treatments-xml/data/03/C2/4E/03C24E22F60D561CFF01C3BBC6B7F41B.xml
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<mods:title id="050586E5C25F268457519C78E4A3AB7A">A review of the late Cenozoic genus Bohra (Diprotodontia: Macropodidae) and the evolution of tree-kangaroos</mods:title>
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<treatment id="03C24E22F60D561CFF01C3BBC6B7F41B" ID-DOI="http://doi.org/10.5281/zenodo.8017920" ID-GBIF-Taxon="209675083" ID-Zenodo-Dep="8017920" LSID="urn:lsid:plazi:treatment:03C24E22F60D561CFF01C3BBC6B7F41B" httpUri="http://treatment.plazi.org/id/03C24E22F60D561CFF01C3BBC6B7F41B" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C371ACBFF60D561CFF01C3BBC643F0A6" box="[151,542,151,178]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BD4FF34F60D561CFF01C3BBC643F0A6" blockId="20.[151,542,151,178]" box="[151,542,151,178]" pageId="20" pageNumber="21">
<heading id="D09C4858F60D561CFF01C3BBC643F0A6" bold="true" box="[151,542,151,178]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
<taxonomicName id="4C6B84B7F60D561CFF01C3BBC643F0A6" authority="Flannery &amp; Szalay, 1982" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[151,542,151,178]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFF01C3BBC643F0A6" bold="true" box="[151,542,151,178]" pageId="20" pageNumber="21">
<emphasis id="B91F2326F60D561CFF01C3BBC4B9F0A5" bold="true" box="[151,228,151,178]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
<bibRefCitation id="EFFA82C5F60D561CFF7DC3BBC643F0A6" author="Flannery, T. F. &amp; Szalay, F. S." box="[235,542,151,177]" pageId="20" pageNumber="21" pagination="83 - 94" refId="ref50999" refString="Flannery, T. F. &amp; Szalay, F. S. (1982) Bohra paulae: a new giant fossil tree kangaroo (Marsupialia: Macropodidae) from New South Wales, Australia. Australian Mammalogy, 5 (2), 83 - 94." type="journal article" year="1982">Flannery &amp; Szalay, 1982</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C371ACBFF60D561CFF01C3F0C75BF2DF" pageId="20" pageNumber="21" type="type_taxon">
<paragraph id="8BD4FF34F60D561CFF01C3F0C6E0F0E3" blockId="20.[151,701,219,244]" box="[151,701,219,244]" pageId="20" pageNumber="21">
<emphasis id="B91F2326F60D561CFF01C3F0C57EF0E4" bold="true" box="[151,291,219,244]" pageId="20" pageNumber="21">
<typeStatus id="54D04196F60D561CFF01C3F0C491F0E3" box="[151,204,220,244]" pageId="20" pageNumber="21">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6B84B7F60D561CFEBCC3F0C6E7F0E3" authority="Flannery &amp; Szalay, 1982" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[298,698,220,244]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="paulae">
<emphasis id="B91F2326F60D561CFEBCC3F0C5E9F0E3" box="[298,436,220,244]" italics="true" pageId="20" pageNumber="21">Bohra paulae</emphasis>
<bibRefCitation id="EFFA82C5F60D561CFE2DC3F0C6E7F0E3" author="Flannery, T. F. &amp; Szalay, F. S." box="[443,698,220,244]" pageId="20" pageNumber="21" pagination="83 - 94" refId="ref50999" refString="Flannery, T. F. &amp; Szalay, F. S. (1982) Bohra paulae: a new giant fossil tree kangaroo (Marsupialia: Macropodidae) from New South Wales, Australia. Australian Mammalogy, 5 (2), 83 - 94." type="journal article" year="1982">Flannery &amp; Szalay, 1982</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4FF34F60D561CFF01C20DC7D5F203" blockId="20.[151,1437,289,1036]" pageId="20" pageNumber="21">
<emphasis id="B91F2326F60D561CFF01C20DC525F12C" bold="true" box="[151,376,289,315]" pageId="20" pageNumber="21">Revised diagnosis.</emphasis>
Species of
<taxonomicName id="4C6B84B7F60D561CFD99C20EC608F12B" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[527,597,290,316]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFD99C20EC608F12B" box="[527,597,290,316]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
</taxonomicName>
are distinguished from those of
<taxonomicName id="4C6B84B7F60D561CFC7EC20EC024F12B" ID-CoL="62PHB" authorityName="Muller" authorityYear="1840" box="[1000,1145,290,316]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFC7EC20EC024F12B" box="[1000,1145,290,316]" italics="true" pageId="20" pageNumber="21">Dendrolagus</emphasis>
</taxonomicName>
based on the following craniodental features: larger cheek teeth relative to the size of the cranium; higher-crowned upper incisors; narrower P3, which lacks a large, distinct posterobuccal accessory cusp and deep cleft posterior to the anterior cusp of the main crest; relatively narrower molars, with a better developed postparacrista and no parametacristid. All known species of
<taxonomicName id="4C6B84B7F60D561CFE9AC29EC50FF1DB" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[268,338,434,460]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFE9AC29EC50FF1DB" box="[268,338,434,460]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
</taxonomicName>
are also larger than those of
<taxonomicName id="4C6B84B7F60D561CFD06C29EC77CF1DB" ID-CoL="62PHB" authorityName="Muller" authorityYear="1840" box="[656,801,434,460]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFD06C29EC77CF1DB" box="[656,801,434,460]" italics="true" pageId="20" pageNumber="21">Dendrolagus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6B84B7F60D561CFCB9C29EC728F1DB" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[815,885,434,460]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFCB9C29EC728F1DB" box="[815,885,434,460]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C6B84B7F60D561CFB15C29FC0A8F1DB" ID-CoL="632VV" authorityName="Schlegel &amp; Muller" authorityYear="1845" box="[1155,1269,435,460]" class="Mammalia" family="Macropodidae" genus="Dorcopsis" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFB15C29FC0A8F1DB" box="[1155,1269,435,460]" italics="true" pageId="20" pageNumber="21">Dorcopsis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B84B7F60D561CFABCC29FC1DDF1DB" authorityName="Flannery, Hoch &amp; Aplin" authorityYear="1989" box="[1322,1408,435,460]" class="Mammalia" family="Macropodidae" genus="Watutia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFABCC29FC1DDF1DB" box="[1322,1408,435,460]" italics="true" pageId="20" pageNumber="21">Watutia</emphasis>
</taxonomicName>
in molar morphology, but the premetacristid, cristid obliqua and postprotocrista are less developed than in
<taxonomicName id="4C6B84B7F60D561CFAB5C2FBC1C8F1E7" ID-CoL="632VV" authorityName="Schlegel &amp; Muller" authorityYear="1845" box="[1315,1429,471,496]" class="Mammalia" family="Macropodidae" genus="Dorcopsis" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFAB5C2FBC1C8F1E7" box="[1315,1429,471,496]" italics="true" pageId="20" pageNumber="21">Dorcopsis</emphasis>
</taxonomicName>
, and the molars are narrower relative to their length than in
<taxonomicName id="4C6B84B7F60D561CFCBDC2D7C7DCF203" authorityName="Flannery, Hoch &amp; Aplin" authorityYear="1989" box="[811,897,507,532]" class="Mammalia" family="Macropodidae" genus="Watutia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFCBDC2D7C7DCF203" box="[811,897,507,532]" italics="true" pageId="20" pageNumber="21">Watutia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD4FF34F60D561CFF51C132C75BF2DF" blockId="20.[151,1437,289,1036]" pageId="20" pageNumber="21">
Species of
<taxonomicName id="4C6B84B7F60D561CFED0C132C5D1F22F" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[326,396,542,568]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFED0C132C5D1F22F" box="[326,396,542,568]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
</taxonomicName>
may be distinguished from those of
<taxonomicName id="4C6B84B7F60D561CFCA2C132C798F22F" ID-CoL="62PHB" authorityName="Muller" authorityYear="1840" box="[820,965,542,568]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFCA2C132C798F22F" box="[820,965,542,568]" italics="true" pageId="20" pageNumber="21">Dendrolagus</emphasis>
</taxonomicName>
on the basis of the following postcranial features: slightly less robust humerus, with a more pronounced pectoral crest, a slightly less developed deltoid tubercle, and a larger, more projecting medial epicondyle; relatively more robust femoral diaphysis; less sinuous tibial diaphysis with a relatively longer distal fibular facet; less obliquely orientated trochlear crests and groove on the talus; shorter and more robust metatarsals IV and V.
</paragraph>
</subSubSection>
<subSubSection id="C371ACBFF60D561CFF51C1FEC6B7F41B" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BD4FF34F60D561CFF51C1FEC76EF36B" blockId="20.[151,1437,289,1036]" pageId="20" pageNumber="21">
<emphasis id="B91F2326F60D561CFF51C1FEC513F2FB" bold="true" box="[199,334,722,748]" pageId="20" pageNumber="21">Etymology.</emphasis>
According to a legend of the Euahlayi people of inland eastern
<collectingCountry id="F37CBFA4F60D561CFBB1C1FEC0CCF2FB" box="[1063,1169,722,748]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
,
<taxonomicName id="4C6B84B7F60D561CFB0BC1FEC0BEF2FB" authorityName="Flannery &amp; Szalay" authorityYear="1982" box="[1181,1251,722,748]" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B91F2326F60D561CFB0BC1FEC0BEF2FB" box="[1181,1251,722,748]" italics="true" pageId="20" pageNumber="21">Bohra</emphasis>
</taxonomicName>
was a kangaroo that went about on four limbs and possessed canines before men removed them (
<bibRefCitation id="EFFA82C5F60D561CFBB0C1DAC0E8F306" author="Parker, K. L." box="[1062,1205,758,785]" pageId="20" pageNumber="21" refId="ref54520" refString="Parker, K. L. (1953) Australian Legendary Tales. Halstead Press, Sydney, 237 pp." type="book" year="1953">Parker 1953</bibRefCitation>
).
<bibRefCitation id="EFFA82C5F60D561CFB5FC1DAC4BFF323" author="Flannery, T. F. &amp; Szalay, F. S." pageId="20" pageNumber="21" pagination="83 - 94" refId="ref50999" refString="Flannery, T. F. &amp; Szalay, F. S. (1982) Bohra paulae: a new giant fossil tree kangaroo (Marsupialia: Macropodidae) from New South Wales, Australia. Australian Mammalogy, 5 (2), 83 - 94." type="journal article" year="1982">Flannery &amp; Szalay (1982)</bibRefCitation>
considered this evocative of extant tree-kangaroos, which have relatively more equally proportioned fore- and hindlimbs and retain small canines. They did not designate a gender, but with the naming of
<taxonomicName id="4C6B84B7F60D561CFA96C013C5BAF36B" authority=", Prideaux &amp; Warburton (2008)" authorityName=", Prideaux &amp; Warburton" authorityYear="2008" class="Mammalia" family="Macropodidae" genus="Bohra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="illuminata">
<emphasis id="B91F2326F60D561CFA96C013C1C8F34F" box="[1280,1429,830,856]" italics="true" pageId="20" pageNumber="21">B. illuminata</emphasis>
,
<bibRefCitation id="EFFA82C5F60D561CFF01C04EC5BAF36B" author="Prideaux, G. J. &amp; Warburton, N. M." box="[151,487,866,892]" pageId="20" pageNumber="21" pagination="463 - 478" refId="ref55336" refString="Prideaux, G. J. &amp; Warburton, N. M. (2008) A new Pleistocene tree-kangaroo (Diprotodontia: Macropodidae) from the Nullarbor Plain of south-central Australia. Journal of Vertebrate Paleontology, 28 (2), 463 - 478. https: // doi. org / 10.1671 / 0272 - 4634 (2008) 28 [463: ANPTDM] 2.0. CO; 2" type="journal article" year="2008">Prideaux &amp; Warburton (2008)</bibRefCitation>
</taxonomicName>
inferred that it was feminine.
</paragraph>
<paragraph id="8BD4FF34F60D561CFF51C0AAC6B7F41B" blockId="20.[151,1437,289,1036]" pageId="20" pageNumber="21">
<emphasis id="B91F2326F60D561CFF51C0AAC6FAF388" bold="true" box="[199,679,901,928]" pageId="20" pageNumber="21">Temporal and geographic distribution.</emphasis>
Late Pliocene of southeastern Queensland, central eastern New South Wales, northern South Australia, and
<collectingCountry id="F37CBFA4F60D561CFD12C086C73CF3D3" box="[644,865,938,964]" name="Papua New Guinea" pageId="20" pageNumber="21">Papua New Guinea</collectingCountry>
. Early and/or middle Pleistocene of central eastern New South Wales and southern South Australia. Middle Pleistocene of eastern Queensland, southeastern Western Australia and northern South Australia. See
<figureCitation id="1350E3B1F60D561CFD13C0DEC6B9F41B" box="[645,740,1010,1036]" captionStart="FIGURE 1" captionStartId="5.[151,250,1691,1715]" captionTargetBox="[207,1381,795,1665]" captionTargetId="figure-278@5.[207,1381,795,1665]" captionTargetPageId="5" captionText="FIGURE 1. Map of Australia and New Guinea showing localities that have produced fossils attributable, or potentially attributable, to species of Bohra." figureDoi="http://doi.org/10.5281/zenodo.8012668" httpUri="https://zenodo.org/record/8012668/files/figure.png" pageId="20" pageNumber="21">Figure 1</figureCitation>
.
</paragraph>
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