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<document id="59AF5E3588E1B19AB0CF377D642DFB51" ID-DOI="10.15553/c2022v771a7" ID-ISSN="2235-3658" ID-Zenodo-Dep="7183082" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665435833373" checkinUser="felipe" docAuthor="Stone, Robert Douglas" docDate="2022" docId="03BD87EFFFAEFFEBFFEE555366C0FE7C" docLanguage="en" docName="Candollea.77.1.81-103.pdf" docOrigin="Candollea 77 (1)" docSource="http://dx.doi.org/10.15553/c2022v771a7" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Memecylon conjugens R. D. Stone 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="86" masterDocId="FF84FF97FFADFFEEFF9F53126367FFEF" masterDocTitle="Ten new species of Memecylon (Melastomataceae) from Madagascar" masterLastPageNumber="103" masterPageNumber="81" pageNumber="84" updateTime="1699384657421" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="01ABCDC48EFEEF8685A424F5091196FF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4FE5EC82D11453C5F2933C5AE0487FB3">Ten new species of Memecylon (Melastomataceae) from Madagascar</mods:title>
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<mods:namePart id="95FE43946FA429E3753AD537B3F3E90C">Stone, Robert Douglas</mods:namePart>
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<mods:title id="9A6BDBCF6A6FF0E42F9F97CD4FF5EA3F">Candollea</mods:title>
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<mods:part id="341055D38D79819CCD2CB0F91A43FB5D">
<mods:date id="4579EB6591DD4EE78C835B44383041A7">2022</mods:date>
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<mods:number id="44200293E2EDFA9E046C85385A3B47D4">2022-06-01</mods:number>
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<mods:identifier id="0190EEA75645E301F58DFCA133C90590" type="DOI">10.15553/c2022v771a7</mods:identifier>
<mods:identifier id="8779F66A28F13E080EF8738733BABF1F" type="ISSN">2235-3658</mods:identifier>
<mods:identifier id="DCD34B24ABA83FD015799C72FDC1DA7C" type="Zenodo-Dep">7183082</mods:identifier>
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<treatment id="03BD87EFFFAEFFEBFFEE555366C0FE7C" LSID="urn:lsid:plazi:treatment:03BD87EFFFAEFFEBFFEE555366C0FE7C" httpUri="http://treatment.plazi.org/id/03BD87EFFFAEFFEBFFEE555366C0FE7C" lastPageId="5" lastPageNumber="86" pageId="3" pageNumber="84">
<subSubSection id="C30E6572FFAEFFEDFFEE55536229F8EB" pageId="3" pageNumber="84" type="nomenclature">
<paragraph id="8BAB36F9FFAEFFEDFFEE555361F7F9B5" blockId="3.[113,656,1601,1626]" box="[113,656,1601,1626]" pageId="3" pageNumber="84">
<heading id="D0E38195FFAEFFEDFFEE555361F7F9B5" box="[113,656,1601,1626]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="84" reason="2">
<taxonomicName id="4C144D7AFFAEFFEDFFEE555362B2F9B5" ID-CoL="9WGRK" authority="R.D. Stone" authorityName="R. D. Stone" authorityYear="2022" box="[113,469,1601,1626]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="84" phylum="Tracheophyta" rank="species" species="conjugens" status="sp. nov.">
<emphasis id="B960EAEBFFAEFFEDFFEE55536233F9B6" bold="true" box="[113,340,1601,1625]" italics="true" pageId="3" pageNumber="84">Memecylon conjugens</emphasis>
R.D. Stone
</taxonomicName>
,
<emphasis id="B960EAEBFFAEFFEDFE4055506154F9B5" bold="true" box="[479,563,1602,1626]" pageId="3" pageNumber="84">
<taxonomicNameLabel id="A2535790FFAEFFEDFE4055506154F9B5" box="[479,563,1602,1626]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132F2A7CFFAEFFEDFDDE555061E4F9B5" box="[577,643,1602,1626]" captionStart="Fig" captionStartId="4.[136,166,1915,1935]" captionTargetBox="[264,1332,317,1821]" captionTargetId="figure-17@4.[268,1367,299,1823]" captionTargetPageId="4" captionText="Fig. 2. – Memecylon conjugens R.D. Stone. A. Flowering branch; B–D. Leaves; E. Inflorescence; F. Floral bud; G. Petal; H. Stamen; I–K. Fruit." figureDoi="http://doi.org/10.5281/zenodo.7183084" httpUri="https://zenodo.org/record/7183084/files/figure.png" pageId="3" pageNumber="84">Fig. 2</figureCitation>
).
</heading>
</paragraph>
<paragraph id="8BAB36F9FFAEFFEDFF08557E6229F8EB" blockId="3.[151,763,1644,1796]" pageId="3" pageNumber="84">
<materialsCitation id="3B7C3CA4FFAEFFEDFF08557E62D9F92B" collectionCode="H, R, S, N" country="Madagascar" elevation="1350" latitude="-14.482499" location="Marojejy" longLatPrecision="21" longitude="49.63667" municipality="Andapa" pageId="3" pageNumber="84" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="B960EAEBFFAEFFEDFF08557E6250F94B" bold="true" pageId="3" pageNumber="84">
<collectionCode id="ED05AE3CFFAEFFEDFF08557E63C9F96B" box="[151,174,1644,1668]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="3" pageNumber="84" type="Herbarium">H</collectionCode>
o l o t y p u s:
<collectingCountry id="F3037669FFAEFFEDFEA3557E6288F96B" box="[316,495,1644,1668]" name="Madagascar" pageId="3" pageNumber="84">MADAGASCAR</collectingCountry>
.
<collectionCode id="ED05AE3CFFAEFFEDFD98557E617CF96B" box="[519,539,1644,1668]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="84">R</collectionCode>
e g.
<collectionCode id="ED05AE3CFFAEFFEDFDCE557E6107F96B" box="[593,608,1644,1668]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="3" pageNumber="84" type="Herbarium">S</collectionCode>
AVA [Pr o v.
<collectingRegion id="49D0F81BFFAEFFEDFF08559E624CF94B" box="[151,299,1676,1700]" country="Madagascar" name="Antsiranana" pageId="3" pageNumber="84">Antsiranana</collectingRegion>
]:
</emphasis>
<collectionCode id="ED05AE3CFFAEFFEDFEA0559F6233F94B" box="[319,340,1677,1700]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="84" type="Herbarium">N</collectionCode>
of
<collectingMunicipality id="6BCFAC83FFAEFFEDFEE4559E62B4F94B" box="[379,467,1676,1700]" pageId="3" pageNumber="84">Andapa</collectingMunicipality>
, RNI
<location id="8ECB6022FFAEFFEDFD84559E6119F94B" LSID="urn:lsid:plazi:treatment:03BD87EFFFAEFFEBFFEE555366C0FE7C:8ECB6022FFAEFFEDFD84559E6119F94B" box="[539,638,1676,1700]" country="Madagascar" latitude="-14.482499" longLatPrecision="21" longitude="49.63667" municipality="Andapa" name="Marojejy" pageId="3" pageNumber="84" stateProvince="Antsiranana">Marojejy</location>
,
<geoCoordinate id="EE20503EFFAEFFEDFD16559E619CF94B" box="[649,763,1676,1700]" degrees="14" direction="south" minutes="28" orientation="latitude" pageId="3" pageNumber="84" precision="15" seconds="57" value="-14.482499">14°28'57&quot;S</geoCoordinate>
<geoCoordinate id="EE20503EFFAEFFEDFF0855BE6274F92B" box="[151,275,1708,1732]" degrees="49" direction="east" minutes="38" orientation="longitude" pageId="3" pageNumber="84" precision="15" seconds="12" value="49.63667">49°38'12&quot;E</geoCoordinate>
,
<quantity id="4CEC9B1CFFAEFFEDFEBE55BE62D9F92B" box="[289,446,1707,1732]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="3" pageNumber="84" unit="m" value="1350.0" valueMax="1600.0" valueMin="1100.0">
<elevation id="0039D1CAFFAEFFEDFEBE55BE62D9F92B" box="[289,446,1707,1732]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="3" pageNumber="84" unit="m" value="1350.0" valueMax="1600.0" valueMin="1100.0">1100– 1600 m</elevation>
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,
<date id="FFAA1039FFAEFFEDFE5355BE6173F92B" box="[460,532,1708,1732]" pageId="3" pageNumber="84" value="1994-01">I.1994</date>
,
<materialsCitation id="3B7C3CA4FFAEFFEDFDBE55BE622DF8EB" collectionCode="K, NU, P" country="Madagascar" pageId="3" pageNumber="84" specimenCode="P00516010" specimenCount="1">
fl. buds,
<emphasis id="B960EAEBFFAEFFEDFD1A55BE6261F90B" italics="true" pageId="3" pageNumber="84">Malcomber et al. 2687</emphasis>
(CAS-926817!; iso-:
<collectionCode id="ED05AE3CFFAEFFEDFE6D55DE616EF90B" box="[498,521,1740,1764]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="84" type="Herbarium">K</collectionCode>
!, MO-6262414!,
<collectionCode id="ED05AE3CFFAEFFEDFD5A55DE6194F90B" box="[709,755,1740,1764]" country="Thailand" name="Department of Microbiology, Faculty of Science" pageId="3" pageNumber="84">NU</collectionCode>
!,
<collectionCode id="ED05AE3CFFAEFFEDFF0855FF63C0F8EB" box="[151,167,1773,1796]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="84" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBB29E82FFAEFFEDFF2B55FE625EF8EB" box="[180,313,1772,1796]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="84" type="Herbarium">P00516010</specimenCode>
]!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6572FFAEFFEBFF08540566C0FE7C" lastPageId="5" lastPageNumber="86" pageId="3" pageNumber="84" type="description">
<paragraph id="8BAB36F9FFAEFFEDFF085405660EFDC0" blockId="3.[151,764,1815,1935]" lastBlockId="3.[839,1451,279,559]" pageId="3" pageNumber="84">
<emphasis id="B960EAEBFFAEFFEDFF085405625DF860" italics="true" pageId="3" pageNumber="84">
Ob folia caudato-acuminata Memecyloni longicuspi Baker et
<taxonomicName id="4C144D7AFFAEFFEDFF2E542A62BCF8A0" authority="H. Perrier" authorityName="H. Perrier" box="[177,475,1848,1871]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="84" phylum="Tracheophyta" rank="species" species="peracuminato">M. peracuminato H. Perrier</taxonomicName>
simile, sed a primo ramulis teretibus (non quadrangularibus et anguste alatis), hypanthocalyce majore 3
</emphasis>
×
<emphasis id="B960EAEBFFAEFFEDFEC654656149F860" box="[345,558,1910,1935]" italics="true" pageId="3" pageNumber="84">
<quantity id="4CEC9B1CFFAEFFEDFEC6546562ACF860" box="[345,459,1910,1935]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="3" pageNumber="84" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5– 4 mm</quantity>
(non 1.5
</emphasis>
×
<emphasis id="B960EAEBFFAEFFEDFDD2546560E1FDC0" italics="true" pageId="3" pageNumber="84">
<quantity id="4CEC9B1CFFAEFFEDFDD2546561C3F861" box="[589,676,1911,1935]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="84" unit="mm" value="2.5">2.5 mm</quantity>
), calycis margine sinuato-dentato (non truncato 4-denticulatoque), fructus corona calycina circa
<quantity id="4CEC9B1CFFAEFFEDFBC1522567F2FEA0" box="[1118,1173,311,335]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="84" unit="mm" value="1.0">1 mm</quantity>
longa (non subprominente); a secundo petiolis brevioribus
<quantity id="4CEC9B1CFFAEFFEDFB065245679DFE80" box="[1177,1274,342,367]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="84" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2 – 3 mm</quantity>
(non
<quantity id="4CEC9B1CFFAEFFEDFADE524566C5FE81" box="[1345,1442,342,367]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="84" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3 – 5 mm</quantity>
) longis, cymularum bracteis deciduis (non persistentibus), floribus pedicellatis (non subsessilibus) pedicellis plerumque
<quantity id="4CEC9B1CFFAEFFEDFCD852A560D6FE20" box="[839,945,438,463]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="3" pageNumber="84" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
longis et corolla in alabastro conico-apiculata (non conico-obtusa); ab ambobus cymulis longioribus 2(– 3) cm, pedunculis
<quantity id="4CEC9B1CFFAEFFEDFC5F52E56741FDE0" box="[960,1062,502,527]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="3" pageNumber="84" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">6 – 11 mm</quantity>
longis et fructibus multo majoribus 13–19
</emphasis>
×
<emphasis id="B960EAEBFFAEFFEDFC005105660EFDC0" box="[927,1385,534,559]" italics="true" pageId="3" pageNumber="84">
<quantity id="4CEC9B1CFFAEFFEDFC005105676FFDC0" box="[927,1032,534,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="3" pageNumber="84" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 mm</quantity>
ellipsoideis vel late ovoideis differt.
</emphasis>
</paragraph>
<paragraph id="8BAB36F9FFAEFFEDFCD8514566F4FA41" blockId="3.[801,1452,598,1455]" pageId="3" pageNumber="84">
<emphasis id="B960EAEBFFAEFFEDFCD8514560F4FD80" box="[839,915,599,623]" italics="true" pageId="3" pageNumber="84">Shrubs</emphasis>
or small
<emphasis id="B960EAEBFFAEFFEDFB92514A6727FD80" box="[1037,1088,600,623]" italics="true" pageId="3" pageNumber="84">trees</emphasis>
<quantity id="4CEC9B1CFFAEFFEDFBD4514567FCFD81" box="[1099,1179,598,623]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="3" pageNumber="84" unit="m" value="6.0" valueMax="8.0" valueMin="4.0">4 – 8 m</quantity>
high, evergreen; ultimate branchlets slender, the youngest compressed, soon becoming terete with age; nodes ± thickened; internodes (1.4 –)2 – 4(– 5.3) cm long.
<emphasis id="B960EAEBFFAEFFEDFBD851AA67F7FD20" box="[1095,1168,696,719]" italics="true" pageId="3" pageNumber="84">Leaves</emphasis>
subcoriaceous, petiolate, dark green and somewhat glossy above, paler and dull below, minutely roughened when dry; petioles
<quantity id="4CEC9B1CFFAEFFEDFB5351E56641FCE1" box="[1228,1318,758,783]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="84" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long; blades elliptic to narrowly elliptic or elliptic-ovate, (3–)4 –5.5(– 7) × (1.1–)1.8–2.4(–2.7) cm, base cuneate, apex caudate-acuminate, acumen (4 –)7 – 12(– 18) mm long, obtuse; midnerve finely canaliculate adaxially, conspicuous abaxially but not prominent or very slightly so towards the base of the blade; intramarginal nerves and transverse veins invisible.
<emphasis id="B960EAEBFFAEFFEDFAD350A566E9FC21" box="[1356,1422,951,974]" italics="true" pageId="3" pageNumber="84">Cymes</emphasis>
to c. 2(–3) cm long, 3–5(–6)-flowered, solitary or geminate in the leaf axils, sometimes terminally on the branchlets, and frequently also at the recently defoliated nodes; peduncles slender, compressed, (4 –)6 – 11(– 14) mm long; axis often extended by a slender internode (1 –)
<quantity id="4CEC9B1CFFAEFFEDFB5F57456647FB81" box="[1216,1312,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="3" pageNumber="84" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3– 6 mm</quantity>
long; bracts rapidly deciduous, not seen.
<emphasis id="B960EAEBFFAEFFEDFBC6576567CBFB60" box="[1113,1196,1143,1167]" italics="true" pageId="3" pageNumber="84">Flowers</emphasis>
white (at least in bud), on slender pedicels (1.5 –)2–3.5(–4.5) mm long; hypanthocalyx obconic to campanulate, c. 3 ×
<quantity id="4CEC9B1CFFAEFFEDFB4B57A5662BFB21" box="[1236,1356,1206,1231]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="3" pageNumber="84" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5 – 4 mm</quantity>
, margin shallowly sinuate-dentate, lobes broadly rounded, scarious margined; corolla in bud conical-apiculate, c.
<quantity id="4CEC9B1CFFAEFFEDFAB557E5660CFAE1" box="[1322,1387,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="84" unit="mm" value="2.0">2 mm</quantity>
long; open flowers not seen; anthers in bud c.
<quantity id="4CEC9B1CFFAEFFEDFB7F56056647FAC1" box="[1248,1312,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="84" unit="mm" value="2.0">2 mm</quantity>
long, thecae fronto-ventral, dorsal oil-gland narrowly elliptic,
<quantity id="4CEC9B1CFFAEFFEDFACA562566CCFAA1" box="[1365,1451,1335,1359]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="84" unit="mm" value="0.7">0.7 mm</quantity>
long, posterior extremity of connective acute.
<emphasis id="B960EAEBFFAEFFEDFA94564A662CFA80" box="[1291,1355,1368,1391]" italics="true" pageId="3" pageNumber="84">Fruits</emphasis>
purplish at maturity, broadly ovoid to ellipsoid, 13– 19 ×
<quantity id="4CEC9B1CFFAEFFEDFAAB566566C1FA61" box="[1332,1446,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="3" pageNumber="84" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11– 15 mm</quantity>
; persistent calycinal crown c.
<quantity id="4CEC9B1CFFAEFFEDFBC5568567F0FA41" box="[1114,1175,1431,1455]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="84" unit="mm" value="1.0">1 mm</quantity>
high, margin truncate.
</paragraph>
<paragraph id="8BAB36F9FFAEFFEDFCD856C56609F9A0" blockId="3.[801,1452,1495,1615]" pageId="3" pageNumber="84">
<emphasis id="B960EAEBFFAEFFEDFCD856C560D4FA00" box="[839,947,1495,1519]" italics="true" pageId="3" pageNumber="84">Etymology</emphasis>
. – The epithet
<taxonomicName id="4C144D7AFFAEFFEDFBCF56CA67C8FA00" box="[1104,1199,1496,1519]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="84" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFAEFFEDFBCF56CA67C8FA00" box="[1104,1199,1496,1519]" italics="true" pageId="3" pageNumber="84">conjugens</emphasis>
</taxonomicName>
is an adjective meaning “joined together”, in reference to the fact that the flowering material from Marojejy and fruiting material from Anjanaharibe-Sud were at first considered to be separate species.
</paragraph>
<paragraph id="8BAB36F9FFAEFFEDFCD855656792F901" blockId="3.[801,1452,1655,1775]" pageId="3" pageNumber="84">
<emphasis id="B960EAEBFFAEFFEDFCD855656731F960" box="[839,1110,1655,1679]" italics="true" pageId="3" pageNumber="84">Distribution and ecology</emphasis>
. – Northeastern
<collectingCountry id="F3037669FFAEFFEDFABD556566CCF960" box="[1314,1451,1655,1679]" name="Madagascar" pageId="3" pageNumber="84">Madagascar</collectingCountry>
(SAVA region), known only from the Marojejy and Anjanaharibe massifs near the town of Andapa. Habitat in montane, humid forests at elevations of
<quantity id="4CEC9B1CFFAEFFEDFBFE55C56796F901" box="[1121,1265,1750,1775]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.35" metricValueMin="1.1" pageId="3" pageNumber="84" unit="m" value="1225.0" valueMax="1350.0" valueMin="1100.0">1100–1350 m</quantity>
.
</paragraph>
<paragraph id="8BAB36F9FFAEFFEDFCD8540566CCF860" blockId="3.[801,1451,1815,1935]" pageId="3" pageNumber="84">
<emphasis id="B960EAEBFFAEFFEDFCD854056768F8C0" box="[839,1039,1815,1839]" italics="true" pageId="3" pageNumber="84">Conservation status</emphasis>
. –
<taxonomicName id="4C144D7AFFAEFFEDFBAF54056660F8C0" box="[1072,1287,1815,1839]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="84" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFAEFFEDFBAF54056660F8C0" box="[1072,1287,1815,1839]" italics="true" pageId="3" pageNumber="84">Memecylon conjugens</emphasis>
</taxonomicName>
is known from three locations with an estimated EOO of
<quantity id="4CEC9B1CFFAEFFEDFA9754256629F8A0" box="[1288,1358,1847,1871]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="84" unit="km" value="50.0">50 km</quantity>
² and an AOO of
<quantity id="4CEC9B1CFFAEFFEDFC1B544560A1F880" box="[900,966,1879,1903]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="84" unit="km" value="12.0">12 km</quantity>
². All of the known locations are in protected areas including the Marojejy National Park (60,050 ha) and
</paragraph>
<caption id="DF6B6671FFA9FFEAFF175469661BF860" ID-DOI="http://doi.org/10.5281/zenodo.7183084" ID-Zenodo-Dep="7183084" box="[136,1404,1915,1935]" httpUri="https://zenodo.org/record/7183084/files/figure.png" pageId="4" pageNumber="85" startId="4.[136,166,1915,1935]" targetBox="[264,1332,317,1821]" targetPageId="4">
<paragraph id="8BAB36F9FFA9FFEAFF175469661BF860" blockId="4.[136,1462,1915,1960]" box="[136,1404,1915,1935]" pageId="4" pageNumber="85">
<emphasis id="B960EAEBFFA9FFEAFF17546963B7F860" bold="true" box="[136,208,1915,1935]" pageId="4" pageNumber="85">Fig. 2. –</emphasis>
<taxonomicName id="4C144D7AFFA9FFEAFF4A54696119F860" authority="R. D. Stone. A. Flowering" box="[213,638,1915,1935]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA9FFEAFF4A546962F9F860" box="[213,414,1915,1935]" italics="true" pageId="4" pageNumber="85">Memecylon conjugens</emphasis>
R.D. Stone.
<emphasis id="B960EAEBFFA9FFEAFD9354696178F860" bold="true" box="[524,543,1915,1935]" pageId="4" pageNumber="85">A.</emphasis>
Flowering
</taxonomicName>
branch;
<emphasis id="B960EAEBFFA9FFEAFD525469619CF860" bold="true" box="[717,763,1915,1935]" pageId="4" pageNumber="85">B–D.</emphasis>
Leaves;
<emphasis id="B960EAEBFFA9FFEAFCD65469603CF860" bold="true" box="[841,859,1915,1935]" pageId="4" pageNumber="85">E.</emphasis>
Inflorescence;
<emphasis id="B960EAEBFFA9FFEAFC775469609FF860" bold="true" box="[1000,1016,1915,1935]" pageId="4" pageNumber="85">F.</emphasis>
Floral bud;
<emphasis id="B960EAEBFFA9FFEAFBFD54696710F860" bold="true" box="[1122,1143,1915,1935]" pageId="4" pageNumber="85">G.</emphasis>
Petal;
<emphasis id="B960EAEBFFA9FFEAFB2B546967AFF860" bold="true" box="[1204,1224,1915,1935]" pageId="4" pageNumber="85">H.</emphasis>
Stamen;
<emphasis id="B960EAEBFFA9FFEAFA8154696622F860" bold="true" box="[1310,1349,1915,1935]" pageId="4" pageNumber="85">I–K.</emphasis>
Fruit.
</paragraph>
</caption>
<paragraph id="8BAB36F9FFA9FFEAFF17548666D1F847" blockId="4.[136,1462,1915,1960]" box="[136,1462,1938,1960]" pageId="4" pageNumber="85">
[
<emphasis id="B960EAEBFFA9FFEAFF115486638DF847" bold="true" box="[142,234,1940,1960]" pageId="4" pageNumber="85">A, B, E–H:</emphasis>
<emphasis id="B960EAEBFFA9FFEAFF70548062D1F849" box="[239,438,1938,1958]" italics="true" pageId="4" pageNumber="85">Malcomber et al. 2687</emphasis>
, CAS,NU,P;
<emphasis id="B960EAEBFFA9FFEAFDB65486611FF847" bold="true" box="[553,632,1940,1960]" pageId="4" pageNumber="85">C, D, I, J:</emphasis>
<emphasis id="B960EAEBFFA9FFEAFDE354806021F849" box="[636,838,1938,1958]" italics="true" pageId="4" pageNumber="85">Ravelonarivo et al. 319</emphasis>
, MO,P;
<emphasis id="B960EAEBFFA9FFEAFC12548660C7F847" bold="true" box="[909,928,1940,1960]" pageId="4" pageNumber="85">K:</emphasis>
<emphasis id="B960EAEBFFA9FFEAFC3A548067A5F849" box="[933,1218,1938,1958]" italics="true" pageId="4" pageNumber="85">Ravelonarivo &amp; Rabesonina 486</emphasis>
, NU] [Drawing: S. Burrows]
</paragraph>
<paragraph id="8BAB36F9FFA8FFEBFFEE52056277FCA0" blockId="5.[113,764,279,847]" pageId="5" pageNumber="86">
the Réserve Spéciale d’Anjanaharibe-Sud (26,903 ha), both of which are managed by
<collectingCountry id="F3037669FFA8FFEBFEC1522562BCFEA0" box="[350,475,311,335]" name="Madagascar" pageId="5" pageNumber="86">Madagascar</collectingCountry>
National Parks (
<bibRefCitation id="EF854B08FFA8FFEBFD1652256385FE80" author="GOODMAN, S. M. &amp; M. J. RAHERILALAO &amp; S. WOHLHAUSER" pageId="5" pageNumber="86" refId="ref14816" refString="GOODMAN, S. M., M. J. RAHERILALAO &amp; S. WOHLHAUSER (ed.) (2021). The terrestrial protected areas of Madagascar: their history, description, and biota. Vol. 2. Association Vahatra, Antananarivo." type="book" year="2021">GOODMAN et al., 2021</bibRefCitation>
). Marojejy has seen a relatively minor loss of 846 ha (1.7 %) of moist evergreen forest between the years 1996 and 2016, and at Anjanaharibe-Sud the deforestation rate has been similarly low, 404 ha (1.6 %) over the same time period (
<bibRefCitation id="EF854B08FFA8FFEBFFE452C56213FE00" author="GOODMAN, S. M. &amp; M. J. RAHERILALAO &amp; S. WOHLHAUSER" box="[123,372,471,495]" pageId="5" pageNumber="86" refId="ref14816" refString="GOODMAN, S. M., M. J. RAHERILALAO &amp; S. WOHLHAUSER (ed.) (2021). The terrestrial protected areas of Madagascar: their history, description, and biota. Vol. 2. Association Vahatra, Antananarivo." type="book" year="2021">GOODMAN et al., 2021</bibRefCitation>
). Ongoing anthropogenic pressures include exploitation of hardwoods, collection of non-woody forest products, uncontrolled fires (Marojejy), slash-and-burn agriculture (Anjanaharibe-Sud), and small-scale mineral extraction (Anjanaharibe-Sud) (
<bibRefCitation id="EF854B08FFA8FFEBFE5D514561CBFD80" author="GOODMAN, S. M. &amp; M. J. RAHERILALAO &amp; S. WOHLHAUSER" box="[450,684,599,623]" pageId="5" pageNumber="86" refId="ref14816" refString="GOODMAN, S. M., M. J. RAHERILALAO &amp; S. WOHLHAUSER (ed.) (2021). The terrestrial protected areas of Madagascar: their history, description, and biota. Vol. 2. Association Vahatra, Antananarivo." type="book" year="2021">GOODMAN et al., 2021</bibRefCitation>
). Based on its limited AOO and the apparent threats,
<taxonomicName id="4C144D7AFFA8FFEBFDED516A619CFD60" box="[626,763,632,655]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA8FFEBFDED516A619CFD60" box="[626,763,632,655]" italics="true" pageId="5" pageNumber="86">M. conjugens</emphasis>
</taxonomicName>
would meet the criterion B for listing as “Endangered” in accordance with the IUCN Red List Categories and Criteria (
<bibRefCitation id="EF854B08FFA8FFEBFF0051C5624AFD00" author="IUCN" box="[159,301,727,751]" pageId="5" pageNumber="86" refId="ref14864" refString="IUCN (2012). IUCN Red List Categories and Criteria. Version 3.1. Ed. 2. IUCN Species Survival Commission, Gland and Cambridge." type="book" year="2012">IUCN, 2012</bibRefCitation>
), but it would be better assessed as “Near Threatened” [NT] contingent upon the continued effectiveness of habitat-specific conservation and management measures (
<bibRefCitation id="EF854B08FFA8FFEBFFE650256263FCA0" author="IUCN" box="[121,260,823,847]" pageId="5" pageNumber="86" refId="ref14892" refString="IUCN (2019). Guidelines for using the IUCN Red List Categories and Criteria. Version 14. IUCN Standards and Petitions Committee. [https: // www. iucnredlist. org / resources / redlistguidelines]" type="url" year="2019">IUCN, 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BAB36F9FFA8FFEBFF08506A6169FB80" blockId="5.[113,765,887,1935]" pageId="5" pageNumber="86">
<emphasis id="B960EAEBFFA8FFEBFF08506A63B7FC60" box="[151,208,888,911]" italics="true" pageId="5" pageNumber="86">Notes</emphasis>
. – The presence of an anther-gland and the shallowly sinuate-dentate calyx margin of
<taxonomicName id="4C144D7AFFA8FFEBFD825085619CFC40" box="[541,763,919,943]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA8FFEBFD825085619CFC40" box="[541,763,919,943]" italics="true" pageId="5" pageNumber="86">Memecylon conjugens</emphasis>
</taxonomicName>
indicate that it would be placed in
<taxonomicName id="4C144D7AFFA8FFEBFE4050A562FAFC00" authority="sect. Memecylon sensu JACQUES-FELIX (1985" authorityName="JACQUES-FELIX" authorityYear="1985" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="section" section="Memecylon">
<emphasis id="B960EAEBFFA8FFEBFE4050A56137FC20" box="[479,592,951,975]" italics="true" pageId="5" pageNumber="86">Memecylon</emphasis>
sect.
<emphasis id="B960EAEBFFA8FFEBFD1550A5619CFC20" box="[650,763,951,975]" italics="true" pageId="5" pageNumber="86">Memecylon</emphasis>
sensu
<bibRefCitation id="EF854B08FFA8FFEBFF2B50C362FAFC00" author="JACQUES-FELIX, H." box="[180,413,977,1007]" pageId="5" pageNumber="86" pagination="3 - 58" refId="ref14998" refString="JACQUES-FELIX, H. (1985 b). Les Memecyleae (Melastomataceae) de Madagascar (2 E partie). Bull. Mus. Natl. Hist. Nat., B, Adansonia 7: 3 - 58." type="journal article" year="1985">JACQUES-FÉLIX (1985</bibRefCitation>
</taxonomicName>
a,
<bibRefCitation id="EF854B08FFA8FFEBFE2B50C56292FC00" author="JACQUES-FELIX, H." box="[436,501,983,1007]" pageId="5" pageNumber="86" pagination="3 - 58" refId="ref14998" refString="JACQUES-FELIX, H. (1985 b). Les Memecyleae (Melastomataceae) de Madagascar (2 E partie). Bull. Mus. Natl. Hist. Nat., B, Adansonia 7: 3 - 58." type="journal article" year="1985">1985b</bibRefCitation>
). However, the limits of this section are in need of revision since molecular results suggest the group is not monophyletic (
<bibRefCitation id="EF854B08FFA8FFEBFD86570561C6FBC0" author="STONE, R. D." box="[537,673,1047,1071]" pageId="5" pageNumber="86" pagination="539 - 561" refId="ref15377" refString="STONE, R. D. (2014). The species-rich, paleotropical genus Memecylon (Melastomataceae): molecular phylogenetics and revised infrageneric classification of the African species. Taxon 63: 539 - 561." type="journal article" year="2014">STONE, 2014</bibRefCitation>
, unpubl. data; AMARASINGHE et al., 2021), and the
<typeStatus id="54AF885BFFA8FFEBFDF1572561C7FBA0" box="[622,672,1079,1103]" pageId="5" pageNumber="86">type</typeStatus>
species (
<taxonomicName id="4C144D7AFFA8FFEBFFE5574A627BFB80" box="[122,284,1111,1135]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="capitellatum">
<emphasis id="B960EAEBFFA8FFEBFFE5574A627BFB80" box="[122,284,1111,1135]" italics="true" pageId="5" pageNumber="86">M. capitellatum</emphasis>
</taxonomicName>
L.) is from
<collectingCountry id="F3037669FFA8FFEBFE025745616CFB80" box="[413,523,1111,1135]" name="Sri Lanka" pageId="5" pageNumber="86">Sri Lanka</collectingCountry>
.
</paragraph>
<paragraph id="8BAB36F9FFA8FFEBFF08576562C2FA80" blockId="5.[113,765,887,1935]" pageId="5" pageNumber="86">
The leaves of the new species resemble those of
<taxonomicName id="4C144D7AFFA8FFEBFD145765627BFB40" authority="Baker" authorityName="Baker" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="longicuspe">
<emphasis id="B960EAEBFFA8FFEBFD14576563B1FB40" italics="true" pageId="5" pageNumber="86">Memecylon longicuspe</emphasis>
Baker
</taxonomicName>
and
<taxonomicName id="4C144D7AFFA8FFEBFECF578A61E0FB40" authority="H. Perrier" authorityName="H. Perrier" box="[336,647,1175,1199]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="peracuminatum">
<emphasis id="B960EAEBFFA8FFEBFECF578A6173FB40" box="[336,532,1176,1199]" italics="true" pageId="5" pageNumber="86">M. peracuminatum</emphasis>
H. Perrier
</taxonomicName>
in appearing 1-nerved with apex caudate-acuminate and acumen narrowly obtuse at the summit. However, this may not be an indication of its true affinity, and both
<taxonomicName id="4C144D7AFFA8FFEBFDA257EA61ACFAE0" box="[573,715,1271,1295]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="longicuspe">
<emphasis id="B960EAEBFFA8FFEBFDA257EA61ACFAE0" box="[573,715,1271,1295]" italics="true" pageId="5" pageNumber="86">M. longicuspe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C144D7AFFA8FFEBFFEE560A6258FAC0" box="[113,319,1304,1327]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="peracuminatum">
<emphasis id="B960EAEBFFA8FFEBFFEE560A6258FAC0" box="[113,319,1304,1327]" italics="true" pageId="5" pageNumber="86">M. peracuminatum</emphasis>
</taxonomicName>
are from east-central and southeastern
<collectingCountry id="F3037669FFA8FFEBFFEE56256392FAA0" box="[113,245,1335,1359]" name="Madagascar" pageId="5" pageNumber="86">Madagascar</collectingCountry>
, while
<taxonomicName id="4C144D7AFFA8FFEBFEDA562A62A9FAA0" box="[325,462,1336,1359]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA8FFEBFEDA562A62A9FAA0" box="[325,462,1336,1359]" italics="true" pageId="5" pageNumber="86">M. conjugens</emphasis>
</taxonomicName>
has a different and limited distribution in the northeast.
</paragraph>
<paragraph id="8BAB36F9FFA8FFEBFF08566561C9F860" blockId="5.[113,765,887,1935]" pageId="5" pageNumber="86">
The fruits of
<taxonomicName id="4C144D7AFFA8FFEBFEBC5665629CFA60" box="[291,507,1399,1423]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA8FFEBFEBC5665629CFA60" box="[291,507,1399,1423]" italics="true" pageId="5" pageNumber="86">Memecylon conjugens</emphasis>
</taxonomicName>
are variable, an unusual feature since other species of Malagasy
<taxonomicName id="4C144D7AFFA8FFEBFDAA568561CCFA40" box="[565,683,1431,1455]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="genus">
<emphasis id="B960EAEBFFA8FFEBFDAA568561CCFA40" box="[565,683,1431,1455]" italics="true" pageId="5" pageNumber="86">Memecylon</emphasis>
</taxonomicName>
have a more-or-less consistent (often diagnostic) fruit size and shape. Among the additional specimens (
<typeStatus id="54AF885BFFA8FFEBFE7F56C5612FFA00" box="[480,584,1495,1519]" pageId="5" pageNumber="86" type="paratype">paratypes</typeStatus>
) cited below, the fruits of
<emphasis id="B960EAEBFFA8FFEBFF5056E562A7F9E0" box="[207,448,1527,1551]" italics="true" pageId="5" pageNumber="86">Ravelonarivo et al. 319</emphasis>
are broadly ovoid (conspicuously wider below the middle), as are those of the NU and P sheets of
<emphasis id="B960EAEBFFA8FFEBFF4B55256142F9A0" box="[212,549,1591,1615]" italics="true" pageId="5" pageNumber="86">Ravelonarivo &amp; Rabesonina 486</emphasis>
. However, the fruits of the CAS and MO sheets of
<emphasis id="B960EAEBFFA8FFEBFE2D5545619CF980" box="[434,763,1623,1647]" italics="true" pageId="5" pageNumber="86">Ravelonarivo &amp; Rabesonina 486</emphasis>
are differently shaped, i.e. broadly ellipsoid or even obovoid (widest above the middle). The ovoid fruits of
<taxonomicName id="4C144D7AFFA8FFEBFDED558A619CF940" box="[626,763,1688,1711]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="conjugens">
<emphasis id="B960EAEBFFA8FFEBFDED558A619CF940" box="[626,763,1688,1711]" italics="true" pageId="5" pageNumber="86">M. conjugens</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C144D7AFFA8FFEBFEC255AA6129F920" authority="Jacq." authorityName="Jacq." box="[349,590,1719,1743]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="pedunculatum">
<emphasis id="B960EAEBFFA8FFEBFEC255AA6177F920" box="[349,528,1719,1743]" italics="true" pageId="5" pageNumber="86">M. pedunculatum</emphasis>
Jacq.
</taxonomicName>
-Fél., except that they are larger (11–15 vs.
<quantity id="4CEC9B1CFFA8FFEBFE1855C562A0F901" box="[391,455,1751,1775]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="86" unit="mm" value="8.0">8 mm</quantity>
in diam.), and the calycinal crown is prominent (vs. appressed to the summit of the ovary). Both of these species occur in the same region of northeastern
<collectingCountry id="F3037669FFA8FFEBFFEE54256396F8A0" box="[113,241,1847,1871]" name="Madagascar" pageId="5" pageNumber="86">Madagascar</collectingCountry>
but are not closely related since
<taxonomicName id="4C144D7AFFA8FFEBFDD6542A619CF8A0" box="[585,763,1847,1871]" class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="species" species="pedunculatum">
<emphasis id="B960EAEBFFA8FFEBFDD6542A619CF8A0" box="[585,763,1847,1871]" italics="true" pageId="5" pageNumber="86">M. pedunculatum</emphasis>
</taxonomicName>
lacks an anther-gland and has been placed in
<taxonomicName id="4C144D7AFFA8FFEBFDC95445625DF860" authority="Jacq." authorityName="Jacq." class="Magnoliopsida" family="Melastomataceae" genus="Memecylon" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="86" phylum="Tracheophyta" rank="section" section="Clavistamina">
<emphasis id="B960EAEBFFA8FFEBFDC9544561A0F880" box="[598,711,1879,1903]" italics="true" pageId="5" pageNumber="86">Memecylon</emphasis>
sect.
<emphasis id="B960EAEBFFA8FFEBFFEE5465639BF860" box="[113,252,1911,1935]" italics="true" pageId="5" pageNumber="86">Clavistamina</emphasis>
Jacq.
</taxonomicName>
-Fél. sensu
<bibRefCitation id="EF854B08FFA8FFEBFE30546361F1F860" author="JACQUES-FELIX, H." box="[431,662,1905,1935]" pageId="5" pageNumber="86" pagination="3 - 58" refId="ref14998" refString="JACQUES-FELIX, H. (1985 b). Les Memecyleae (Melastomataceae) de Madagascar (2 E partie). Bull. Mus. Natl. Hist. Nat., B, Adansonia 7: 3 - 58." type="journal article" year="1985">JACQUES-FÉLIX (1985</bibRefCitation>
a).
</paragraph>
<paragraph id="8BAB36F9FFA8FFEBFCD8520766C0FE7C" blockId="5.[801,1456,277,403]" pageId="5" pageNumber="86">
<emphasis id="B960EAEBFFA8FFEBFCD85207673FFEC7" box="[839,1112,277,296]" italics="true" pageId="5" pageNumber="86">Additional specimens examined</emphasis>
. –
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<emphasis id="B960EAEBFFA8FFEBFBE95207660BFEC7" bold="true" box="[1142,1388,277,296]" pageId="5" pageNumber="86">
<collectingCountry id="F3037669FFA8FFEBFBE952076791FEC7" box="[1142,1270,277,296]" name="Madagascar" pageId="5" pageNumber="86">MADAGASCAR</collectingCountry>
.
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<collectionCode id="ED05AE3CFFA8FFEBFAAB5207660BFEC7" box="[1332,1388,277,296]" pageId="5" pageNumber="86">SAVA</collectionCode>
</emphasis>
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<materialsCitation id="3B7C3CA4FFA8FFEBFAEC520766C5FE7C" collectingDate="1994-07-08" collectingDateMax="1994-11-03" collectingDateMin="1994-07-08" collectionCode="CAS, G, MO, NU, P, TAN" collectorName="RS d'Anjanaharibe-Sud &amp; Ravelonarivo &amp; Rabesonina" country="Madagascar" elevation="1336" latitude="-14.745" location="Ravelonarivo" longLatPrecision="21" longitude="49.461666" pageId="5" pageNumber="86" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="B960EAEBFFA8FFEBFAEC520760CFFEAC" bold="true" pageId="5" pageNumber="86">
[Prov.
<collectingRegion id="49D0F81BFFA8FFEBFCBE522260F9FEAC" box="[801,926,304,323]" country="Madagascar" name="Antsiranana" pageId="5" pageNumber="86">Antsiranana</collectingRegion>
]:
</emphasis>
<collectorName id="26E1532FFFA8FFEBFC2E522267FFFEAC" box="[945,1176,304,323]" pageId="5" pageNumber="86">RS d’Anjanaharibe-Sud</collectorName>
, env. du sommet,
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,
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,
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<collectingDate id="EFEEE9D1FFA8FFEBFC4852596751FEB1" box="[983,1078,331,350]" pageId="5" pageNumber="86" value="1994-07-08">8.VII.1994</collectingDate>
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, fr.,
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<location id="8ECB6022FFA8FFEBFBC2525867A8FEB2" LSID="urn:lsid:plazi:treatment:03BD87EFFFAEFFEBFFEE555366C0FE7C:8ECB6022FFA8FFEBFBC2525867A8FEB2" box="[1117,1231,330,349]" country="Madagascar" latitude="-14.745" longLatPrecision="21" longitude="49.461666" name="Ravelonarivo" pageId="5" pageNumber="86" stateProvince="Antsiranana">Ravelonarivo</location>
et al. 319
</emphasis>
(
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,
<collectionCode id="ED05AE3CFFA8FFEBFAFA52596610FEB1" box="[1381,1399,331,350]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="86" type="Herbarium">G</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFA1E525966CFFEB1" box="[1409,1448,331,350]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="86" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFCBE52776025FE97" box="[801,834,357,376]" country="Thailand" name="Department of Microbiology, Faculty of Science" pageId="5" pageNumber="86">NU</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFCD352776030FE97" box="[844,855,357,376]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="86" type="Herbarium">P</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFCFF527760F6FE97" box="[864,913,357,376]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="5" pageNumber="86" type="Herbarium">TAN</collectionCode>
); ibid. loco, Camp no 2,
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,
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</quantity>
,
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<collectingDate id="EFEEE9D1FFA8FFEBFCBE52926013FE7C" box="[801,884,384,403]" pageId="5" pageNumber="86" value="1994-11-03">3.XI.1994</collectingDate>
</date>
, fr.,
<emphasis id="B960EAEBFFA8FFEBFC07529267C1FE7C" box="[920,1190,384,403]" italics="true" pageId="5" pageNumber="86">
<collectorName id="26E1532FFFA8FFEBFC0752926760FE7C" box="[920,1031,384,403]" pageId="5" pageNumber="86">Ravelonarivo</collectorName>
&amp;
<collectorName id="26E1532FFFA8FFEBFBBC529267E6FE7C" box="[1059,1153,384,403]" pageId="5" pageNumber="86">Rabesonina</collectorName>
486
</emphasis>
(
<collectionCode id="ED05AE3CFFA8FFEBFB2E529267BAFE7C" box="[1201,1245,384,403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="5" pageNumber="86" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFB795292679FFE7C" box="[1254,1272,384,403]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="86" type="Herbarium">G</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFA9F52926640FE7C" box="[1280,1319,384,403]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="86" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFAAF52926637FE7C" box="[1328,1360,384,403]" country="Thailand" name="Department of Microbiology, Faculty of Science" pageId="5" pageNumber="86">NU</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFAC652926603FE7C" box="[1369,1380,384,403]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="86" type="Herbarium">P</collectionCode>
,
<collectionCode id="ED05AE3CFFA8FFEBFAF4529266FBFE7C" box="[1387,1436,384,403]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="5" pageNumber="86" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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