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<document id="AE5C6D9DCEB4C026D43D25E86C539566" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BC1FF65AF89FA00FA423851" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera nitens Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens 2021" docType="key" docVersion="9" lastPageNumber="161" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="161" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="C99A936EEEDF9B2B1FDC4E03E99E0BC9">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="6E73169DAB12C03FB77A64904E6BAF6C">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="5F198BC776A98D2568E4139F625D919B">Liu, Wan-Gang</mods:namePart>
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<mods:nameIdentifier id="0666902CA08D39E395BD48EB938AD644" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="2765DFFBDA60ED65E78C4509C7E7340B">Bai, Ming</mods:namePart>
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<mods:nameIdentifier id="E98E428A65EA875810BB0A4DA748C790" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:name id="EF13EC9AEF30EF127D58471FBF2A12A7" type="personal">
<mods:role id="9E05F8013DA69F774548B45FFE96946A">
<mods:roleTerm id="B27FA1D143946B0087D5B16EC7278626">Author</mods:roleTerm>
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<mods:namePart id="27F4F5F2994D44C16BC6E45315DE5278">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="3255A4A338C613326E10FADD85EA61D5" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="846DEA58F3F6A8862A35BD5C2F74350B" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="08AD24DDDB6130723800AF95BF2C36DB">Ahrens, Dirk</mods:namePart>
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<mods:date id="C034F1C9C22DAB9B2E789DA1F9A846A9">2021</mods:date>
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<mods:number id="E739B5D266A7BA44364FF09431050948">4922</mods:number>
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<mods:identifier id="6D58256FC8B9CF47A3C3F37C420665DA" type="CLB-Dataset">8257</mods:identifier>
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<treatment id="03BD87E66BC1FF65AF89FA00FA423851" ID-DOI="http://doi.org/10.5281/zenodo.4534611" ID-GBIF-Taxon="177323571" ID-Zenodo-Dep="4534611" LSID="urn:lsid:plazi:treatment:03BD87E66BC1FF65AF89FA00FA423851" httpUri="http://treatment.plazi.org/id/03BD87E66BC1FF65AF89FA00FA423851" lastPageNumber="161" pageId="160" pageNumber="161">
<subSubSection id="C30E657B6BC1FF65AF89FA00FDBE3FC9" box="[151,608,618,645]" pageId="160" pageNumber="161" type="nomenclature">
<paragraph id="8BAB36F06BC1FF65AF89FA00FDBE3FC9" blockId="160.[151,608,618,645]" box="[151,608,618,645]" pageId="160" pageNumber="161">
<heading id="D0E3819C6BC1FF65AF89FA00FDBE3FC9" bold="true" box="[151,608,618,645]" fontSize="11" level="2" pageId="160" pageNumber="161" reason="4">
<emphasis id="B960EAE26BC1FF65AF89FA00FDBE3FC9" bold="true" box="[151,608,618,645]" pageId="160" pageNumber="161">
<taxonomicName id="4C144D736BC1FF65AF89FA00FDD03FC8" authority="(Moser, 1915)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[151,526,618,645]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B960EAE26BC1FF65AF89FA00FE833FC9" bold="true" box="[151,349,618,645]" italics="true" pageId="160" pageNumber="161">Maladera nitens</emphasis>
(Moser, 1915)
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BC1FF65AF89FAD9FA423851" pageId="160" pageNumber="161" type="key">
<paragraph id="8BAB36F06BC1FF65AF89FAD9FB723F83" blockId="160.[151,1196,690,719]" box="[151,1196,690,719]" pageId="160" pageNumber="161">
<emphasis id="B960EAE26BC1FF65AF89FAD9FB723F83" bold="true" box="[151,1196,690,719]" pageId="160" pageNumber="161">
Key to species of the
<taxonomicName id="4C144D736BC1FF65AE83FAD8FCCA3F80" authority="(Moser, 1915)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[413,788,690,717]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B960EAE26BC1FF65AE83FAD8FDBC3F81" bold="true" box="[413,610,690,717]" italics="true" pageId="160" pageNumber="161">Maladera nitens</emphasis>
(Moser, 1915)
</taxonomicName>
group occurring in
<collectingCountry id="F30376606BC1FF65AB10FAD8FB853F80" box="[1038,1115,690,716]" name="China" pageId="160" pageNumber="161">China</collectingCountry>
(³³):
</emphasis>
</paragraph>
<key id="FF8561C16BC1FF65AF89FA92FA423851" pageId="160" pageNumber="161">
<keyStep id="30E02E556BC1FF65AF89FA92FA423E29" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FA92FA433E66" blockId="160.[151,1437,760,1310]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FA92FA433E66" pageId="160" pageNumber="161">
1 Dorsal surface dull, body size larger (&gt;
<quantity id="4CEC9B156BC1FF65AD7BFA92FD713E41" box="[613,687,760,781]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="160" pageNumber="161" unit="mm" value="10.0">10 mm</quantity>
). Anterior margin of metafemur without serrate line behind anterior margin. Metatibia without serrate line.........................................
<taxonomicName id="4C144D736BC1FF65ACF7FB7FFABE3E67" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[1001,1376,789,811]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACF7FB7FFB5C3E67" box="[1001,1154,789,811]" italics="true" pageId="160" pageNumber="161">Maladera songi</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FB7FFA433E66" box="[1385,1437,789,810]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FB58FA423E29" blockId="160.[151,1437,760,1310]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FB58FA423E29" pageId="160" pageNumber="161">
1’ Dorsal surface shiny, body size smaller (&lt;
<quantity id="4CEC9B156BC1FF65AD64FB58FD1E3E04" box="[634,704,818,840]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="160" pageNumber="161" unit="mm" value="10.0">10 mm</quantity>
). Anterior margin of metafemur with a serrate line immediately behind anterior margin. Metatibia with serrate line in basal half.......................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556BC1FF65AF89FB07FA423ED3" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FB07FA423ECE" blockId="160.[151,1437,760,1310]" box="[151,1436,877,899]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FB07FA423ECE" box="[151,1436,877,899]" pageId="160" pageNumber="161">2 Parameres equally in length............................................................................. 4</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FBE0FA423ED3" blockId="160.[151,1437,760,1310]" box="[151,1436,906,928]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FBE0FA423ED3" box="[151,1436,906,928]" pageId="160" pageNumber="161">2’ Right paramere distinctly shorter than the left one............................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556BC1FF65AF89FBC2FA433E96" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FBC2FA433EF1" blockId="160.[151,1437,760,1310]" box="[151,1437,936,958]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FBC2FA433EF1" box="[151,1437,936,958]" pageId="160" pageNumber="161">
3 Right paramere narrowly emarginate before apex.....................
<taxonomicName id="4C144D736BC1FF65ACA0FBC2FABE3EF2" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[958,1376,936,958]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="allonitens" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACA0FBC2FB5C3EF1" box="[958,1154,936,957]" italics="true" pageId="160" pageNumber="161">Maladera allonitens</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FBC2FA433EF1" box="[1385,1437,936,957]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FBAFFA433E96" blockId="160.[151,1437,760,1310]" box="[151,1437,965,987]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FBAFFA433E96" box="[151,1437,965,987]" pageId="160" pageNumber="161">
3’ Right paramere deeply and widely emarginate before apex..............
<taxonomicName id="4C144D736BC1FF65ACA3FBAFFABE3E97" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[957,1376,965,987]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="daanensis" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACA3FBAFFB5C3E96" box="[957,1154,965,986]" italics="true" pageId="160" pageNumber="161">Maladera daanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FBAFFA433E96" box="[1385,1437,965,986]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556BC1FF65AF89FB88FA423959" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FB88FA423EB4" blockId="160.[151,1437,760,1310]" box="[151,1436,994,1016]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FB88FA423EB4" box="[151,1436,994,1016]" pageId="160" pageNumber="161">4 Right paramere in basal half convexly widened.............................................................. 6</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FC6AFA423959" blockId="160.[151,1437,760,1310]" box="[151,1436,1023,1046]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FC6AFA423959" box="[151,1436,1023,1046]" pageId="160" pageNumber="161">4’ Right paramere in basal half not particularly widened......................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556BC1FF65AF89FC77FA4339C6" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FC77FA43391C" blockId="160.[151,1437,760,1310]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FC77FA43391C" pageId="160" pageNumber="161">
5 Parameres bent laterally (to the right) at apex. Phallobase strongly narrowed towards apex, half as wide as maximum width (dorsal view)..................................................
<taxonomicName id="4C144D736BC1FF65ACA8FC50FABE391C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[950,1376,1082,1104]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="paranitens" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACA8FC50FB5C391C" box="[950,1154,1082,1104]" italics="true" pageId="160" pageNumber="161">Maladera paranitens</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FC51FA43391C" box="[1385,1437,1083,1104]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FC3DFA4339C6" blockId="160.[151,1437,760,1310]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FC3DFA4339C6" pageId="160" pageNumber="161">
5’ Parameres straight. Phallobase moderately narrowed towards apex, ¾ as wide as maximum width (dorsal view)......................................................................
<taxonomicName id="4C144D736BC1FF65ACBEFC1FFABE39C7" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[928,1376,1141,1163]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="pseudonitens" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACBEFC1FFB5F39C6" box="[928,1153,1141,1162]" italics="true" pageId="160" pageNumber="161">Maladera pseudonitens</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FC1FFA4339C6" box="[1385,1437,1141,1162]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556BC1FF65AF89FCF8FA433989" pageId="160" pageNumber="161">
<paragraph id="8BAB36F06BC1FF65AF89FCF8FA4239E4" blockId="160.[151,1437,760,1310]" box="[151,1436,1170,1192]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FCF8FA4239E4" box="[151,1436,1170,1192]" pageId="160" pageNumber="161">6 Right paramere laterally convex in basal half............................................................... 7</keyLead>
</paragraph>
<paragraph id="8BAB36F06BC1FF65AF89FCDAFA433989" blockId="160.[151,1437,760,1310]" box="[151,1437,1200,1222]" pageId="160" pageNumber="161">
<keyLead id="30E595C56BC1FF65AF89FCDAFA433989" box="[151,1437,1200,1222]" pageId="160" pageNumber="161">
6’ Right paramere laterally concave in basal half.....................
<taxonomicName id="4C144D736BC1FF65AC8FFCDAFABE398A" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[913,1376,1200,1222]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="jiucailingensis" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65AC8FFCDAFB5C3989" box="[913,1154,1200,1221]" italics="true" pageId="160" pageNumber="161">Maladera jiucailingensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC1FF65AA77FCDAFA433989" box="[1385,1437,1200,1221]" pageId="160" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
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<paragraph id="8BAB36F06BC1FF65AF89FCA7FA43384C" blockId="160.[151,1437,760,1310]" pageId="160" pageNumber="161">
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7 Right paramere abruptly and strongly bent in apical half (dorsal view,
<figureCitation id="132F2A756BC1FF65AC9AFCA7FC0539AF" box="[900,987,1229,1251]" captionStart="FIGURE 38" captionStartId="171.[151,250,1943,1967]" captionTargetBox="[222,1371,191,1903]" captionTargetId="figure-20@171.[205,1382,181,1920]" captionTargetPageId="171" captionText="FIGURE 38. A–D: Maladera rufonitida Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. songi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. detersa (Erichson, 1834) (Vietnam: DaNang). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496408" httpUri="https://zenodo.org/record/4496408/files/figure.png" pageId="160" pageNumber="161">Fig. 38B</figureCitation>
)................................................................................................
<taxonomicName id="4C144D736BC1FF65ACA3FC80FABE384C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[957,1376,1258,1280]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="rufonitida" status="sp. nov.">
<emphasis id="B960EAE26BC1FF65ACA3FC80FB5C39B3" box="[957,1154,1258,1279]" italics="true" pageId="160" pageNumber="161">Maladera rufonitida</emphasis>
Ahrens, Fabrizi &amp; Liu
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,
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7’ Right paramere weakly bent in apical half (dorsal view,
<figureCitation id="132F2A756BC1FF65ADE2FD62FC8A3852" box="[764,852,1288,1310]" captionStart="FIGURE 36" captionStartId="162.[151,250,1944,1968]" captionTargetBox="[202,1389,196,1909]" captionTargetId="figure-19@162.[190,1398,181,1921]" captionTargetPageId="162" captionText="FIGURE 36. A–D: Maladera nitens (Moser, 1915) (China: Guadun, Fujian); E–H: M. allonitens Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. daanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496402" httpUri="https://zenodo.org/record/4496402/files/figure.png" pageId="160" pageNumber="161">Fig. 36B</figureCitation>
).......................
<taxonomicName id="4C144D736BC1FF65AB6EFD62FA423851" authority="(Moser, 1915)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1136,1436,1287,1309]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="160" pageNumber="161" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B960EAE26BC1FF65AB6EFD62FAD03851" box="[1136,1294,1288,1309]" italics="true" pageId="160" pageNumber="161">Maladera nitens</emphasis>
(Moser, 1915)
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