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<document id="6084AFDA444DFD11E68865D683E17BC5" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BCDFF6BAF89FF28FD103BF1" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera kalawensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="175" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="173" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="950688E2DFE28D7B9540DE27F06E8AC9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="23C0C107D86486D9D205543C7A1B803A">
<mods:title id="B36B5F8A0176638C26771E2A5BE4C096">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="D48E1828B7C1F90E07D490CD517327F2" type="personal">
<mods:role id="F0B1576095E8B66491D076309070D6F4">
<mods:roleTerm id="D1C8EC7F2E1D1331E0280CD7E8719C8B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FA960484575124403ECD312812115C5">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="DDD615CBED4F05F4F9875217BEB20539" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="331EC115486F7CDAF7952EB419B75FA5" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="ABEDB5CC22A4BDBB8A07DFC364FACFC8" type="personal">
<mods:role id="37460DF2C31081B8302C8CBD10F5FF9E">
<mods:roleTerm id="55BCAF588A78AC159F96C11479F2C96C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0EFC4700022971B6103D32F330A8F61">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="D952539EE92E2E0437AB18328D785CA1" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="57573E2DE77B73E8A6AD3FE951071F08" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="142F167FE6724804F3757E86B7833D07" type="personal">
<mods:role id="DDC1EB2B4D522FFC374A7C0742720751">
<mods:roleTerm id="F0925B1787D9B0D16D7A5B6FEF970A4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1B304C5F76047B3EEB88E2DA5E00F6A">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="B9F6A962C7CA63F875AA925B71D6B307" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="92F72A4974D9DBBE9BC62CAE4DB2CF36" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="CDC6DE0C05C6352AB8E72943BE9FA0B7" type="personal">
<mods:role id="2A3D9836263AC6313353309F85DC771D">
<mods:roleTerm id="1D4025594CFB1D5498FFC88F01BDF5A1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0383EB1B11EB5DE04E632AC69FC1665B">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="3592D245AE333A753A1C3557EE3565C9" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="31289F2462B100FB982C5B74712838D9" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="C32862B02A0DBB410AA727105B9E73ED" type="personal">
<mods:role id="7CB2BD3EACF9BFEE110C8E40B3413584">
<mods:roleTerm id="C52D41C3E8952F814AA0199432111B7E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A4A66E857F3C1854B839AA56EBE462C">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="E644196BBB29F63A5768FC9B71B1309F" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="13F231A2C784395608118A82E200E1E4" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="AF6EC40025B3C29AC103E18762BBFF29">text</mods:typeOfResource>
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<mods:titleInfo id="8A2C2F096C3870A36DC808A43B3AF1B4">
<mods:title id="E19F5E86DCF968399794F085596A010E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C6D1C02E61BDA29F518AEEC1C742499F">
<mods:date id="CB5855BE433F23CA2202B785EF7BA9A0">2021</mods:date>
<mods:detail id="1C070D43F57BE0567D343C34E0C43126" type="pubDate">
<mods:number id="68CF8C81D88E5B3E3DF069F7D40AADE4">2021-02-03</mods:number>
</mods:detail>
<mods:detail id="5487F87FBB6A310C8BD6B63CC3F7B06C" type="volume">
<mods:number id="D49E017C306AC786CDBBC39DCD546105">4922</mods:number>
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<mods:detail id="200E81D2ED246939B22B93CC8D554E9E" type="issue">
<mods:number id="AC178C7F8017EC62EB26ED6397FF7E8D">1</mods:number>
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<mods:classification id="6F5736C353E5ECCA5545EB4A3A785B65">journal article</mods:classification>
<mods:identifier id="EE5D2FC90CBD1FACC20E082584DA26F1" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="CCECB6E80E08E6A877620357E17EB131" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="6F48564F92CAF2C7A411A4BD1E8B0FDA" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="268692ED639CA198C9D6B44EA82EB6EF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A3B05D67645330A5A70A63FA4E81AA0D" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="C0D40214763A0DC4F9FA1E02CAA0A33C" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
</mods:mods>
<treatment id="03BD87E66BCDFF6BAF89FF28FD103BF1" ID-DOI="http://doi.org/10.5281/zenodo.4534636" ID-GBIF-Taxon="177323578" ID-Zenodo-Dep="4534636" LSID="urn:lsid:plazi:treatment:03BD87E66BCDFF6BAF89FF28FD103BF1" httpUri="http://treatment.plazi.org/id/03BD87E66BCDFF6BAF89FF28FD103BF1" lastPageId="174" lastPageNumber="175" pageId="172" pageNumber="173">
<subSubSection id="C30E657B6BCDFF69AF89FF28FCD63A11" box="[151,776,1858,1885]" pageId="172" pageNumber="173" type="nomenclature">
<paragraph id="8BAB36F06BCDFF69AF89FF28FCD63A11" blockId="172.[151,776,1858,1921]" box="[151,776,1858,1885]" pageId="172" pageNumber="173">
<heading id="D0E3819C6BCDFF69AF89FF28FCD63A11" bold="true" box="[151,776,1858,1885]" fontSize="11" level="2" pageId="172" pageNumber="173" reason="4">
<emphasis id="B960EAE26BCDFF69AF89FF28FCD63A11" bold="true" box="[151,776,1858,1885]" pageId="172" pageNumber="173">
<taxonomicName id="4C144D736BCDFF69AF89FF28FD673A10" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,697,1858,1885]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="kalawensis" status="sp. nov.">
<emphasis id="B960EAE26BCDFF69AF89FF28FE463A11" bold="true" box="[151,408,1858,1885]" italics="true" pageId="172" pageNumber="173">Maladera kalawensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCDFF69ADD8FF29FCD63A11" box="[710,776,1859,1885]" pageId="172" pageNumber="173" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCDFF69AF89FF0DFEA33ACD" box="[151,381,1895,1921]" pageId="172" pageNumber="173" type="description">
<paragraph id="8BAB36F06BCDFF69AF89FF0DFEA33ACD" blockId="172.[151,776,1858,1921]" box="[151,381,1895,1921]" pageId="172" pageNumber="173">
<figureCitation id="132F2A756BCDFF69AF89FF0DFED03ACD" box="[151,270,1895,1921]" captionStart="FIGURE 39" captionStartId="173.[151,250,1933,1957]" captionTargetBox="[234,1351,285,1897]" captionTargetId="figure-61@173.[226,1361,273,1909]" captionTargetPageId="173" captionText="FIGURE 39. A–D: Maladera kalawensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. liaochengensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. paradetersa Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496410" httpUri="https://zenodo.org/record/4496410/files/figure.png" pageId="172" pageNumber="173">Figures 39</figureCitation>
A–D, 107
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCDFF68AF89FFC5FE113DB5" lastPageId="173" lastPageNumber="174" pageId="172" pageNumber="173" type="materials_examined">
<paragraph id="8BAB36F06BCDFF68AF89FFC5FE113DB5" blockId="172.[151,1436,1966,2029]" lastBlockId="173.[151,1436,150,249]" lastPageId="173" lastPageNumber="174" pageId="172" pageNumber="173">
<emphasis id="B960EAE26BCDFF69AF89FFC5FE1D3A84" bold="true" box="[151,451,1966,1993]" pageId="172" pageNumber="173">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BCDFF69AED2FFC5FE403AA1" ID-GBIF-Occurrence="3034556063" collectingDate="1997-06-05" collectionCode="ZFMK" collectorName="J. Rejsek" location="Taunggyi" pageId="172" pageNumber="173" specimenCount="1" specimenCount-male="1" stateProvince="Shan" typeStatus="holotype">
<typeStatus id="54AF88526BCDFF69AED2FFC5FDE73A85" box="[460,569,1967,1993]" pageId="172" pageNumber="173" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796BCDFF69AD5BFFC5FD843A85" box="[581,602,1967,1993]" pageId="172" pageNumber="173" type="male">♂</specimenCount>
“Burma (Myanmar) SW
<collectingRegion id="49D0F8126BCDFF69AC9EFFC5FC2A3A85" box="[896,1012,1967,1993]" country="Myanmar" name="Shan" pageId="172" pageNumber="173">Shan state</collectingRegion>
Kalaw
<quantity id="4CEC9B156BCDFF69AB51FFC5FB463A85" box="[1103,1176,1967,1993]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="172" pageNumber="173" unit="km" value="70.0">70 km</quantity>
W
<location id="8ECB602B6BCDFF69ABDDFFC5FAF23A85" LSID="urn:lsid:plazi:treatment:03BD87E66BCDFF6BAF89FF28FD103BF1:8ECB602B6BCDFF69ABDDFFC5FAF23A85" box="[1219,1324,1967,1993]" name="Taunggyi" pageId="172" pageNumber="173" stateProvince="Shan">Taunggyi</location>
<collectorName id="26E153266BCDFF69AA2BFFC5FA423A85" box="[1333,1436,1967,1993]" pageId="172" pageNumber="173">J. Rejsek</collectorName>
14.-
<date id="FFAA10306BCDFF69AFA3FFB8FEF63AA1" box="[189,296,2002,2029]" pageId="172" pageNumber="173" value="1997-06-05">
<collectingDate id="EFEEE9D86BCDFF69AFA3FFB8FEF63AA1" box="[189,296,2002,2029]" pageId="172" pageNumber="173" value="1997-06-05">5.6.1997</collectingDate>
</date>
” (
<collectionCode id="ED05AE356BCDFF69AE5CFFB9FE483AA1" box="[322,406,2003,2029]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="172" pageNumber="173" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BCDFF68AEB4FFB9FCDB3DFD" ID-GBIF-Occurrence="3034556160" collectingDate="1997-06-10" collectingDateMax="1997-06-11" collectingDateMin="1997-06-10" collectionCode="CP" collectorName="J. Kalab" country="Myanmar" lastPageId="173" lastPageNumber="174" location="Taunggyi" pageId="172" pageNumber="173" specimenCount="1" specimenCount-male="1" stateProvince="Shan" typeStatus="paratype">
<typeStatus id="54AF88526BCDFF69AEB4FFB9FDC53AA1" box="[426,539,2003,2029]" pageId="172" pageNumber="173" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B960EAE26BCDFF69AD39FFB9FD763AA1" bold="true" box="[551,680,2003,2029]" pageId="172" pageNumber="173">
<collectingCountry id="F30376606BCDFF69AD39FFB9FD7D3AA1" box="[551,675,2003,2029]" name="Myanmar" pageId="172" pageNumber="173">Myanmar</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BCDFF69ADAEFFB9FD053AA1" box="[688,731,2003,2029]" pageId="172" pageNumber="173" type="male">1 ♂</specimenCount>
“Burma: SW
<collectingRegion id="49D0F8126BCDFF69AC60FFB9FC283AA1" box="[894,1014,2003,2029]" country="Myanmar" name="Shan" pageId="172" pageNumber="173">Shan State</collectingRegion>
Kalaw,
<date id="FFAA10306BCDFF69AB4AFFB9FB253AA1" box="[1108,1275,2003,2029]" pageId="172" pageNumber="173" value="1997-06-10" valueMax="1997-06-11" valueMin="1997-06-10">
<collectingDate id="EFEEE9D86BCDFF69AB4AFFB9FB253AA1" box="[1108,1275,2003,2029]" pageId="172" pageNumber="173" value="1997-06-10" valueMax="1997-06-11" valueMin="1997-06-10">10.-11.vi.1997</collectingDate>
</date>
; (
<quantity id="4CEC9B156BCDFF69AA11FFB9FA8E3AA1" box="[1295,1360,2003,2029]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="172" pageNumber="173" unit="km" value="70.0">70km</quantity>
WSW
<location id="8ECB602B6BCCFF68AF89F8FDFED83DFE" LSID="urn:lsid:plazi:treatment:03BD87E66BCDFF6BAF89FF28FD103BF1:8ECB602B6BCCFF68AF89F8FDFED83DFE" box="[151,262,151,178]" country="Myanmar" name="Taunggyi" pageId="173" pageNumber="174" stateProvince="Shan">Taunggyi</location>
);
<collectorName id="26E153266BCCFF68AE07F8FDFEA83DFD" box="[281,374,151,177]" pageId="173" pageNumber="174">J. Kalab</collectorName>
leg./ 274
<taxonomicName id="4C144D736BCCFF68AEF9F8FDFDE03DFD" box="[487,574,151,177]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356BCCFF68ADCBF8FDFD223DFD" box="[725,764,151,177]" pageId="173" pageNumber="174">CP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BCCFF68AC0EF8FCFD843D99" ID-GBIF-Occurrence="3034555461" collectingDate="1993-05-01" collectingDateMax="1993-05-08" collectingDateMin="1993-05-01" collectionCode="CP" collectorName="Dembicky" country="Thailand" elevation="1800" location="Thai" pageId="173" pageNumber="174" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B960EAE26BCCFF68AC0EF8FCFC583DFC" bold="true" box="[784,902,150,176]" pageId="173" pageNumber="174">
<collectingCountry id="F30376606BCCFF68AC0EF8FCFC5F3DFC" box="[784,897,150,176]" name="Thailand" pageId="173" pageNumber="174">Thailand</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BCCFF68AC90F8FDFC663DFD" box="[910,952,151,177]" pageId="173" pageNumber="174" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6BCCFF68ACD4F8FDFBDE3DFD" LSID="urn:lsid:plazi:treatment:03BD87E66BCDFF6BAF89FF28FD103BF1:8ECB602B6BCCFF68ACD4F8FDFBDE3DFD" box="[970,1024,151,177]" country="Thailand" name="Thai" pageId="173" pageNumber="174">Thai</location>
<date id="FFAA10306BCCFF68AB19F8FDFB5D3DFD" box="[1031,1155,151,177]" pageId="173" pageNumber="174" value="1993-05-01" valueMax="1993-05-08" valueMin="1993-05-01">
<collectingDate id="EFEEE9D86BCCFF68AB19F8FDFB5D3DFD" box="[1031,1155,151,177]" pageId="173" pageNumber="174" value="1993-05-01" valueMax="1993-05-08" valueMin="1993-05-01">1-8.V.1993</collectingDate>
</date>
Soppong Pai
<quantity id="4CEC9B156BCCFF68AA3FF8FDFAAE3DFD" box="[1313,1392,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="173" pageNumber="174" unit="m" value="1800.0">
<elevation id="0039D1C36BCCFF68AA3FF8FDFAAE3DFD" box="[1313,1392,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="173" pageNumber="174" unit="m" value="1800.0">1800m</elevation>
</quantity>
Pacholatko &amp;
<collectorName id="26E153266BCCFF68AE04F8D1FE513D99" box="[282,399,187,213]" pageId="173" pageNumber="174">Dembicky</collectorName>
leg./ TS 22” (
<collectionCode id="ED05AE356BCCFF68AD34F8D1FD8F3D99" box="[554,593,187,213]" pageId="173" pageNumber="174">CP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BCCFF68AD7AF8D0FE153DB5" ID-GBIF-Occurrence="3034555320" collectingDate="1980-04-23" collectionCode="IZAS, ZFMK" collectorName="Li Hongxing" country="China" location="Yunnan" pageId="173" pageNumber="174" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B960EAE26BCCFF68AD7AF8D0FD683D98" bold="true" box="[612,694,186,212]" pageId="173" pageNumber="174">
<collectingCountry id="F30376606BCCFF68AD7AF8D0FD6F3D98" box="[612,689,186,212]" name="China" pageId="173" pageNumber="174">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BCCFF68ADA3F8D1FD233D99" box="[701,765,187,213]" pageId="173" pageNumber="174" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D12FD796BCCFF68AC17F8D1FCEC3D99" box="[777,818,187,213]" pageId="173" pageNumber="174" type="female">1 ♀</specimenCount>
“[China] Yingjiang,
<collectingRegion id="49D0F8126BCCFF68AB04F8D1FBAD3D99" box="[1050,1139,187,213]" country="China" name="Yunnan" pageId="173" pageNumber="174">Yunnan</collectingRegion>
,
<date id="FFAA10306BCCFF68AB63F8D1FB223D99" box="[1149,1276,187,213]" pageId="173" pageNumber="174" value="1980-04-23">
<collectingDate id="EFEEE9D86BCCFF68AB63F8D1FB223D99" box="[1149,1276,187,213]" pageId="173" pageNumber="174" value="1980-04-23">
23.IV.
<quantity id="4CEC9B156BCCFF68ABDFF8D1FB223D99" box="[1217,1276,187,213]" metricMagnitude="6" metricUnit="m" metricValue="1.9808" pageId="173" pageNumber="174" unit="m" value="1980800.0">1980</quantity>
</collectingDate>
</date>
, 800m, leg.
<collectorName id="26E153266BCCFF68AA9DF8D1FED43DB5" pageId="173" pageNumber="174">Li Hongxing</collectorName>
” (
<collectionCode id="ED05AE356BCCFF68AE3DF8B5FEBB3DB5" box="[291,357,223,249]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="173" pageNumber="174">IZAS</collectionCode>
,
<collectionCode id="ED05AE356BCCFF68AE6EF8B5FE1D3DB5" box="[368,451,223,249]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="173" pageNumber="174" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6B66786BCCFF68AF89FFE7FA443AA1" ID-DOI="http://doi.org/10.5281/zenodo.4496410" ID-Zenodo-Dep="4496410" httpUri="https://zenodo.org/record/4496410/files/figure.png" pageId="173" pageNumber="174" startId="173.[151,250,1933,1957]" targetBox="[234,1351,285,1897]" targetPageId="173">
<paragraph id="8BAB36F06BCCFF68AF89FFE7FA443AA1" blockId="173.[151,1437,1933,2029]" pageId="173" pageNumber="174">
<emphasis id="B960EAE26BCCFF68AF89FFE7FE803AE9" bold="true" box="[151,350,1933,1957]" pageId="173" pageNumber="174">FIGURE 39. A–D:</emphasis>
<taxonomicName id="4C144D736BCCFF68AE7BFFE7FCF63AE9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[357,808,1933,1957]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="kalawensis" status="sp. nov.">
<emphasis id="B960EAE26BCCFF68AE7BFFE7FDE13AE9" box="[357,575,1933,1957]" italics="true" pageId="173" pageNumber="174">Maladera kalawensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCCFF68AC2CFFE7FCB73AE8" box="[818,873,1933,1956]" pageId="173" pageNumber="174" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26BCCFF68ACF8FFE7FBFE3AE9" bold="true" box="[998,1056,1933,1957]" pageId="173" pageNumber="174">E–H:</emphasis>
<taxonomicName id="4C144D736BCCFF68AB39FFE4FF633A85" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="liaochengensis" status="sp. nov.">
<emphasis id="B960EAE26BCCFF68AB39FFE4FB3E3AE9" box="[1063,1248,1933,1957]" italics="true" pageId="173" pageNumber="174">M. liaochengensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCCFF68AFD9FFDBFF233A84" box="[199,253,1969,1992]" pageId="173" pageNumber="174" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26BCCFF68AE67FFDBFE763A85" bold="true" box="[377,424,1969,1993]" pageId="173" pageNumber="174">I–L:</emphasis>
<taxonomicName id="4C144D736BCCFF68AEB1FFD8FCEC3A85" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[431,818,1969,1993]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="paradetersa" status="sp. nov.">
<emphasis id="B960EAE26BCCFF68AEB1FFD8FD943A85" box="[431,586,1969,1993]" italics="true" pageId="173" pageNumber="174">M. paradetersa</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCCFF68AC25FFDBFCAF3A84" box="[827,881,1969,1992]" pageId="173" pageNumber="174" rank="species">sp. n.</taxonomicNameLabel>
(holotype).
<emphasis id="B960EAE26BCCFF68ACF3FFDBFBE43A85" bold="true" box="[1005,1082,1969,1993]" pageId="173" pageNumber="174">A, E, I:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26BCCFF68AA9AFFDBFF0D3AA1" bold="true" pageId="173" pageNumber="174">C, G, K:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26BCCFF68AD33FFBFFDA43AA1" bold="true" box="[557,634,2005,2029]" pageId="173" pageNumber="174">B, F, J:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26BCCFF68AC66FFBFFC0E3AA1" bold="true" box="[888,976,2005,2029]" pageId="173" pageNumber="174">D, H, L:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<subSubSection id="C30E657B6BCFFF6BAFD9F8FCFD79388D" pageId="174" pageNumber="175" type="description">
<paragraph id="8BAB36F06BCFFF6BAFD9F8FCFBDF3D99" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">
<emphasis id="B960EAE26BCFFF6BAFD9F8FCFE873DFC" bold="true" box="[199,345,150,176]" pageId="174" pageNumber="175">Description.</emphasis>
Length:
<quantity id="4CEC9B156BCFFF6BAEDAF8FCFDC13DFD" box="[452,543,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="174" pageNumber="175" unit="mm" value="5.9">5.9 mm</quantity>
, length of elytra: 4.0 mm, width:
<quantity id="4CEC9B156BCFFF6BACB5F8FDFBD93DFD" box="[939,1031,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="174" pageNumber="175" unit="mm" value="3.2">3.2 mm</quantity>
. Body oblong-oval, dark yellowish brown, antenna yellow, shiny, labroclypeus shiny, dorsal face nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9F8B5FC9A3CB9" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">
Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent anteriorly, anterior angles moderately rounded, anterior margin weakly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex medially, finely, densely punctate, glabrous; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, twice as wide as long; ocular canthus short and narrow (1/4 of ocular diameter), finely densely punctate, terminal seta lacking. Frons with dense and fine punctures, glabrous. Eyes small, ratio diameter/ interocular width: 0.65. Antenna with ten antennomeres, club composed of three antennomeres and straight, distinctly shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126BCFFF6BAE0AF9B1FEAD3CB9" box="[276,371,475,501]" country="Myanmar" name="Chin" pageId="174" pageNumber="175">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9F995FABC3FE5" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">Pronotum transverse, widest at base, lateral margins in basal half nealy straight and subparallel, in anterior half evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin straight, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures.</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9FAD9FD0E3E76" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">Elytra widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures and with minute setae in punctures glabrous; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9FB29FCCC3EA1" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">Ventral surface shiny, finely and densely punctate, metasternum including disc sparsely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Pygidium strongly convex, shiny, finely and densely punctate, without smooth midline, glabrous.</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9FB9DFB6038D1" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">Legs short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, strongly widened at ventral apex, dorsal posterior margin smooth, neither serrate, with a few single long setae. Metatibia short and moderately wide, widest at middle, ratio of width/length: 1/2.67, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae; lateral face longitudinally convex, except a few superficial punctures on sides and base smooth and glabrous; ventral margin finely serrate, with five equidistant robust setae; medial face smooth and glabrous; apex finely serrate, weakly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06BCFFF6BAFD9FDCDFD79388D" blockId="174.[151,1437,150,1725]" box="[199,679,1447,1473]" pageId="174" pageNumber="175">
Aedeagus:
<figureCitation id="132F2A756BCFFF6BAE5AFDCDFE4A388D" box="[324,404,1447,1473]" captionStart="FIGURE 39" captionStartId="173.[151,250,1933,1957]" captionTargetBox="[234,1351,285,1897]" captionTargetId="figure-61@173.[226,1361,273,1909]" captionTargetPageId="173" captionText="FIGURE 39. A–D: Maladera kalawensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. liaochengensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. paradetersa Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496410" httpUri="https://zenodo.org/record/4496410/files/figure.png" pageId="174" pageNumber="175">Fig. 39</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BCFFF6BAD23FDCDFD7A388D" box="[573,676,1447,1473]" captionStart="FIGURE 39" captionStartId="173.[151,250,1933,1957]" captionTargetBox="[234,1351,285,1897]" captionTargetId="figure-61@173.[226,1361,273,1909]" captionTargetPageId="173" captionText="FIGURE 39. A–D: Maladera kalawensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. liaochengensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. paradetersa Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496410" httpUri="https://zenodo.org/record/4496410/files/figure.png" pageId="174" pageNumber="175">Fig. 39D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCFFF6BAFD9FDA0FA893B45" pageId="174" pageNumber="175" type="diagnosis">
<paragraph id="8BAB36F06BCFFF6BAFD9FDA0FA893B45" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">
<emphasis id="B960EAE26BCFFF6BAFD9FDA0FE9C38A8" bold="true" box="[199,322,1482,1508]" pageId="174" pageNumber="175">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BCFFF6BAE55FDA1FC9738A9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[331,841,1483,1509]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="kalawensis" status="sp. nov.">
<emphasis id="B960EAE26BCFFF6BAE55FDA1FDE338A9" box="[331,573,1483,1509]" italics="true" pageId="174" pageNumber="175">Maladera kalawensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCFFF6BAC48FDA1FC4A38A9" box="[854,916,1483,1509]" pageId="174" pageNumber="175" rank="species">sp. n.</taxonomicNameLabel>
differs from other species of the
<taxonomicName id="4C144D736BCFFF6BAA09FDA1FA4238A9" box="[1303,1436,1483,1509]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="detsersa">
<emphasis id="B960EAE26BCFFF6BAA09FDA1FA4238A9" box="[1303,1436,1483,1509]" italics="true" pageId="174" pageNumber="175">M. detsersa</emphasis>
</taxonomicName>
group possessing parameres as long or longer than phallobase, in having distinctly asymmetrical parameres.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCFFF6BAFD9FE79FA933B62" box="[199,1357,1555,1582]" pageId="174" pageNumber="175" type="etymology">
<paragraph id="8BAB36F06BCFFF6BAFD9FE79FA933B62" blockId="174.[151,1437,150,1725]" box="[199,1357,1555,1582]" pageId="174" pageNumber="175">
<emphasis id="B960EAE26BCFFF6BAFD9FE79FE903B61" bold="true" box="[199,334,1555,1581]" pageId="174" pageNumber="175">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54AF88526BCFFF6BADFAFE79FCCA3B61" box="[740,788,1555,1581]" pageId="174" pageNumber="175">type</typeStatus>
locality, Kalaw (adjective in nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCFFF6BAFD9FE5CFE963BD5" pageId="174" pageNumber="175" type="description">
<paragraph id="8BAB36F06BCFFF6BAFD9FE5CFE963BD5" blockId="174.[151,1437,150,1725]" pageId="174" pageNumber="175">
<emphasis id="B960EAE26BCFFF6BAFD9FE5CFEE13B1C" bold="true" box="[199,319,1590,1616]" pageId="174" pageNumber="175">Variation.</emphasis>
Length:
<quantity id="4CEC9B156BCFFF6BAEB6FE5CFDEA3B1D" box="[424,564,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.8" metricValueMin="5.8" pageId="174" pageNumber="175" unit="mm" value="6.3" valueMax="6.8" valueMin="5.8">5.8–6.8 mm</quantity>
, length of elytra: 4.0–
<quantity id="4CEC9B156BCFFF6BAC2AFE5CFC533B1D" box="[820,909,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="174" pageNumber="175" unit="mm" value="4.5">4.5 mm</quantity>
, width:
<quantity id="4CEC9B156BCFFF6BACF2FE5DFBA63B1D" box="[1004,1144,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="174" pageNumber="175" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
. Female: eyes as large as in male; antennal club with three antennomeres, little shorter than remaining antennomeres combined; pygidium weakly convex.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCFFF6BAFD9FEC8FD103BF1" box="[199,718,1698,1725]" pageId="174" pageNumber="175" type="distribution">
<paragraph id="8BAB36F06BCFFF6BAFD9FEC8FD103BF1" blockId="174.[151,1437,150,1725]" box="[199,718,1698,1725]" pageId="174" pageNumber="175">
<emphasis id="B960EAE26BCFFF6BAFD9FEC8FEBF3BF0" bold="true" box="[199,353,1698,1724]" pageId="174" pageNumber="175">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BCFFF6BAEC8FEC9FDE83BF1" box="[470,566,1699,1725]" captionStart="FIGURE 107" captionStartId="355.[151,250,1033,1057]" captionTargetBox="[211,1378,188,1001]" captionTargetId="figure-20@355.[202,1385,181,1009]" captionTargetPageId="355" captionText="FIGURE 107. Distribution of the Maladera detersa group: Maladera detersa (Erichson, 1834), M. kalawensis sp. n., M. liaochengensis sp. n., M. paradetersa sp. n. (not precisely localized collection site indicated by a “?”)." figureDoi="http://doi.org/10.5281/zenodo.4496608" httpUri="https://zenodo.org/record/4496608/files/figure.png" pageId="174" pageNumber="175">Fig. 107</figureCitation>
) and
<tableCitation id="C696034B6BCFFF6BAD6AFEC9FD163BF1" box="[628,712,1699,1725]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="174" pageNumber="175">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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