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<document id="0C7F2698DB5DA0E2D5372FA08A80780C" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BF0FF56AF89FDF1FD103835" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera guomenshanica Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="148" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="146" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="BD17AFCFD750EF8FBA9AAC20F0C296C7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6E8F1C9E0AA3E86CDE7851A5489E3791">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:name id="1D0B1324443DDE8A5C9A5819F22F932E" type="personal">
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<mods:namePart id="CE6C9AF549549146EE17F642050D7043">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="7D96BE827DDC306467C88D642F3B0449" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="0BF8D00DAA4E4719B9A43462EC1D03FB" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:name id="849639693245CD9E8E3A3FE21F2C9F3A" type="personal">
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<mods:namePart id="7ABA89E762FD99729E191E0642479282">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="162DC4C8FF82DE2C2770F021DC347C3F" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="1A7C4760F13F792C13F567BA5FA933B0" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:roleTerm id="1E7000C9434E041DF474B5422372E07E">Author</mods:roleTerm>
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<mods:namePart id="716727EC097174628911661E29FFB61E">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="934064B5272ED6E2A93189317231A51C" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="C4F494FD47B9C5464257A139BCE7E064" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="96CFC4DF6AB173094E4A92974D14D1CF" type="personal">
<mods:role id="0471B5828348E974506B16DABBFE2549">
<mods:roleTerm id="839587AE457D2EA3FD2F74B252B017B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D299BFBFCEC341721E2A83DA3CADD3A">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="CF7870986224A9A1BE2001B48E39F6FE" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="077582E7648CD9CB43A322F7A6849C67" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="ACA37030BB5CBFEFA8DE7825B64B5705">Ahrens, Dirk</mods:namePart>
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<mods:nameIdentifier id="58CBEFDED1ECF80EBC2783150B381E72" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="8AA3E33A1EB8A7F24986CD00832A7D21">Zootaxa</mods:title>
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<mods:part id="6FEA94E96C50D29CC8BCA27503102AF0">
<mods:date id="80941F758300F42DD30F127CD0BDE85D">2021</mods:date>
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<mods:number id="0A4803CB0026AE5E392C76E51086E325">2021-02-03</mods:number>
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<mods:detail id="72B9389D8819939F77C74C1A856D423D" type="volume">
<mods:number id="1CECFED175FEEEC394263BF23501ADED">4922</mods:number>
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<mods:classification id="00930E2B17F25D31EA460EF9CE3846BB">journal article</mods:classification>
<mods:identifier id="F0D53521BFB4818D928FF1938B888DE8" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="BA766AD2B7E15C035C1C0417F97B6F36" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
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<treatment id="03BD87E66BF0FF56AF89FDF1FD103835" ID-DOI="http://doi.org/10.5281/zenodo.4534595" ID-GBIF-Taxon="177323540" ID-Zenodo-Dep="4534595" LSID="urn:lsid:plazi:treatment:03BD87E66BF0FF56AF89FDF1FD103835" httpUri="http://treatment.plazi.org/id/03BD87E66BF0FF56AF89FDF1FD103835" lastPageId="147" lastPageNumber="148" pageId="145" pageNumber="146">
<subSubSection id="C30E657B6BF0FF54AF89FDF1FC9D38FA" box="[151,835,1435,1462]" pageId="145" pageNumber="146" type="nomenclature">
<paragraph id="8BAB36F06BF0FF54AF89FDF1FC9D38FA" blockId="145.[151,835,1435,1498]" box="[151,835,1435,1462]" pageId="145" pageNumber="146">
<heading id="D0E3819C6BF0FF54AF89FDF1FC9D38FA" bold="true" box="[151,835,1435,1462]" fontSize="11" level="2" pageId="145" pageNumber="146" reason="4">
<emphasis id="B960EAE26BF0FF54AF89FDF1FC9D38FA" bold="true" box="[151,835,1435,1462]" pageId="145" pageNumber="146">
<taxonomicName id="4C144D736BF0FF54AF89FDF1FD2A38FA" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,756,1435,1462]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="145" pageNumber="146" phylum="Arthropoda" rank="species" species="guomenshanica" status="sp. nov.">
<emphasis id="B960EAE26BF0FF54AF89FDF1FE0A38FA" bold="true" box="[151,468,1435,1462]" italics="true" pageId="145" pageNumber="146">Maladera guomenshanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BF0FF54AC1EFDF6FC9D38FA" box="[768,835,1436,1462]" pageId="145" pageNumber="146" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF0FF54AF89FDAAFEA33896" box="[151,381,1472,1498]" pageId="145" pageNumber="146" type="description">
<paragraph id="8BAB36F06BF0FF54AF89FDAAFEA33896" blockId="145.[151,835,1435,1498]" box="[151,381,1472,1498]" pageId="145" pageNumber="146">
<figureCitation id="132F2A756BF0FF54AF89FDAAFED03896" box="[151,270,1472,1498]" captionStart="FIGURE 32" captionStartId="146.[151,250,1909,1933]" captionTargetBox="[234,1349,199,1875]" captionTargetId="figure-19@146.[222,1366,181,1885]" captionTargetPageId="146" captionText="FIGURE 32. A–D: Maladera guomenshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. hansmalickyi Ahrens, Fabrizi &amp; Liu, sp. n.; I–M: M. jaintiaensis Ahrens &amp; Fabrizi, 2016 (holotype). A, E, I: aedeagus, left side lateral view; C, G, L: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; K: parameres, ventral view;D, H, M: habitus. Scale: 0.5 mm. Habitus not to scale.;" figureDoi="http://doi.org/10.5281/zenodo.4496390" httpUri="https://zenodo.org/record/4496390/files/figure.png" pageId="145" pageNumber="146">Figures 32</figureCitation>
A–D, 102
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF0FF54AF89FE62FF213B26" pageId="145" pageNumber="146" type="materials_examined">
<paragraph id="8BAB36F06BF0FF54AF89FE62FF213B26" blockId="145.[151,1437,1543,2039]" pageId="145" pageNumber="146">
<emphasis id="B960EAE26BF0FF54AF89FE62FE623B6E" bold="true" box="[151,444,1544,1570]" pageId="145" pageNumber="146">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BF0FF54AEDCFE62FEE83B0A" ID-GBIF-Occurrence="3034556039" collectingDate="2009-04-26" collectionCode="ISAZ" country="China" location="Guomenshan" municipality="Nabanhe Nature Reserve" pageId="145" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54AF88526BF0FF54AEDCFE62FDF13B6E" box="[450,559,1544,1570]" pageId="145" pageNumber="146" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796BF0FF54AD29FE63FD923B6F" box="[567,588,1545,1571]" pageId="145" pageNumber="146" type="male">♂</specimenCount>
“[
<collectingCountry id="F30376606BF0FF54AD7DFE6DFD733B6E" box="[611,685,1543,1570]" name="China" pageId="145" pageNumber="146">China</collectingCountry>
]
<collectingRegion id="49D0F8126BF0FF54ADA6FE63FCCF3B6F" box="[696,785,1545,1571]" country="China" name="Yunnan" pageId="145" pageNumber="146">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCFAC8A6BF0FF54AC04FE62FBEC3B6F" box="[794,1074,1544,1571]" pageId="145" pageNumber="146">Nabanhe Nature Reserve</collectingMunicipality>
,
<location id="8ECB602B6BF0FF54AB22FE62FB0D3B6E" LSID="urn:lsid:plazi:treatment:03BD87E66BF0FF56AF89FDF1FD103835:8ECB602B6BF0FF54AB22FE62FB0D3B6E" box="[1084,1235,1544,1570]" country="China" municipality="Nabanhe Nature Reserve" name="Guomenshan" pageId="145" pageNumber="146" stateProvince="Yunnan">Guomenshan</location>
,
<date id="FFAA10306BF0FF54ABC2FE62FA823B6E" box="[1244,1372,1544,1570]" pageId="145" pageNumber="146" value="2009-04-26">
<collectingDate id="EFEEE9D86BF0FF54ABC2FE62FA823B6E" box="[1244,1372,1544,1570]" pageId="145" pageNumber="146" value="2009-04-26">2009-IV-26</collectingDate>
</date>
/ LW-1165” (
<collectionCode id="ED05AE356BF0FF54AFF4FE46FEF03B0A" box="[234,302,1580,1606]" pageId="145" pageNumber="146">ISAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BF0FF54AE5FFE47FF253B26" ID-GBIF-Occurrence="3034556332" collectingDate="2009-04-26" collectionCode="ZFMK" country="China" location="Guomenshan" municipality="Nabanhe Nature Reserve" pageId="145" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54AF88526BF0FF54AE5FFE47FE793B0B" box="[321,423,1581,1607]" pageId="145" pageNumber="146" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D12FD796BF0FF54AEACFE46FE023B0B" box="[434,476,1580,1607]" pageId="145" pageNumber="146" type="male">1 ♂</specimenCount>
“[
<collectingCountry id="F30376606BF0FF54AEEAFE41FDE03B0A" box="[500,574,1579,1606]" name="China" pageId="145" pageNumber="146">China</collectingCountry>
]
<collectingRegion id="49D0F8126BF0FF54AD55FE47FD7A3B0B" box="[587,676,1581,1607]" country="China" name="Yunnan" pageId="145" pageNumber="146">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCFAC8A6BF0FF54ADB0FE46FC173B0B" box="[686,969,1580,1607]" pageId="145" pageNumber="146">Nabanhe Nature Reserve</collectingMunicipality>
,
<location id="8ECB602B6BF0FF54ACCBFE46FBB23B0A" LSID="urn:lsid:plazi:treatment:03BD87E66BF0FF56AF89FDF1FD103835:8ECB602B6BF0FF54ACCBFE46FBB23B0A" box="[981,1132,1580,1606]" country="China" municipality="Nabanhe Nature Reserve" name="Guomenshan" pageId="145" pageNumber="146" stateProvince="Yunnan">Guomenshan</location>
,
<date id="FFAA10306BF0FF54AB68FE46FB293B0A" box="[1142,1271,1580,1606]" pageId="145" pageNumber="146" value="2009-04-26">
<collectingDate id="EFEEE9D86BF0FF54AB68FE46FB293B0A" box="[1142,1271,1580,1606]" pageId="145" pageNumber="146" value="2009-04-26">2009-IV-26</collectingDate>
</date>
/ LW-1165bis” (
<collectionCode id="ED05AE356BF0FF54AF81FE3AFF2D3B26" box="[159,243,1616,1642]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="145" pageNumber="146" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF0FF56AFD9FE1EFCA93915" lastPageId="147" lastPageNumber="148" pageId="145" pageNumber="146" type="description">
<paragraph id="8BAB36F06BF0FF54AFD9FE1EFC2B3BFE" blockId="145.[151,1437,1543,2039]" pageId="145" pageNumber="146">
<emphasis id="B960EAE26BF0FF54AFD9FE1EFE873BC2" bold="true" box="[199,345,1652,1678]" pageId="145" pageNumber="146">Description.</emphasis>
Length: 9.0 mm, length of elytra:
<quantity id="4CEC9B156BF0FF54ADFDFE1EFCE23BC3" box="[739,828,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="145" pageNumber="146" unit="mm" value="6.2">6.2 mm</quantity>
, width:
<quantity id="4CEC9B156BF0FF54AC86FE1EFC2C3BC3" box="[920,1010,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="145" pageNumber="146" unit="mm" value="5.2">5.2 mm</quantity>
. Body oblong-oval, yellowish brown, dull, labroclypeus shiny, with a few long setae on head, pronotum and elytra.
</paragraph>
<paragraph id="8BAB36F06BF0FF54AFD9FED6FC9A3ABA" blockId="145.[151,1437,1543,2039]" pageId="145" pageNumber="146">
Labroclypeus short and wide, subtrapezoidal, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin shallowly emarginate, margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface distinctly carinate along middle, finely, densely punctate, with a few erect, long setae; frontoclypeal suture distinctly incised, bluntly angled medially; smooth area anterior to eye very small and flat, three times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with sparse and fine punctures, with robust and long setae beside eyes and behind frontoclypeal suture. Eyes moderately small, ratio diameter/ interocular width: 0.57. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126BF0FF54AE0AFFB7FEAD3ABB" box="[276,371,2013,2039]" country="Myanmar" name="Chin" pageId="145" pageNumber="146">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<caption id="DF6B66786BF3FF57AF89FF1FFEB33AB5" ID-DOI="http://doi.org/10.5281/zenodo.4496390" ID-Zenodo-Dep="4496390" httpUri="https://zenodo.org/record/4496390/files/figure.png" pageId="146" pageNumber="147" startId="146.[151,250,1909,1933]" targetBox="[234,1349,199,1875]" targetPageId="146">
<paragraph id="8BAB36F06BF3FF57AF89FF1FFEB33AB5" blockId="146.[151,1437,1909,2041]" pageId="146" pageNumber="147">
<emphasis id="B960EAE26BF3FF57AF89FF1FFE803AC1" bold="true" box="[151,350,1909,1933]" pageId="146" pageNumber="147">FIGURE 32. A–D:</emphasis>
<taxonomicName id="4C144D736BF3FF57AE7AFF1FFC883AC1" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[356,854,1909,1933]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="species" species="guomenshanica" status="sp. nov.">
<emphasis id="B960EAE26BF3FF57AE7AFF1FFDB33AC1" box="[356,621,1909,1933]" italics="true" pageId="146" pageNumber="147">Maladera guomenshanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BF3FF57AC7EFF1FFC483AC0" box="[864,918,1909,1932]" pageId="146" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26BF3FF57AB0DFF1FFB923AC1" bold="true" box="[1043,1100,1909,1933]" pageId="146" pageNumber="147">E–H:</emphasis>
<taxonomicName id="4C144D736BF3FF57AB4DFF1CFF063AFD" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="species" species="hansmalickyi" status="sp. nov.">
<emphasis id="B960EAE26BF3FF57AB4DFF1CFB243AC1" box="[1107,1274,1909,1933]" italics="true" pageId="146" pageNumber="147">M. hansmalickyi</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BF3FF57AFFDFFF3FEC33AFC" box="[227,285,1945,1968]" pageId="146" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
;
<emphasis id="B960EAE26BF3FF57AE37FFF3FEBE3AFD" bold="true" box="[297,352,1945,1969]" pageId="146" pageNumber="147">I–M:</emphasis>
<taxonomicName id="4C144D736BF3FF57AE79FFF0FD273AFD" authority="Ahrens &amp; Fabrizi, 2016" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" box="[359,761,1945,1969]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="species" species="jaintiaensis">
<emphasis id="B960EAE26BF3FF57AE79FFF0FE213AFD" box="[359,511,1946,1969]" italics="true" pageId="146" pageNumber="147">M. jaintiaensis</emphasis>
<bibRefCitation id="EF854B016BF3FF57AD1BFFF3FD273AFD" author="Ahrens, D. &amp; Fabrizi, S." box="[517,761,1945,1969]" pageId="146" pageNumber="147" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi, 2016</bibRefCitation>
</taxonomicName>
(holotype).
<emphasis id="B960EAE26BF3FF57AC66FFF3FC173AFD" bold="true" box="[888,969,1945,1969]" pageId="146" pageNumber="147">A, E, I:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26BF3FF57AA05FFF3FAAB3AFD" bold="true" box="[1307,1397,1945,1969]" pageId="146" pageNumber="147">C, G, L:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26BF3FF57AEC9FFD7FDF83A99" bold="true" box="[471,550,1981,2005]" pageId="146" pageNumber="147">B, F, J:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26BF3FF57AC38FFD7FC9C3A99" bold="true" box="[806,834,1981,2005]" pageId="146" pageNumber="147">K:</emphasis>
parameres, ventral view;
<emphasis id="B960EAE26BF3FF57AB5AFFD7FB7B3A99" bold="true" box="[1092,1189,1981,2005]" pageId="146" pageNumber="147">D, H, M:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.;
</paragraph>
</caption>
<paragraph id="8BAB36F06BF2FF56AFD9F8FDFB813C29" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">
Pronotum moderately transverse, widest shortly before base, lateral margins evenly convex and convergent anteriorly and posteriorly, anterior angles distinctly produced and sharp, posterior angles blunt, rounded at tip; anterior margin straight, with fine marginal line, base without marginal line; surface moderately finely and densely punctate, with minute setae in punctures and a very few long, semi-erect setae in coarse punctures on lateral disc (abraded in
<typeStatus id="54AF88526BF2FF56AF89F94DFF223C0D" box="[151,252,295,321]" pageId="147" pageNumber="148" type="holotype">holotype</typeStatus>
, but large punctures visible); anterior and lateral margin robustly densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, moderately dense punctures.
</paragraph>
<paragraph id="8BAB36F06BF2FF56AFD9F905FA463CBA" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, second one flat, with fine, moderately dense punctures and with minute setae in punctures, odd intervals with numerous single long setae in darker coarse punctures; epipleural margin robust, ending half of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06BF2FF56AFD9F995FD7C3F81" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">Ventral surface dull, finely and densely punctate, nearly glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.98. Pygidium weakly convex, dull, finely and moderately densely punctate, without smooth midline, with a few long setae along apical margin, otherwise only with minute setae in punctures.</paragraph>
<paragraph id="8BAB36F06BF2FF56AFD9FABDFA423979" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">
Legs short and wide, dull; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line, anterior row of setae completely reduced; posterior ventral margin smooth, strongly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.4, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, superficially and sparsely punctate, along midline widely smooth, glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres lacking in
<typeStatus id="54AF88526BF2FF56AA2DFB9DFA47395D" box="[1331,1433,1015,1041]" pageId="147" pageNumber="148" type="holotype">holotype</typeStatus>
. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
</paragraph>
<paragraph id="8BAB36F06BF2FF56AFD9FC55FCA93915" blockId="147.[151,1437,151,1401]" box="[199,887,1087,1113]" pageId="147" pageNumber="148">
Aedeagus:
<figureCitation id="132F2A756BF2FF56AE5AFC55FE4A3915" box="[324,404,1087,1113]" captionStart="FIGURE 32" captionStartId="146.[151,250,1909,1933]" captionTargetBox="[234,1349,199,1875]" captionTargetId="figure-19@146.[222,1366,181,1885]" captionTargetPageId="146" captionText="FIGURE 32. A–D: Maladera guomenshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. hansmalickyi Ahrens, Fabrizi &amp; Liu, sp. n.; I–M: M. jaintiaensis Ahrens &amp; Fabrizi, 2016 (holotype). A, E, I: aedeagus, left side lateral view; C, G, L: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; K: parameres, ventral view;D, H, M: habitus. Scale: 0.5 mm. Habitus not to scale.;" figureDoi="http://doi.org/10.5281/zenodo.4496390" httpUri="https://zenodo.org/record/4496390/files/figure.png" pageId="147" pageNumber="148">Fig. 32</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BF2FF56AD23FC55FD7A3915" box="[573,676,1087,1113]" captionStart="FIGURE 32" captionStartId="146.[151,250,1909,1933]" captionTargetBox="[234,1349,199,1875]" captionTargetId="figure-19@146.[222,1366,181,1885]" captionTargetPageId="146" captionText="FIGURE 32. A–D: Maladera guomenshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. hansmalickyi Ahrens, Fabrizi &amp; Liu, sp. n.; I–M: M. jaintiaensis Ahrens &amp; Fabrizi, 2016 (holotype). A, E, I: aedeagus, left side lateral view; C, G, L: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; K: parameres, ventral view;D, H, M: habitus. Scale: 0.5 mm. Habitus not to scale.;" figureDoi="http://doi.org/10.5281/zenodo.4496390" httpUri="https://zenodo.org/record/4496390/files/figure.png" pageId="147" pageNumber="148">Fig. 32D</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF2FF56AFD9FC08FBDC39ED" pageId="147" pageNumber="148" type="diagnosis">
<paragraph id="8BAB36F06BF2FF56AFD9FC08FBDC39ED" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">
<emphasis id="B960EAE26BF2FF56AFD9FC08FE9C3930" bold="true" box="[199,322,1122,1148]" pageId="147" pageNumber="148">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BF2FF56AE55FC09FC5F3931" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[331,897,1123,1149]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="147" pageNumber="148" phylum="Arthropoda" rank="species" species="guomenshanica" status="sp. nov.">
<emphasis id="B960EAE26BF2FF56AE55FC09FDAD3931" box="[331,627,1123,1149]" italics="true" pageId="147" pageNumber="148">Maladera guomenshanica</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BF2FF56AC90FC09FC133931" box="[910,973,1123,1149]" pageId="147" pageNumber="148" rank="species">sp. n.</taxonomicNameLabel>
differs from
<taxonomicName id="4C144D736BF2FF56AB77FC09FAA33931" authority="(Murayama)" baseAuthorityName="Murayama" baseAuthorityYear="1938" box="[1129,1405,1123,1149]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="147" pageNumber="148" phylum="Arthropoda" rank="species" species="aureola">
<emphasis id="B960EAE26BF2FF56AB77FC09FB393931" box="[1129,1255,1123,1149]" italics="true" pageId="147" pageNumber="148">M. aureola</emphasis>
(Murayama)
</taxonomicName>
in having longer parameres, and denser setae on the disc of pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF2FF56AFD9FCC0FB213841" pageId="147" pageNumber="148" type="description">
<paragraph id="8BAB36F06BF2FF56AFD9FCC0FB213841" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">
<emphasis id="B960EAE26BF2FF56AFD9FCC0FEE13988" bold="true" box="[199,319,1194,1220]" pageId="147" pageNumber="148">Variation.</emphasis>
Length: 9.0–
<quantity id="4CEC9B156BF2FF56AECAFCC1FDF53989" box="[468,555,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="147" pageNumber="148" unit="mm" value="9.2">9.2 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BF2FF56ADF1FCC1FCA63989" box="[751,888,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.6" metricValueMin="6.2" pageId="147" pageNumber="148" unit="mm" value="6.4" valueMax="6.6" valueMin="6.2">6.2–6.6 mm</quantity>
, width: 5.0–
<quantity id="4CEC9B156BF2FF56AB1CFCC0FB843989" box="[1026,1114,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="147" pageNumber="148" unit="mm" value="5.2">5.2 mm</quantity>
. Metatarsomeres of
<typeStatus id="54AF88526BF2FF56AA22FCC1FA423989" box="[1340,1436,1195,1221]" pageId="147" pageNumber="148" type="paratype">paratype</typeStatus>
with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF2FF56AFD9FD7DFEED3819" pageId="147" pageNumber="148" type="etymology">
<paragraph id="8BAB36F06BF2FF56AFD9FD7DFEED3819" blockId="147.[151,1437,151,1401]" pageId="147" pageNumber="148">
<emphasis id="B960EAE26BF2FF56AFD9FD7DFE90387D" bold="true" box="[199,334,1303,1329]" pageId="147" pageNumber="148">Etymology.</emphasis>
The name of the new species (adjective in the nominative singular) is derived from its
<typeStatus id="54AF88526BF2FF56AA13FD7DFAE3387D" box="[1293,1341,1303,1329]" pageId="147" pageNumber="148">type</typeStatus>
locality, Guomenshan.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BF2FF56AFD9FD34FD103835" box="[199,718,1374,1401]" pageId="147" pageNumber="148" type="distribution">
<paragraph id="8BAB36F06BF2FF56AFD9FD34FD103835" blockId="147.[151,1437,151,1401]" box="[199,718,1374,1401]" pageId="147" pageNumber="148">
<emphasis id="B960EAE26BF2FF56AFD9FD34FEBF3834" bold="true" box="[199,353,1374,1400]" pageId="147" pageNumber="148">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BF2FF56AEC8FD35FDE83835" box="[470,566,1375,1401]" captionStart="FIGURE 102" captionStartId="352.[151,250,1967,1991]" captionTargetBox="[213,1372,1129,1938]" captionTargetId="figure-44@352.[208,1380,1124,1944]" captionTargetPageId="352" captionText="FIGURE 102. Distribution of the Maladera thomsoni group: Maladera apicalis sp. n., M.aureola (Murayama, 1938), M. botrytibia Nomura, 1974, M. guomenshanica sp. n., M. hansmalickyi sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496592" httpUri="https://zenodo.org/record/4496592/files/figure.png" pageId="147" pageNumber="148">Fig. 102</figureCitation>
) and
<tableCitation id="C696034B6BF2FF56AD6AFD35FD163835" box="[628,712,1375,1401]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="147" pageNumber="148">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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