280,367,393]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">Bohra</taxonomicName>
,
</emphasis>
in contrast to
<taxonomicName id="4C6B84B7F63C562DFF01C2BFC4A1F1BA" authorityName="Prideaux &amp; Warburton" authorityYear="2023" box="[151,252,403,429]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="planei">
<emphasis id="B91F2326F63C562DFF01C2BFC4A1F1BA" box="[151,252,403,429]" italics="true" pageId="37" pageNumber="38">B. planei</emphasis>
</taxonomicName>
(
<figureCitation id="1350E3B1F63C562DFE9DC2BFC5D1F1BA" box="[267,396,403,429]" captionStart="FIGURE 33" captionStartId="62.[151,250,1893,1917]" captionTargetBox="[166,1421,598,1868]" captionTargetId="figure-236@62.[164,1423,597,1870]" captionTargetPageId="62" captionText="FIGURE 33. Calcaneus and talus (right, adult) of the holotype of Bohra planei sp. nov. UCMP 70128, holotype calcaneus (AE). A: plantar view. B: dorsal view. C: cranial view. D: medial view. E: lateral view. NHMD 193283, partial right calcaneus (FG). F: dorsal view. G: medial view. UCMP 70128, holotype talus (JM). H: plantar view. I: dorsal view. J: medial view. K: lateral view. L: caudal view. M: cranial view." figureDoi="http://doi.org/10.5281/zenodo.8012754" httpUri="https://zenodo.org/record/8012754/files/figure.png" pageId="37" pageNumber="38">Figure 33C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C371ACBFF63C562DFF51C29BC7A1F32E" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BD4FF34F63C562DFF51C29BC7A1F32E" blockId="37.[151,1437,151,825]" pageId="37" pageNumber="38">
<emphasis id="B91F2326F63C562DFF51C29BC566F1C6" bold="true" box="[199,315,439,465]" pageId="37" pageNumber="38">Remarks.</emphasis>
The recognition of
<taxonomicName id="4C6B84B7F63C562DFDB6C29BC6BAF1C6" authorityName="Dawson" authorityYear="2004" box="[544,743,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63C562DFDB6C29BC6BAF1C6" box="[544,743,439,465]" italics="true" pageId="37" pageNumber="38">B. wilkinsonorum</emphasis>
</taxonomicName>
as a second member of the genus
<taxonomicName id="4C6B84B7F63C562DFBE3C29BC0E6F1C6" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1141,1211,439,465]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFBE3C29BC0E6F1C6" box="[1141,1211,439,465]" italics="true" pageId="37" pageNumber="38">Bohra</emphasis>
</taxonomicName>
on the basis of
<specimenCount id="9D6D34BDF63C562DFAE2C29BC4ACF1E2" pageId="37" pageNumber="38" type="juvenile">one juvenile</specimenCount>
maxilla fragment (
<bibRefCitation id="EFFA82C5F63C562DFE45C2F7C6DFF1E2" author="Dawson, L." box="[467,642,475,501]" pageId="37" pageNumber="38" pagination="267 - 273" refId="ref49751" refString="Dawson, L. (2004 a) A new Pliocene tree kangaroo species (Marsupialia, Macropodinae) from the Chinchilla Local Fauna, southeastern Queensland. Alcheringa, 28 (1), 267 - 273. https: // doi. org / 10.1080 / 03115510408619284" type="journal article" year="2004">Dawson 2004a</bibRefCitation>
) was a highly perceptive deduction given that the
<typeStatus id="54D04196F63C562DFB4CC2F7C157F1E2" box="[1242,1290,475,501]" pageId="37" pageNumber="38">type</typeStatus>
species was described and is still known only from hindlimb elements from a locality
<quantity id="4C9352D1F63C562DFC41C2D2C070F20E" box="[983,1069,510,537]" metricMagnitude="5" metricUnit="m" metricValue="6.65" pageId="37" pageNumber="38" unit="km" value="665.0">665 km</quantity>
to the south (
<bibRefCitation id="EFFA82C5F63C562DFB51C2D3C48EF22A" author="Flannery, T. F. &amp; Szalay, F. S." pageId="37" pageNumber="38" pagination="83 - 94" refId="ref50999" refString="Flannery, T. F. &amp; Szalay, F. S. (1982) Bohra paulae: a new giant fossil tree kangaroo (Marsupialia: Macropodidae) from New South Wales, Australia. Australian Mammalogy, 5 (2), 83 - 94." type="journal article" year="1982">Flannery &amp; Szalay 1982</bibRefCitation>
). Based on the dimensions of these specimens (
<tableCitation id="C6E9CA8FF63C562DFD7BC10FC71EF22A" box="[749,835,547,573]" captionStart="TABLE 8" captionStartId="52.[151,239,493,519]" captionTargetPageId="52" captionText="TABLE 8. Postcranial element dimensions (mm) for representative specimens of species of Bohra and Dendrolagus. Abbreviation: est., estimated." httpUri="http://table.plazi.org/id/DF14AFBCF62D563CFF01C2C1C2EAF210" pageId="37" pageNumber="38" tableUuid="DF14AFBCF62D563CFF01C2C1C2EAF210">Table 8</tableCitation>
),
<taxonomicName id="4C6B84B7F63C562DFCCCC10FC07DF22A" authorityName="Dawson" authorityYear="2004" box="[858,1056,547,573]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63C562DFCCCC10FC07DF22A" box="[858,1056,547,573]" italics="true" pageId="37" pageNumber="38">B. wilkinsonorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B84B7F63C562DFBCFC10FC099F22A" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1113,1220,547,573]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="paulae">
<emphasis id="B91F2326F63C562DFBCFC10FC099F22A" box="[1113,1220,547,573]" italics="true" pageId="37" pageNumber="38">B. paulae</emphasis>
</taxonomicName>
are the two largest species of
<taxonomicName id="4C6B84B7F63C562DFE9AC16BC50FF276" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[268,338,583,609]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFE9AC16BC50FF276" box="[268,338,583,609]" italics="true" pageId="37" pageNumber="38">Bohra</emphasis>
</taxonomicName>
, and one might be able to hypothesise on those grounds that the fossils belong to the same species. However, the recognition from the Darling Downs region (indeed, probably from the
<typeStatus id="54D04196F63C562DFBCAC147C0D1F292" box="[1116,1164,619,645]" pageId="37" pageNumber="38">type</typeStatus>
locality) of a very large calcaneus clearly referable to a species of
<taxonomicName id="4C6B84B7F63C562DFDF8C1A3C6E9F2BE" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[622,692,655,681]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F63C562DFDF8C1A3C6E9F2BE" box="[622,692,655,681]" italics="true" pageId="37" pageNumber="38">Bohra</emphasis>
</taxonomicName>
that is morphologically distinct from that of
<taxonomicName id="4C6B84B7F63C562DFB3CC1A3C148F2BE" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1194,1301,655,681]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="paulae">
<emphasis id="B91F2326F63C562DFB3CC1A3C148F2BE" box="[1194,1301,655,681]" italics="true" pageId="37" pageNumber="38">B. paulae</emphasis>
</taxonomicName>
supports its taxonomic separation. This calcaneus is much larger than, and differs in morphology from, the other two calcanei from the Chinchilla LF, which we refer to
<emphasis id="B91F2326F63C562DFDEDC1FBC6CEF2E7" box="[635,659,727,752]" italics="true" pageId="37" pageNumber="38">B.</emphasis>
sp. indet. 1. Referral to
<taxonomicName id="4C6B84B7F63C562DFC3FC1FBC032F2E6" authorityName="Dawson" authorityYear="2004" box="[937,1135,727,753]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="wilkinsonorum">
<emphasis id="B91F2326F63C562DFC3FC1FBC032F2E6" box="[937,1135,727,753]" italics="true" pageId="37" pageNumber="38">B. wilkinsonorum</emphasis>
</taxonomicName>
of a loose upper premolar and molar from the putatively late Pliocene or early Pleistocene Childers Cove LF of southern
<collectingRegion id="49AF31D6F63C562DFB28C1D7C145F302" box="[1214,1304,763,789]" country="Australia" name="Victoria" pageId="37" pageNumber="38">Victoria</collectingRegion>
(
<bibRefCitation id="EFFA82C5F63C562DFAB0C1D7C48CF32E" author="Rich, T. H. &amp; Piper, K. J. &amp; Pickering, D. &amp; Wright, S." pageId="37" pageNumber="38" pagination="133 - 140" refId="ref56292" refString="Rich, T. H., Piper, K. J., Pickering, D. &amp; Wright, S. (2006) Further Ektopodontidae (Phalangeroidea, Mammalia) from southwestern Victoria. Alcheringa, 30 (1), 133 - 140. https: // doi. org / 10.1080 / 03115510608619349" type="journal article" year="2006">
Rich
<emphasis id="B91F2326F63C562DFAF2C1D7C1C0F302" box="[1380,1437,763,789]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2006
</bibRefCitation>
) represents a major range extension of
<quantity id="4C9352D1F63C562DFD1CC032C6A9F32E" box="[650,756,798,825]" metricMagnitude="6" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="km" value="1500.0">1,500 km</quantity>
to the south (
<figureCitation id="1350E3B1F63C562DFC1AC033C7B6F32E" box="[908,1003,799,825]" captionStart="FIGURE 1" captionStartId="5.[151,250,1691,1715]" captionTargetBox="[207,1381,795,1665]" captionTargetId="figure-278@5.[207,1381,795,1665]" captionTargetPageId="5" captionText="FIGURE 1. Map of Australia and New Guinea showing localities that have produced fossils attributable, or potentially attributable, to species of Bohra." figureDoi="http://doi.org/10.5281/zenodo.8012668" httpUri="https://zenodo.org/record/8012668/files/figure.png" pageId="37" pageNumber="38">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>