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<document id="2097A92C7184A849F5DB56EE2CD0755A" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BE0FF47AF89FAD8FD103E85" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera pseudofuscipes Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="9" lastPageNumber="131" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="130" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="103FFF99CB3AC9D4DC09B8B2754EF7B5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D9B2D79B25BDF42F27FF0881203526F6">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:name id="C0131B150EEDC7EE8C7963563BFF98C4" type="personal">
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<mods:roleTerm id="02CFC4D3D7AF4D573E3333752F3FAB86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8480470993787036B431147E0FFF6828">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="54AF8A1A57A04655546A9107279754DF" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="5784E21EF0E16B6D5F34BB6AC43036B6" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:name id="2AD51C173657F662956D513F85FD2743" type="personal">
<mods:role id="B40815D40197C80226331CAA22F17A8B">
<mods:roleTerm id="E112BBF93DB62C5F72BD6F7CFD26E869">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F55BB8295C113D9E83B46059AE7A4368">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="4A97979BBF08BF4A4EB5943FC116933E" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="A4D938A0D3F1397355F7891249D43D62" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="1863DBC9A6E4F343BF258808FECB0262" type="personal">
<mods:role id="07124BC7B43E0CBCB9EFA123B16BFAFE">
<mods:roleTerm id="EA592D329CEC5FCFF1E85311C8F36C24">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7620ADCE2BE50105D2ABCD1A772E3D51">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="201578F8CC5D306098640AF7E6B91901" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="3E2E0D5329984B37823364BE8F4AC320" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="BE6E4DE73274D825FF078F3B31ABFDAF" type="personal">
<mods:role id="F1386F4323FB166358E27363BA74680E">
<mods:roleTerm id="FEB02F4F8FA7921335B3B5B36FAB21A2">Author</mods:roleTerm>
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<mods:namePart id="67019CEA8B251F58402640AE5ACC9CC1">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="5C86928158B246436E1FA7D3D5FCFB1C" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="DE316D25E07999F7A5C1C2AE804C8115" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="655811A3F501738027267BBA77C2D758">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="13ED23656F8B1D7727412D96681250CF" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
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<mods:title id="1C0A0CF2F995EC9090E7A3FB7D25299E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CB15260D9C439930EAD031941A36A8D7">
<mods:date id="68E12ED0560FBA6F1E9AB260E61539D7">2021</mods:date>
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<mods:number id="F29A441EE8170C6CB87838D9EB8E3F73">2021-02-03</mods:number>
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<mods:number id="3AF3FDCB2F031D0E2B841E6D1C188E8B">4922</mods:number>
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<mods:classification id="1C71D0BF50C02E6C3BEBE435D6A2B978">journal article</mods:classification>
<mods:identifier id="76F86775FF40E3B66C5F5E1FC7218974" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="91A42887E1A429DB18F7F8C178B0CA53" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="BB675B93AC74910DC001B0DFD946B9BD" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="E0D8D1F870A16385FB4709A8BB085BF7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BD87E66BE0FF47AF89FAD8FD103E85" ID-DOI="http://doi.org/10.5281/zenodo.4534575" ID-GBIF-Taxon="177323561" ID-Zenodo-Dep="4534575" LSID="urn:lsid:plazi:treatment:03BD87E66BE0FF47AF89FAD8FD103E85" httpUri="http://treatment.plazi.org/id/03BD87E66BE0FF47AF89FAD8FD103E85" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">
<subSubSection id="C30E657B6BE0FF44AF89FAD8FCE43F81" box="[151,826,690,717]" pageId="129" pageNumber="130" type="nomenclature">
<paragraph id="8BAB36F06BE0FF44AF89FAD8FCE43F81" blockId="129.[151,826,690,753]" box="[151,826,690,717]" pageId="129" pageNumber="130">
<heading id="D0E3819C6BE0FF44AF89FAD8FCE43F81" bold="true" box="[151,826,690,717]" fontSize="11" level="2" pageId="129" pageNumber="130" reason="4">
<emphasis id="B960EAE26BE0FF44AF89FAD8FCE43F81" bold="true" box="[151,826,690,717]" pageId="129" pageNumber="130">
<taxonomicName id="4C144D736BE0FF44AF89FAD8FD353F80" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,747,690,717]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="pseudofuscipes" status="sp. nov.">
<emphasis id="B960EAE26BE0FF44AF89FAD8FE153F81" bold="true" box="[151,459,690,717]" italics="true" pageId="129" pageNumber="130">Maladera pseudofuscipes</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BE0FF44ADE6FAD9FCE43F81" box="[760,826,691,717]" pageId="129" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE0FF44AF89FABDFEA33FBD" box="[151,381,727,753]" pageId="129" pageNumber="130" type="description">
<paragraph id="8BAB36F06BE0FF44AF89FABDFEA33FBD" blockId="129.[151,826,690,753]" box="[151,381,727,753]" pageId="129" pageNumber="130">
<figureCitation id="132F2A756BE0FF44AF89FABDFED03FBD" box="[151,270,727,753]" captionStart="FIGURE 28" captionStartId="131.[151,250,1909,1933]" captionTargetBox="[197,1399,191,1873]" captionTargetId="figure-20@131.[188,1399,181,1884]" captionTargetPageId="131" captionText="FIGURE 28. A–D: Maladera pseudofuscipes Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–I: M. snizeki Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. fangana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; F: aedeagus, dorsal view; B, G, K: parameres, dorsal view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496380" httpUri="https://zenodo.org/record/4496380/files/figure.png" pageId="129" pageNumber="130">Figures 28</figureCitation>
A–D, 100
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE0FF44AF89FB75FE523915" pageId="129" pageNumber="130" type="materials_examined">
<paragraph id="8BAB36F06BE0FF44AF89FB75FE523915" blockId="129.[151,1437,798,2013]" pageId="129" pageNumber="130">
<emphasis id="B960EAE26BE0FF44AF89FB75FE1A3E74" bold="true" box="[151,452,798,825]" pageId="129" pageNumber="130">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BE0FF44AED3FB75FD3F3E11" ID-GBIF-Occurrence="3034555332" collectingDate="1999-05-23" collectingDateMax="1999-05-27" collectingDateMin="1999-05-23" collectionCode="ZFMK" collectorName="Ahrens" elevation="1650" location="Sa Pa" pageId="129" pageNumber="130" specimenCount="1" stateProvince="Prov. Lao Cai" typeStatus="holotype">
<typeStatus id="54AF88526BE0FF44AED3FB75FDE43E75" box="[461,570,799,825]" pageId="129" pageNumber="130" type="holotype">Holotype</typeStatus>
: “N-Vietnam,
<collectingRegion id="49D0F8126BE0FF44ADFEFB75FC5C3E75" box="[736,898,799,825]" inferredBySuffix="true" pageId="129" pageNumber="130">Prov. Lao Cai</collectingRegion>
,
<location id="8ECB602B6BE0FF44AC91FB75FC0A3E75" LSID="urn:lsid:plazi:treatment:03BD87E66BE0FF47AF89FAD8FD103E85:8ECB602B6BE0FF44AC91FB75FC0A3E75" box="[911,980,799,825]" name="Sa Pa" pageId="129" pageNumber="130" stateProvince="Prov. Lao Cai">Sa Pa</location>
,
<quantity id="4CEC9B156BE0FF44ACFFFB75FBA23E75" box="[993,1148,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="129" pageNumber="130" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="0039D1C36BE0FF44ACFFFB75FBA23E75" box="[993,1148,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="129" pageNumber="130" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600-1700 m</elevation>
</quantity>
, N22°19‘52‘‘; E103°50‘;
<date id="FFAA10306BE0FF44AF89FB29FEE93E11" box="[151,311,835,861]" pageId="129" pageNumber="130" value="1999-05-23" valueMax="1999-05-27" valueMin="1999-05-23">
<collectingDate id="EFEEE9D86BE0FF44AF89FB29FEE93E11" box="[151,311,835,861]" pageId="129" pageNumber="130" value="1999-05-23" valueMax="1999-05-27" valueMin="1999-05-23">23.-27.V.1999</collectingDate>
</date>
leg.
<collectorName id="26E153266BE0FF44AE71FB29FE1D3E11" box="[367,451,835,861]" pageId="129" pageNumber="130">Ahrens</collectorName>
, Jäger, Fabrizi” (
<collectionCode id="ED05AE356BE0FF44AD9BFB29FD073E11" box="[645,729,835,861]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="129" pageNumber="130" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BE0FF44ADF3FB29FAC53ECD" ID-GBIF-Occurrence="3034556522" collectingDate="1999-05-23" collectingDateMax="1999-05-27" collectingDateMin="1999-05-23" collectionCode="ZFMK" collectorName="Ahrens &amp; Jager &amp; Fabrizi" country="Vietnam" elevation="1650" location="Sa Pa" pageId="129" pageNumber="130" specimenCount="38" specimenCount-female="18" specimenCount-male="20" stateProvince="Lao Cai" typeStatus="paratype">
<typeStatus id="54AF88526BE0FF44ADF3FB29FC803E11" box="[749,862,835,861]" pageId="129" pageNumber="130" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B960EAE26BE0FF44AC74FB28FC073E10" bold="true" box="[874,985,834,860]" pageId="129" pageNumber="130">
<collectingCountry id="F30376606BE0FF44AC74FB28FC0A3E10" box="[874,980,834,860]" name="Vietnam" pageId="129" pageNumber="130">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BE0FF44ACFFFB29FBEC3E11" box="[993,1074,835,861]" pageId="129" pageNumber="130" type="male">20 ♂♂</specimenCount>
,
<specimenCount id="9D12FD796BE0FF44AB21FB29FB533E11" box="[1087,1165,835,861]" pageId="129" pageNumber="130" type="female">18 ♀♀</specimenCount>
“N-Vietnam, Prov.
<collectingRegion id="49D0F8126BE0FF44AA6FFB29FF1F3ECD" country="Vietnam" name="Lao Cai" pageId="129" pageNumber="130">Lao Cai</collectingRegion>
,
<location id="8ECB602B6BE0FF44AFD3FB0DFED13ECD" LSID="urn:lsid:plazi:treatment:03BD87E66BE0FF47AF89FAD8FD103E85:8ECB602B6BE0FF44AFD3FB0DFED13ECD" box="[205,271,871,897]" country="Vietnam" name="Sa Pa" pageId="129" pageNumber="130" stateProvince="Lao Cai">Sa Pa</location>
,
<quantity id="4CEC9B156BE0FF44AE04FB0DFE6D3ECD" box="[282,435,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="129" pageNumber="130" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="0039D1C36BE0FF44AE04FB0DFE6D3ECD" box="[282,435,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="129" pageNumber="130" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600-1700 m</elevation>
</quantity>
, N22°19‘52‘‘; E103°50‘;
<date id="FFAA10306BE0FF44ADC9FB0DFCA83ECD" box="[727,886,871,897]" pageId="129" pageNumber="130" value="1999-05-23" valueMax="1999-05-27" valueMin="1999-05-23">
<collectingDate id="EFEEE9D86BE0FF44ADC9FB0DFCA83ECD" box="[727,886,871,897]" pageId="129" pageNumber="130" value="1999-05-23" valueMax="1999-05-27" valueMin="1999-05-23">23.-27.V.1999</collectingDate>
</date>
leg.
<collectorName id="26E153266BE0FF44ACB2FB0DFBDE3ECD" box="[940,1024,871,897]" pageId="129" pageNumber="130">Ahrens</collectorName>
,
<collectorName id="26E153266BE0FF44AB12FB0DFB973ECD" box="[1036,1097,871,897]" pageId="129" pageNumber="130">Jäger</collectorName>
,
<collectorName id="26E153266BE0FF44AB4AFB0DFB783ECD" box="[1108,1190,871,897]" pageId="129" pageNumber="130">Fabrizi</collectorName>
” (
<collectionCode id="ED05AE356BE0FF44ABA1FB0DFACD3ECD" box="[1215,1299,871,897]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="129" pageNumber="130" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BE0FF44AA38FB0DFBAF3EE9" ID-GBIF-Occurrence="3034555315" collectionCode="CP" country="Vietnam" elevation="1500" location="Hoang Lien" pageId="129" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D12FD796BE0FF44AA38FB0DFA8F3ECD" box="[1318,1361,871,897]" pageId="129" pageNumber="130" type="male">1 ♂</specimenCount>
“Vietnam N 1990 Sa-Pa
<date id="FFAA10306BE0FF44AE70FBE1FDC63EE9" box="[366,536,906,933]" pageId="129" pageNumber="130" value="1500-06-11" valueMax="1500-06-19" valueMin="1500-06-11">
11-19.VI.
<quantity id="4CEC9B156BE0FF44AEFFFBE0FDC63EE9" box="[481,536,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="129" pageNumber="130" unit="m" value="1500.0">
<elevation id="0039D1C36BE0FF44AEFFFBE0FDC63EE9" box="[481,536,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="129" pageNumber="130" unit="m" value="1500.0">1500</elevation>
</quantity>
</date>
m
<location id="8ECB602B6BE0FF44AD2BFBE1FD673EE9" LSID="urn:lsid:plazi:treatment:03BD87E66BE0FF47AF89FAD8FD103E85:8ECB602B6BE0FF44AD2BFBE1FD673EE9" box="[565,697,907,933]" country="Vietnam" name="Hoang Lien" pageId="129" pageNumber="130">Hoang Lien</location>
son prov. Strnad Jan lgt./ VS130” (
<collectionCode id="ED05AE356BE0FF44AB5CFBE1FBB73EE9" box="[1090,1129,907,933]" pageId="129" pageNumber="130">CP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BE0FF44AB62FBE0FA473E85" ID-GBIF-Occurrence="3034555616" collectingDate="2010-05-15" collectingDateMax="2010-05-16" collectingDateMin="2010-05-15" collectionCode="ZFMK" collectorName="L. Dembicky" country="Vietnam" elevation="50" latitude="22.7625" location="12 km NE Cao Bang" longLatPrecision="853" longitude="106.316666" pageId="129" pageNumber="130" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Cao Bang" typeStatus="paratype">
<specimenCount id="9D12FD796BE0FF44AB62FBE0FB623EE9" box="[1148,1212,906,933]" pageId="129" pageNumber="130" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D12FD796BE0FF44ABD6FBE1FAD83EE9" box="[1224,1286,907,933]" pageId="129" pageNumber="130" type="female">2 ♀♀</specimenCount>
“Vietnam-N,
<collectingRegion id="49D0F8126BE0FF44AF89FBC5FE9B3E85" box="[151,325,943,969]" country="Vietnam" name="Cao Bang" pageId="129" pageNumber="130">Cao Bang prov.</collectingRegion>
<location id="8ECB602B6BE0FF44AE54FBC5FDED3E85" LSID="urn:lsid:plazi:treatment:03BD87E66BE0FF47AF89FAD8FD103E85:8ECB602B6BE0FF44AE54FBC5FDED3E85" box="[330,563,943,969]" country="Vietnam" latitude="22.7625" longLatPrecision="853" longitude="106.316666" name="12 km NE Cao Bang" pageId="129" pageNumber="130" stateProvince="Cao Bang">
<quantity id="4CEC9B156BE0FF44AE54FBC5FE4E3E85" box="[330,400,943,969]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="129" pageNumber="130" unit="km" value="12.0">12 km</quantity>
NE
<collectingRegion id="49D0F8126BE0FF44AEDCFBC5FDED3E85" box="[450,563,943,969]" country="Vietnam" name="Cao Bang" pageId="129" pageNumber="130">Cao Bang</collectingRegion>
</location>
, 650+
<quantity id="4CEC9B156BE0FF44AD68FBC4FD763E85" box="[630,680,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="129" pageNumber="130" unit="m" value="50.0">
<elevation id="0039D1C36BE0FF44AD68FBC4FD763E85" box="[630,680,942,969]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="129" pageNumber="130" unit="m" value="50.0">50m</elevation>
</quantity>
<geoCoordinate id="EE2050376BE0FF44ADB1FBC4FCE23E85" box="[687,828,942,969]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="129" pageNumber="130" precision="15" seconds="45" value="22.7625">22°45’45’’N</geoCoordinate>
<geoCoordinate id="EE2050376BE0FF44AC5CFBC5FC703E85" box="[834,942,943,969]" degrees="106" direction="east" minutes="19" orientation="longitude" pageId="129" pageNumber="130" precision="925" value="106.316666">106°19’E</geoCoordinate>
<collectorName id="26E153266BE0FF44ACAAFBC5FB993E85" box="[948,1095,943,969]" pageId="129" pageNumber="130">L. Dembicky</collectorName>
leg.,
<date id="FFAA10306BE0FF44AB9DFBC4FAFA3E85" box="[1155,1316,942,969]" pageId="129" pageNumber="130" value="2010-05-15" valueMax="2010-05-16" valueMin="2010-05-15">
<collectingDate id="EFEEE9D86BE0FF44AB9DFBC4FAFA3E85" box="[1155,1316,942,969]" pageId="129" pageNumber="130" value="2010-05-15" valueMax="2010-05-16" valueMin="2010-05-15">15.-16.V.2010</collectingDate>
</date>
” (
<collectionCode id="ED05AE356BE0FF44AA23FBC5FA4F3E85" box="[1341,1425,943,969]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="129" pageNumber="130" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BE0FF44AF89FBB8FDB7395D" ID-GBIF-Occurrence="3034556552" collectionCode="CP" country="China" elevation="1550" location="Yunnan" pageId="129" pageNumber="130" specimenCount="11" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B960EAE26BE0FF44AF89FBB8FF343EA0" bold="true" box="[151,234,978,1004]" pageId="129" pageNumber="130">
<collectingCountry id="F30376606BE0FF44AF89FBB8FF3A3EA0" box="[151,228,978,1004]" name="China" pageId="129" pageNumber="130">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BE0FF44AFEBFBB9FEFA3EA1" box="[245,292,979,1005]" pageId="129" pageNumber="130" type="male">1 ♂</specimenCount>
(+
<specimenCount id="9D12FD796BE0FF44AE56FBB9FE4C3EA1" box="[328,402,979,1005]" pageId="129" pageNumber="130" type="generic">10 ex.</specimenCount>
unsexed) “
<collectingCountry id="F30376606BE0FF44AD06FBB9FDBE3EA1" box="[536,608,979,1005]" name="China" pageId="129" pageNumber="130">China</collectingCountry>
, SE
<collectingRegion id="49D0F8126BE0FF44AD85FBB9FD2A3EA1" box="[667,756,979,1005]" country="China" name="Yunnan" pageId="129" pageNumber="130">Yunnan</collectingRegion>
, Xichou—E env.,
<quantity id="4CEC9B156BE0FF44ACD0FBB9FBBE3EA1" box="[974,1120,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="129" pageNumber="130" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">
<elevation id="0039D1C36BE0FF44ACD0FBB9FBBE3EA1" box="[974,1120,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="129" pageNumber="130" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400-1700m</elevation>
</quantity>
, 13.-18.5.95 23°22-26’[N]/ 104°41-49’[E] L.+R. Businsky lgt.” (
<collectionCode id="ED05AE356BE0FF44AD24FB9DFDBF395D" box="[570,609,1015,1041]" pageId="129" pageNumber="130">CP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BE0FF44AD6BFB9DFECE3979" ID-GBIF-Occurrence="3034556494" collectingDate="1994-06-24" collectingDateMax="1996-05-19" collectingDateMin="1994-06-24" collectionCode="CP" collectorName="Vit Kuban" country="Vietnam" elevation="1375" latitude="23.14" location="Hanoi" longLatPrecision="754" longitude="104.34" pageId="129" pageNumber="130" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi" typeStatus="paratype">
<specimenCount id="9D12FD796BE0FF44AD6BFB9DFD41395D" box="[629,671,1015,1041]" pageId="129" pageNumber="130" type="male">1 ♂</specimenCount>
“Yunnan
<quantity id="4CEC9B156BE0FF44AC11FB9CFC40395D" box="[783,926,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.5" metricValueMin="1.25" pageId="129" pageNumber="130" unit="m" value="1375.0" valueMax="1500.0" valueMin="1250.0">
<elevation id="0039D1C36BE0FF44AC11FB9CFC40395D" box="[783,926,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.5" metricValueMin="1.25" pageId="129" pageNumber="130" unit="m" value="1375.0" valueMax="1500.0" valueMin="1250.0">1250-1500m</elevation>
</quantity>
<geoCoordinate id="EE2050376BE0FF44ACBBFB9DFC26395D" box="[933,1016,1015,1041]" degrees="23.14" direction="north" orientation="latitude" pageId="129" pageNumber="130" precision="555" value="23.14">23.14N</geoCoordinate>
<geoCoordinate id="EE2050376BE0FF44AB1EFB9DFB83395D" box="[1024,1117,1015,1041]" degrees="104.34" direction="east" orientation="longitude" pageId="129" pageNumber="130" precision="555" value="104.34">104.34E</geoCoordinate>
<collectorName id="26E153266BE0FF44AB7BFB9DFB75395D" box="[1125,1195,1015,1041]" pageId="129" pageNumber="130">Xinjie</collectorName>
<date id="FFAA10306BE0FF44ABACFB9DFAFC395D" box="[1202,1314,1015,1041]" pageId="129" pageNumber="130" value="1994-06-24">
<collectingDate id="EFEEE9D86BE0FF44ABACFB9DFAFC395D" box="[1202,1314,1015,1041]" pageId="129" pageNumber="130" value="1994-06-24">24.6.1994</collectingDate>
</date>
<collectorName id="26E153266BE0FF44AA36FB9DFA43395D" box="[1320,1437,1015,1041]" pageId="129" pageNumber="130">Vit Kuban</collectorName>
leg.” (
<collectionCode id="ED05AE356BE0FF44AFC2FC71FEDD3979" box="[220,259,1051,1077]" pageId="129" pageNumber="130">CP</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B7C3CAD6BE0FF44AE09FC71FE563915" ID-GBIF-Occurrence="3034556282" collectingDate="1994-06-24" collectingDateMax="1996-05-19" collectingDateMin="1994-06-24" collectionCode="CP" collectorName="Pacholátko &amp; Dembicky" country="Vietnam" elevation="1375" latitude="23.14" location="Hanoi" longLatPrecision="754" longitude="104.34" pageId="129" pageNumber="130" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi" typeStatus="paratype">
1 @ “N
<collectingCountry id="F30376606BE0FF44AE6AFC71FE0D3979" box="[372,467,1051,1077]" name="Vietnam" pageId="129" pageNumber="130">Vietnam</collectingCountry>
<geoCoordinate id="EE2050376BE0FF44AEC4FC71FDE83979" box="[474,566,1051,1077]" pageId="129" pageNumber="130">21.27’N</geoCoordinate>
<geoCoordinate id="EE2050376BE0FF44AD20FC70FD423979" box="[574,668,1050,1077]" degrees="105.39" direction="east" orientation="longitude" pageId="129" pageNumber="130" precision="555" value="105.39">105.39E</geoCoordinate>
<quantity id="4CEC9B156BE0FF44ADBDFC71FD3D3979" box="[675,739,1051,1077]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="129" pageNumber="130" unit="km" value="70.0">70km</quantity>
NW
<collectingRegion id="49D0F8126BE0FF44AC3EFC71FCB93979" box="[800,871,1051,1077]" country="Vietnam" name="Ha Noi" pageId="129" pageNumber="130">Hanoi</collectingRegion>
,
<collectorName id="26E153266BE0FF44AC6CFC71FC073979" box="[882,985,1051,1077]" pageId="129" pageNumber="130">Tam Dao</collectorName>
<date id="FFAA10306BE0FF44ACFEFC71FBB13979" box="[992,1135,1051,1077]" pageId="129" pageNumber="130" value="1996-05-09" valueMax="1996-05-19" valueMin="1996-05-09">
<collectingDate id="EFEEE9D86BE0FF44ACFEFC71FBB13979" box="[992,1135,1051,1077]" pageId="129" pageNumber="130" value="1996-05-09" valueMax="1996-05-19" valueMin="1996-05-09">9.-19.v.1996</collectingDate>
</date>
, 900-
<quantity id="4CEC9B156BE0FF44ABB3FC71FB223979" box="[1197,1276,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="129" pageNumber="130" unit="m" value="1200.0">1200m</quantity>
Pacholátko &amp; Dembicky leg.” (
<collectionCode id="ED05AE356BE0FF44AE46FC55FEA13915" box="[344,383,1087,1113]" pageId="129" pageNumber="130">CP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE0FF47AFD9FC08FD793FC9" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130" type="description">
<paragraph id="8BAB36F06BE0FF44AFD9FC08FD533989" blockId="129.[151,1437,798,2013]" pageId="129" pageNumber="130">
<emphasis id="B960EAE26BE0FF44AFD9FC08FE873930" bold="true" box="[199,345,1122,1148]" pageId="129" pageNumber="130">Description.</emphasis>
Length:
<quantity id="4CEC9B156BE0FF44AEA5FC08FDCC3931" box="[443,530,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="129" pageNumber="130" unit="mm" value="5.9">5.9 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BE0FF44ADCCFC09FCF73931" box="[722,809,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="129" pageNumber="130" unit="mm" value="3.8">3.8 mm</quantity>
, width:
<quantity id="4CEC9B156BE0FF44AC9EFC09FC063931" box="[896,984,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="129" pageNumber="130" unit="mm" value="3.9">3.9 mm</quantity>
. Body oval, black to dark reddish brown, club of antenna yellowish, parts of pronotum and head with greenish shine, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06BE0FF44AFD9FCA5FE233B45" blockId="129.[151,1437,798,2013]" pageId="129" pageNumber="130">
Labroclypeus wide and trapezoidal, shiny, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, medially angled; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons with fine, sparse punctures and a few single erect setae beside eyes. Eyes very small, ratio diameter/ interocular width: 0.38. Antenna with nine or ten antennomeres; club with three antennomeres and straight, 1.1 times as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126BE0FF44ABC9FDA1FAE838A9" box="[1239,1334,1483,1509]" country="Myanmar" name="Chin" pageId="129" pageNumber="130">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BE0FF44AFD9FE79FD0B3BF1" blockId="129.[151,1437,798,2013]" pageId="129" pageNumber="130">Pronotum transverse, widest at base, lateral margins evenly convex, distinctly convergent anteriorly, anterior angles moderately produced and rectangular, posterior angles blunt, weakly rounded at tip; anterior margin convex, with fine and complete marginal line, base without marginal line; surface finely and densely punctate, glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures, smooth at middle.</paragraph>
<paragraph id="8BAB36F06BE0FF44AFD9FEADFBFD3A02" blockId="129.[151,1437,798,2013]" pageId="129" pageNumber="130">Elytra widest at middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with only a few short, setae on odd intervals; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra not membraneous, without visible rim of microtrichomes (at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06BE0FF47AFD9FF3DFE623D99" blockId="129.[151,1437,798,2013]" lastBlockId="130.[151,1437,151,969]" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few robust setae along apical margin.</paragraph>
<paragraph id="8BAB36F06BE3FF47AFD9F8B5FAC03F2D" blockId="130.[151,1437,151,969]" pageId="130" pageNumber="131">Legs wide and short, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, without serrated line behind anterior margin, setae of anterior row very short, posterior margin smooth and widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.45, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short robust setae basally; lateral face longitudinally convex, sparsely punctate, glabrous; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06BE3FF47AFD9FA01FD793FC9" blockId="130.[151,1437,151,969]" box="[199,679,619,645]" pageId="130" pageNumber="131">
Aedeagus:
<figureCitation id="132F2A756BE3FF47AE5AFA01FE4A3FC9" box="[324,404,619,645]" captionStart="FIGURE 28" captionStartId="131.[151,250,1909,1933]" captionTargetBox="[197,1399,191,1873]" captionTargetId="figure-20@131.[188,1399,181,1884]" captionTargetPageId="131" captionText="FIGURE 28. A–D: Maladera pseudofuscipes Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–I: M. snizeki Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. fangana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; F: aedeagus, dorsal view; B, G, K: parameres, dorsal view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496380" httpUri="https://zenodo.org/record/4496380/files/figure.png" pageId="130" pageNumber="131">Fig. 28</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BE3FF47AD23FA01FD7A3FC9" box="[573,676,619,645]" captionStart="FIGURE 28" captionStartId="131.[151,250,1909,1933]" captionTargetBox="[197,1399,191,1873]" captionTargetId="figure-20@131.[188,1399,181,1884]" captionTargetPageId="131" captionText="FIGURE 28. A–D: Maladera pseudofuscipes Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–I: M. snizeki Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. fangana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, J: aedeagus, left side lateral view; C, H, L: aedeagus, right side lateral view; F: aedeagus, dorsal view; B, G, K: parameres, dorsal view; D, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496380" httpUri="https://zenodo.org/record/4496380/files/figure.png" pageId="130" pageNumber="131">Fig. 28D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE3FF47AFD9FAE4FBAE3FBD" pageId="130" pageNumber="131" type="diagnosis">
<paragraph id="8BAB36F06BE3FF47AFD9FAE4FBAE3FBD" blockId="130.[151,1437,151,969]" pageId="130" pageNumber="131">
<emphasis id="B960EAE26BE3FF47AFD9FAE4FE9C3FE4" bold="true" box="[199,322,654,680]" pageId="130" pageNumber="131">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BE3FF47AE57FAE5FDB43FE5" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[329,618,655,681]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="pseudofuscipes">
<emphasis id="B960EAE26BE3FF47AE57FAE5FDB43FE5" box="[329,618,655,681]" italics="true" pageId="130" pageNumber="131">Maladera pseudofuscipes</emphasis>
</taxonomicName>
is very closely related to
<taxonomicName id="4C144D736BE3FF47AC8EFAE5FBCE3FE5" authorityName="Moser" authorityYear="1915" box="[912,1040,655,681]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="fuscipes">
<emphasis id="B960EAE26BE3FF47AC8EFAE5FBCE3FE5" box="[912,1040,655,681]" italics="true" pageId="130" pageNumber="131">M. fuscipes</emphasis>
</taxonomicName>
, it differs from the latter in having the antennal club slightly shorter in the male as well as by the shape of parameres: the dorsobasal lobe of the right paramere is divided into two longer and sharp processes which diverge from each other.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE3FF47AFD9FA91FA893E75" pageId="130" pageNumber="131" type="etymology">
<paragraph id="8BAB36F06BE3FF47AFD9FA91FA893E75" blockId="130.[151,1437,151,969]" pageId="130" pageNumber="131">
<emphasis id="B960EAE26BE3FF47AFD9FA91FE903E59" bold="true" box="[199,334,763,789]" pageId="130" pageNumber="131">Etymology.</emphasis>
The name of the new species (noun in apposition) is derived from the combined Gree prefix
<emphasis id="B960EAE26BE3FF47AF89FB75FF383E75" box="[151,230,799,825]" italics="true" pageId="130" pageNumber="131">pseudo</emphasis>
- (false) and the species name ‘
<taxonomicName id="4C144D736BE3FF47AD5DFB75FD433E75" authorityName="Moser" authorityYear="1915" box="[579,669,799,825]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="fuscipes">
<emphasis id="B960EAE26BE3FF47AD5DFB75FD433E75" box="[579,669,799,825]" italics="true" pageId="130" pageNumber="131">fuscipes</emphasis>
</taxonomicName>
’, with reference to the similarity of the species to
<taxonomicName id="4C144D736BE3FF47ABCFFB75FA8E3E75" authorityName="Moser" authorityYear="1915" box="[1233,1360,799,825]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="fuscipes">
<emphasis id="B960EAE26BE3FF47ABCFFB75FA8E3E75" box="[1233,1360,799,825]" italics="true" pageId="130" pageNumber="131">M. fuscipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE3FF47AFD9FB28FE6A3EE9" pageId="130" pageNumber="131" type="description">
<paragraph id="8BAB36F06BE3FF47AFD9FB28FE6A3EE9" blockId="130.[151,1437,151,969]" pageId="130" pageNumber="131">
<emphasis id="B960EAE26BE3FF47AFD9FB28FEE13E10" bold="true" box="[199,319,834,860]" pageId="130" pageNumber="131">Variation.</emphasis>
Length:
<quantity id="4CEC9B156BE3FF47AEBAFB28FE233E11" box="[420,509,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="130" pageNumber="131" unit="mm" value="5.9">5.9 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BE3FF47ADDAFB29FCC23E11" box="[708,796,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="130" pageNumber="131" unit="mm" value="3.8">3.8 mm</quantity>
, width:
<quantity id="4CEC9B156BE3FF47AC69FB29FC0E3E11" box="[887,976,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="130" pageNumber="131" unit="mm" value="3.9">3.9 mm</quantity>
. Female: eyes little smaller than in male, ratio of diameter/interocular width: 0.34; antennal club with three antennomeres, as long as remaining antennomeres combined; pygidium flat.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BE3FF47AFD9FBC4FD103E85" box="[199,718,942,969]" pageId="130" pageNumber="131" type="distribution">
<paragraph id="8BAB36F06BE3FF47AFD9FBC4FD103E85" blockId="130.[151,1437,151,969]" box="[199,718,942,969]" pageId="130" pageNumber="131">
<emphasis id="B960EAE26BE3FF47AFD9FBC4FEBF3E84" bold="true" box="[199,353,942,968]" pageId="130" pageNumber="131">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BE3FF47AEC8FBC5FDE83E85" box="[470,566,943,969]" captionStart="FIGURE 100" captionStartId="351.[151,250,1973,1997]" captionTargetBox="[206,1379,1125,1944]" captionTargetId="figure-48@351.[202,1385,1119,1950]" captionTargetPageId="351" captionText="FIGURE 100. Distribution of the Maladera sikkimensis group: Maladera exima (Arrow, 1946), M. fangana sp. n., M. fuscipes (Moser, 1915), M. pseudoexima sp. n., M. pseudofuscipes sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496586" httpUri="https://zenodo.org/record/4496586/files/figure.png" pageId="130" pageNumber="131">Fig. 100</figureCitation>
) and
<tableCitation id="C696034B6BE3FF47AD6AFBC5FD163E85" box="[628,712,943,969]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="130" pageNumber="131">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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