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<mods:title id="4A374D7370DFB80491CF67521150EAEA">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<taxonomicName id="4C144D736BBCFF18AF89FD9CFD943B5C" authority="(Frey, 1972)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[151,586,1526,1553]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="221" pageNumber="222" phylum="Arthropoda" rank="species" species="brunnescens" status="comb. nov.">
<emphasis id="B960EAE26BBCFF18AF89FD9CFE733B5D" bold="true" box="[151,429,1526,1553]" italics="true" pageId="221" pageNumber="222">Maladera brunnescens</emphasis>
(Frey, 1972)
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<taxonomicNameLabel id="A25357996BBCFF18AD4FFD9DFD623B5D" box="[593,700,1527,1553]" pageId="221" pageNumber="222" rank="species">comb. n.</taxonomicNameLabel>
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<figureCitation id="132F2A756BBCFF18AF89FE71FED33B79" box="[151,269,1562,1589]" captionStart="FIGURE 51" captionStartId="222.[151,250,1374,1398]" captionTargetBox="[159,1424,191,1339]" captionTargetId="figure-20@222.[151,1436,181,1351]" captionTargetPageId="222" captionText="FIGURE 51. A–D: Maladera brunnescens (Frey, 1972) (holotype); E–H: M. parabrunnescens Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496446" httpUri="https://zenodo.org/record/4496446/files/figure.png" pageId="221" pageNumber="222">Figures 51</figureCitation>
A–D, 116
</paragraph>
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<treatmentCitation id="0AB510E16BBCFF18AF89FE30FDCF3B3E" author="Frey, G." box="[151,529,1626,1650]" page="166" pageId="221" pageNumber="222" year="1972">
<taxonomicName id="4C144D736BBCFF18AF89FE30FDCF3B3E" authority="Frey, 1972 a: 166" authorityName="G. Frey" authorityPageNumber="166" authorityYear="1972" box="[151,529,1626,1650]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="221" pageNumber="222" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B960EAE26BBCFF18AF89FE30FE833B3E" box="[151,349,1626,1650]" italics="true" pageId="221" pageNumber="222">Serica brunnescens</emphasis>
<bibRefCitation id="EF854B016BBCFF18AE7DFE30FDCF3B3E" author="Frey, G." box="[355,529,1626,1650]" pageId="221" pageNumber="222" pagination="162 - 177" refId="ref332211" refString="Frey, G. (1972 a) Neue Sericinen der Klapperich-Ausbeute aus Fukien des Alexander Koenig Museum in Bonn (Col., Scarab., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey, 23, 162 - 177." type="journal article" year="1972">Frey, 1972a: 166</bibRefCitation>
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;
<treatmentCitation id="0AB510E16BBCFF18AD04FE30FD013B3E" author="Ahrens, D." box="[538,735,1626,1650]" page="245" pageId="221" pageNumber="222" year="2006">
<bibRefCitation id="EF854B016BBCFF18AD04FE30FD013B3E" author="Ahrens, D." box="[538,735,1626,1650]" pageId="221" pageNumber="222" refId="ref329946" refString="Ahrens, D. (2006 b) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book" year="2006">Ahrens 2006b: 245</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16BBCFF18ADF7FE30FC8C3B3E" author="Ahrens, D." box="[745,850,1626,1650]" page="36" pageId="221" pageNumber="222" year="2007">
<bibRefCitation id="EF854B016BBCFF18ADF7FE30FC8C3B3E" author="Ahrens, D." box="[745,850,1626,1650]" pageId="221" pageNumber="222" pagination="1 - 51" refId="ref330155" refString="Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1" type="journal article" year="2007">2007b: 36</bibRefCitation>
</treatmentCitation>
.
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<emphasis id="B960EAE26BBCFF18AF89FEF1FE653BF8" bold="true" box="[151,443,1690,1717]" pageId="221" pageNumber="222">Type material examined.</emphasis>
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:
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“Kuatun (
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<collectorName id="26E153266BBCFF18AB1AFEF1FB6B3BF9" box="[1028,1205,1691,1717]" pageId="221" pageNumber="222">L. J. Klapperich</collectorName>
<date id="FFAA10306BBCFF18ABA5FEF1FAF13BF9" box="[1211,1327,1691,1717]" pageId="221" pageNumber="222" value="1938-11-06">
<collectingDate id="EFEEE9D86BBCFF18ABA5FEF1FAF13BF9" box="[1211,1327,1691,1717]" pageId="221" pageNumber="222" value="1938-11-06">11.6. 1938</collectingDate>
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(Fukien)/ Type
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<emphasis id="B960EAE26BBCFF18AFCBFED5FE703B95" box="[213,430,1727,1753]" italics="true" pageId="221" pageNumber="222">Serica brunnescens</emphasis>
G. Frey 1972
</taxonomicName>
” (
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)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BBCFF18ADDBFED5FBD63BB1" ID-GBIF-Occurrence="3034556624" collectingDate="1938-06-17" collectionCode="ZMFK" elevation="2300" latitude="27.4" longLatPrecision="742" longitude="-117.4" pageId="221" pageNumber="222" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54AF88526BBCFF18ADDBFED5FCE83B95" box="[709,822,1727,1753]" pageId="221" pageNumber="222" type="paratype">Paratypes</typeStatus>
:
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“Kuatun (
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<geoCoordinate id="EE2050376BBCFF18AB21FED5FB4A3B95" box="[1087,1172,1727,1753]" degrees="27, 40" direction="north" orientation="latitude" pageId="221" pageNumber="222" precision="555" value="27.4">27,40 n</geoCoordinate>
.Br.
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.L. J. Klapperich
<date id="FFAA10306BBCFF18AFFBFE89FE833BB1" box="[229,349,1763,1789]" pageId="221" pageNumber="222" value="1938-06-17">
<collectingDate id="EFEEE9D86BBCFF18AFFBFE89FE833BB1" box="[229,349,1763,1789]" pageId="221" pageNumber="222" value="1938-06-17">17.6. 1938</collectingDate>
</date>
(Fukien)/ Type
<taxonomicName id="4C144D736BBCFF18AD0BFE89FC4C3BB1" authority="G. Frey 1972" authorityName="G. Frey" authorityYear="1972" box="[533,914,1763,1789]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="221" pageNumber="222" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B960EAE26BBCFF18AD0BFE89FD2F3BB1" box="[533,753,1763,1789]" italics="true" pageId="221" pageNumber="222">Serica brunnescens</emphasis>
G. Frey 1972
</taxonomicName>
” (
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,
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<specimenCount id="9D12FD796BBCFF18AB0AFE89FBE13BB1" box="[1044,1087,1763,1789]" pageId="221" pageNumber="222" type="female">1 ♀</specimenCount>
“Kuatun (
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<geoCoordinate id="EE2050376BBCFF18AA01FE89FAA83BB1" box="[1311,1398,1763,1789]" degrees="27, 40" direction="north" orientation="latitude" pageId="221" pageNumber="222" precision="555" value="27.4">27,40 n</geoCoordinate>
.Br.
<geoCoordinate id="EE2050376BBCFF18AF89FF6DFF243A6D" box="[151,250,1799,1825]" degrees="117, 40" direction="west" orientation="longitude" pageId="221" pageNumber="222" precision="555" value="-117.4">117,40 ö</geoCoordinate>
.
<collectorName id="26E153266BBCFF18AFE0FF6DFE6D3A6D" box="[254,435,1799,1825]" pageId="221" pageNumber="222">L. J. Klapperich</collectorName>
<date id="FFAA10306BBCFF18AEA5FF6DFDFA3A6D" box="[443,548,1799,1825]" pageId="221" pageNumber="222" value="1938-06-08">
<collectingDate id="EFEEE9D86BBCFF18AEA5FF6DFDFA3A6D" box="[443,548,1799,1825]" pageId="221" pageNumber="222" value="1938-06-08">8.6. 1938</collectingDate>
</date>
(Fukien)/ Type
<taxonomicName id="4C144D736BBCFF18ADC6FF6DFB8E3A6D" authority="G. Frey 1972" authorityName="G. Frey" authorityYear="1972" box="[728,1104,1799,1825]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="221" pageNumber="222" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B960EAE26BBCFF18ADC6FF6DFC6C3A6D" box="[728,946,1799,1825]" italics="true" pageId="221" pageNumber="222">Serica brunnescens</emphasis>
G. Frey 1972
</taxonomicName>
” (
<collectionCode id="ED05AE356BBCFF18AB77FF6DFB633A6D" box="[1129,1213,1799,1825]" pageId="221" pageNumber="222">ZMFK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36F06BBCFF18AFD9FF40FE5C3AB5" blockId="221.[151,1437,1690,2041]" pageId="221" pageNumber="222">
<emphasis id="B960EAE26BBCFF18AFD9FF40FDE53A08" bold="true" box="[199,571,1834,1860]" pageId="221" pageNumber="222">Additional material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BBCFF18AD5BFF41FBAB3A25" ID-GBIF-Occurrence="3034555694" collectingDate="1938-11-06" collectionCode="NMPC" collectorName="L. J. Klapperich &amp; Coll. V. Balthasar National Museum" country="Czech Republic" elevation="2300" latitude="27.4" location="Kuatun" longLatPrecision="742" longitude="-117.4" pageId="221" pageNumber="222" specimenCount="41" stateProvince="Praha">
<specimenCount id="9D12FD796BBCFF18AD5BFF41FDA13A09" box="[581,639,1835,1861]" pageId="221" pageNumber="222" type="generic">1 ex.</specimenCount>
“
<location id="8ECB602B6BBCFF18AD8DFF41FD363A09" LSID="urn:lsid:plazi:treatment:03BD87E66BBCFF1AAF89FD9CFD133931:8ECB602B6BBCFF18AD8DFF41FD363A09" box="[659,744,1835,1861]" country="Czech Republic" latitude="27.4" longLatPrecision="742" longitude="-117.4" name="Kuatun" pageId="221" pageNumber="222" stateProvince="Praha">Kuatun</location>
(
<quantity id="4CEC9B156BBCFF18ADE4FF41FC8B3A09" box="[762,853,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="221" pageNumber="222" unit="m" value="2300.0">
<elevation id="0039D1C36BBCFF18ADE4FF41FC8B3A09" box="[762,853,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="221" pageNumber="222" unit="m" value="2300.0">2300 m</elevation>
</quantity>
)
<geoCoordinate id="EE2050376BBCFF18AC76FF41FC1F3A09" box="[872,961,1835,1861]" degrees="27, 40" direction="north" orientation="latitude" pageId="221" pageNumber="222" precision="555" value="27.4">
27,
<specimenCount id="9D12FD796BBCFF18AC97FF41FC1F3A09" box="[905,961,1835,1861]" pageId="221" pageNumber="222" type="generic">40 n</specimenCount>
</geoCoordinate>
.Br.
<geoCoordinate id="EE2050376BBCFF18ACECFF41FB863A09" box="[1010,1112,1835,1861]" degrees="117, 40" direction="west" orientation="longitude" pageId="221" pageNumber="222" precision="555" value="-117.4">117,40 ö</geoCoordinate>
.
<collectorName id="26E153266BBCFF18AB42FF41FAC63A09" box="[1116,1304,1835,1861]" pageId="221" pageNumber="222">L. J. Klapperich</collectorName>
<date id="FFAA10306BBCFF18AA3DFF41FA423A09" box="[1315,1436,1835,1861]" pageId="221" pageNumber="222" value="1938-11-06">
<collectingDate id="EFEEE9D86BBCFF18AA3DFF41FA423A09" box="[1315,1436,1835,1861]" pageId="221" pageNumber="222" value="1938-11-06">11.6. 1938</collectingDate>
</date>
(Fukien)/ ex.
<collectorName id="26E153266BBCFF18AE28FF25FD043A25" box="[310,730,1871,1897]" pageId="221" pageNumber="222">Coll. V. Balthasar National Museum</collectorName>
<collectingRegion id="49D0F8126BBCFF18ADFBFF25FCE83A25" box="[741,822,1871,1897]" country="Czech Republic" name="Praha" pageId="221" pageNumber="222">Prague</collectingRegion>
,
<collectingCountry id="F30376606BBCFF18AC5AFF25FC223A25" box="[836,1020,1871,1897]" name="Czech Republic" pageId="221" pageNumber="222">Czech Republic</collectingCountry>
” (
<collectionCode id="ED05AE356BBCFF18AB06FF25FBB33A25" box="[1048,1133,1871,1897]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="221" pageNumber="222">NMPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BBCFF18AB9AFF25FEA83AFD" ID-GBIF-Occurrence="3034555333" collectingDate="1938-06-08" collectionCode="NMPC" collectorName="L. J. Klapperich &amp; Coll. V. Balthasar National Museum" country="Czech Republic" elevation="2300" latitude="27.4" location="Kuatun" longLatPrecision="742" longitude="-117.4" pageId="221" pageNumber="222" specimenCount="41" stateProvince="Praha">
<specimenCount id="9D12FD796BBCFF18AB9AFF25FB603A25" box="[1156,1214,1871,1897]" pageId="221" pageNumber="222" type="generic">1 ex.</specimenCount>
“
<location id="8ECB602B6BBCFF18ABCCFF25FAF93A25" LSID="urn:lsid:plazi:treatment:03BD87E66BBCFF1AAF89FD9CFD133931:8ECB602B6BBCFF18ABCCFF25FAF93A25" box="[1234,1319,1871,1897]" country="Czech Republic" latitude="27.4" longLatPrecision="742" longitude="-117.4" name="Kuatun" pageId="221" pageNumber="222" stateProvince="Praha">Kuatun</location>
(
<quantity id="4CEC9B156BBCFF18AA27FF25FA4A3A25" box="[1337,1428,1871,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="221" pageNumber="222" unit="m" value="2300.0">
<elevation id="0039D1C36BBCFF18AA27FF25FA4A3A25" box="[1337,1428,1871,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="221" pageNumber="222" unit="m" value="2300.0">2300 m</elevation>
</quantity>
)
<geoCoordinate id="EE2050376BBCFF18AF89FF19FF313AC1" box="[151,239,1907,1933]" degrees="27, 40" direction="north" orientation="latitude" pageId="221" pageNumber="222" precision="555" value="27.4">
27,
<specimenCount id="9D12FD796BBCFF18AFA6FF19FF313AC1" box="[184,239,1907,1933]" pageId="221" pageNumber="222" type="generic">40 n</specimenCount>
</geoCoordinate>
.Br.
<geoCoordinate id="EE2050376BBCFF18AE00FF19FE5C3AC1" box="[286,386,1907,1933]" degrees="117, 40" direction="west" orientation="longitude" pageId="221" pageNumber="222" precision="555" value="-117.4">117,40 ö</geoCoordinate>
.
<collectorName id="26E153266BBCFF18AE98FF19FDE03AC1" box="[390,574,1907,1933]" pageId="221" pageNumber="222">L. J. Klapperich</collectorName>
<date id="FFAA10306BBCFF18AD59FF19FD6F3AC1" box="[583,689,1907,1933]" pageId="221" pageNumber="222" value="1938-06-08">
<collectingDate id="EFEEE9D86BBCFF18AD59FF19FD6F3AC1" box="[583,689,1907,1933]" pageId="221" pageNumber="222" value="1938-06-08">8.6. 1938</collectingDate>
</date>
(Fukien)/ ex.
<collectorName id="26E153266BBCFF18AC4AFF19FB2F3AC1" box="[852,1265,1907,1933]" pageId="221" pageNumber="222">Coll. V. Balthasar National Museum</collectorName>
<collectingRegion id="49D0F8126BBCFF18ABE7FF19FA943AC1" box="[1273,1354,1907,1933]" country="Czech Republic" name="Praha" pageId="221" pageNumber="222">Prague</collectingRegion>
,
<collectingCountry id="F30376606BBCFF18AA48FF19FF213AFD" name="Czech Republic" pageId="221" pageNumber="222">Czech Republic</collectingCountry>
” (
<collectionCode id="ED05AE356BBCFF18AE06FFFDFEB33AFD" box="[280,365,1943,1969]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="221" pageNumber="222">NMPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BBCFF18AE9CFFFDFEA03A99" ID-GBIF-Occurrence="3034556496" collectionCode="ZFMK" collectorName="Jaroslav Turna" country="China" elevation="1350" latitude="27.016666" location="Emei Feng" longLatPrecision="1239" longitude="117.066666" pageId="221" pageNumber="222" specimenCount="43" stateProvince="Fujian">
<specimenCount id="9D12FD796BBCFF18AE9CFFFDFE193AFD" box="[386,455,1943,1969]" pageId="221" pageNumber="222" type="generic">43 ex.</specimenCount>
“
<collectingCountry id="F30376606BBCFF18AEC7FFFDFDFF3AFD" box="[473,545,1943,1969]" name="China" pageId="221" pageNumber="222">China</collectingCountry>
, W
<collectingRegion id="49D0F8126BBCFF18AD50FFFDFD473AFD" box="[590,665,1943,1969]" country="China" name="Fujian" pageId="221" pageNumber="222">Fujian</collectingRegion>
, 3.-4.VI.
<location id="8ECB602B6BBCFF18AC1BFFFDFC5C3AFD" LSID="urn:lsid:plazi:treatment:03BD87E66BBCFF1AAF89FD9CFD133931:8ECB602B6BBCFF18AC1BFFFDFC5C3AFD" box="[773,898,1943,1969]" country="China" latitude="27.016666" longLatPrecision="1239" longitude="117.066666" name="Emei Feng" pageId="221" pageNumber="222" stateProvince="Fujian">Emei Feng</location>
,
<quantity id="4CEC9B156BBCFF18AC90FFFCFBC03AFD" box="[910,1054,1942,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="221" pageNumber="222" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">
<elevation id="0039D1C36BBCFF18AC90FFFCFBC03AFD" box="[910,1054,1942,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="221" pageNumber="222" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">1200-1500m</elevation>
</quantity>
<geoCoordinate id="EE2050376BBCFF18AB3BFFFDFB583AFD" box="[1061,1158,1943,1969]" degrees="27" direction="north" minutes="01" orientation="latitude" pageId="221" pageNumber="222" precision="925" value="27.016666">27°01‘N</geoCoordinate>
<geoCoordinate id="EE2050376BBCFF18AB90FFFDFB313AFD" box="[1166,1263,1943,1969]" degrees="117" direction="east" minutes="04" orientation="longitude" pageId="221" pageNumber="222" precision="925" value="117.066666">117°04E</geoCoordinate>
<collectorName id="26E153266BBCFF18ABE9FFFDFA423AFD" box="[1271,1436,1943,1969]" pageId="221" pageNumber="222">Jaroslav Turna</collectorName>
leg., 2008” (
<collectionCode id="ED05AE356BBCFF18AE3CFFD1FEA83A99" box="[290,374,1979,2005]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="221" pageNumber="222" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BBCFF18AE97FFD1FEA03AB5" ID-GBIF-Occurrence="3034556017" collectionCode="ZFMK" collectorName="Jaroslav Turna" country="China" elevation="1350" latitude="27.016666" location="Emei Feng" longLatPrecision="1239" longitude="117.066666" pageId="221" pageNumber="222" specimenCount="10" stateProvince="Fujian">
<specimenCount id="9D12FD796BBCFF18AE97FFD1FE103A99" box="[393,462,1979,2005]" pageId="221" pageNumber="222" type="generic">10 ex.</specimenCount>
“
<collectingCountry id="F30376606BBCFF18AEC1FFD1FDF93A99" box="[479,551,1979,2005]" name="China" pageId="221" pageNumber="222">China</collectingCountry>
, W
<collectingRegion id="49D0F8126BBCFF18AD4DFFD1FD403A99" box="[595,670,1979,2005]" country="China" name="Fujian" pageId="221" pageNumber="222">Fujian</collectingRegion>
, 1.-2.VI.
<location id="8ECB602B6BBCFF18AC16FFD1FC583A99" LSID="urn:lsid:plazi:treatment:03BD87E66BBCFF1AAF89FD9CFD133931:8ECB602B6BBCFF18AC16FFD1FC583A99" box="[776,902,1979,2005]" country="China" latitude="27.016666" longLatPrecision="1239" longitude="117.066666" name="Emei Feng" pageId="221" pageNumber="222" stateProvince="Fujian">Emei Feng</location>
,
<quantity id="4CEC9B156BBCFF18AC8FFFD0FBFE3A99" box="[913,1056,1978,2005]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="221" pageNumber="222" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">
<elevation id="0039D1C36BBCFF18AC8FFFD0FBFE3A99" box="[913,1056,1978,2005]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="221" pageNumber="222" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">1200-1500m</elevation>
</quantity>
<geoCoordinate id="EE2050376BBCFF18AB39FFD1FB563A99" box="[1063,1160,1979,2005]" degrees="27" direction="north" minutes="01" orientation="latitude" pageId="221" pageNumber="222" precision="925" value="27.016666">27°01‘N</geoCoordinate>
<geoCoordinate id="EE2050376BBCFF18AB91FFD1FB2F3A99" box="[1167,1265,1979,2005]" degrees="117" direction="east" minutes="04" orientation="longitude" pageId="221" pageNumber="222" precision="925" value="117.066666">117°04E</geoCoordinate>
<collectorName id="26E153266BBCFF18ABE6FFD1FA423A99" box="[1272,1436,1979,2005]" pageId="221" pageNumber="222">Jaroslav Turna</collectorName>
leg., 2007” (
<collectionCode id="ED05AE356BBCFF18AE3CFFB5FEA83AB5" box="[290,374,2015,2041]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="221" pageNumber="222" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6B66786BBFFF1BAF89FD34FD8538F3" ID-DOI="http://doi.org/10.5281/zenodo.4496446" ID-Zenodo-Dep="4496446" httpUri="https://zenodo.org/record/4496446/files/figure.png" pageId="222" pageNumber="223" startId="222.[151,250,1374,1398]" targetBox="[159,1424,191,1339]" targetPageId="222">
<paragraph id="8BAB36F06BBFFF1BAF89FD34FD8538F3" blockId="222.[151,1437,1374,1471]" pageId="222" pageNumber="223">
<emphasis id="B960EAE26BBFFF1BAF89FD34FEBE383B" bold="true" box="[151,352,1374,1399]" pageId="222" pageNumber="223">FIGURE 51. A–D:</emphasis>
<taxonomicName id="4C144D736BBFFF1BAE77FD35FD07383B" authority="(Frey, 1972)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[361,729,1375,1399]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="222" pageNumber="223" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B960EAE26BBFFF1BAE77FD35FD8D383B" box="[361,595,1375,1399]" italics="true" pageId="222" pageNumber="223">Maladera brunnescens</emphasis>
(Frey, 1972)
</taxonomicName>
(holotype);
<emphasis id="B960EAE26BBFFF1BAC47FD35FC4C383B" bold="true" box="[857,914,1375,1399]" pageId="222" pageNumber="223">E–H:</emphasis>
<taxonomicName id="4C144D736BBFFF1BAC84FD35FA84383B" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[922,1370,1375,1399]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="222" pageNumber="223" phylum="Arthropoda" rank="species" species="parabrunnescens" status="sp. nov.">
<emphasis id="B960EAE26BBFFF1BAC84FD35FBB5383B" box="[922,1131,1375,1399]" italics="true" pageId="222" pageNumber="223">M. parabrunnescens</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BBFFF1BAA7BFD35FA43383A" box="[1381,1437,1375,1398]" pageId="222" pageNumber="223" rank="species">sp. n.</taxonomicNameLabel>
(holotype).
<emphasis id="B960EAE26BBFFF1BAE10FDE8FE9838D7" bold="true" box="[270,326,1410,1435]" pageId="222" pageNumber="223">A, E:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26BBFFF1BAD8AFDE8FD1138D7" bold="true" box="[660,719,1410,1435]" pageId="222" pageNumber="223">C, G:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26BBFFF1BAB35FDE9FBBF38D7" bold="true" box="[1067,1121,1411,1435]" pageId="222" pageNumber="223">B, F:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26BBFFF1BAA7FFDE9FA4238D7" bold="true" box="[1377,1436,1411,1435]" pageId="222" pageNumber="223">D, H:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<subSubSection id="C30E657B6BBFFF1AAFD9FD86FB663915" lastPageId="223" lastPageNumber="224" pageId="222" pageNumber="223" type="description">
<paragraph id="8BAB36F06BBFFF1BAFD9FD86FCBB3B67" blockId="222.[151,1437,1516,2011]" pageId="222" pageNumber="223">
<emphasis id="B960EAE26BBFFF1BAFD9FD86FEAB3B4A" bold="true" box="[199,373,1516,1542]" pageId="222" pageNumber="223">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156BBFFF1BAEC2FD87FD9D3B4B" box="[476,579,1517,1543]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="222" pageNumber="223" unit="mm" value="10.1">10.1 mm</quantity>
, width: 6.0 mm, length of elytra: 7.0 mm. Body oblong, reddish brown, antenna yellow, dorsal surface shiny, except head almost glabrous.
</paragraph>
<paragraph id="8BAB36F06BBFFF1BAFD9FE5FFE463ADF" blockId="222.[151,1437,1516,2011]" pageId="222" pageNumber="223">
Labroclypeus subtrapezoidal, widest at base, lateral margins slightly curved and strongly convergent to feebly rounded anterior angles, lateral border and ocular canthus produce a distinctly blunt angle, only anterior margin weakly reflexed, anteriorly indistinctly but widely emarginate medially; surface distinctly convex medially and shiny, finely and moderately densely punctate, with numerous coarse punctures bearing fine and erect setae; frontoclypeal suture very indistinctly incised and medially weakly angled; smooth area in front of eye very small, as wide as first antennal joint and approximately 1.5times as wide as long; ocular canthus moderately long and moderately slender, densely finely punctate, with one terminal seta. Frons with fine, sparsely scattered punctures, with a few setae bent backwards. Eyes moderately small, ratio of diameter/ interocular width: 0.57. Antenna yellow, 10-segmented; club with three segments, in male as long as remaining segments together.
<collectingRegion id="49D0F8126BBFFF1BAB44FF3FFB673A23" box="[1114,1209,1877,1903]" country="Myanmar" name="Chin" pageId="222" pageNumber="223">Mentum</collectingRegion>
elevated, anteriorly flattened and glabrous.
</paragraph>
<paragraph id="8BAB36F06BBFFF1AAFD9FFF7FE1F3DB5" blockId="222.[151,1437,1516,2011]" lastBlockId="223.[151,1437,151,1150]" lastPageId="223" lastPageNumber="224" pageId="222" pageNumber="223">Pronotum narrow, widest at base, lateral margins weakly curved and strongly narrowed anteriorly, behind the middle feebly convexly emarginate, anterior angles strongly produced and sharp, anterior margin without marginal line and weakly produced medially, posterior angles sharp; surface moderately finely and densely punctate, punctures denser basally, glabrous; anterior and lateral border setaceous. Scutellum wide, triangular, with fine and very dense punctures, glabrous.</paragraph>
<paragraph id="8BAB36F06BBEFF1AAFD9F969FAD93CC5" blockId="223.[151,1437,151,1150]" pageId="223" pageNumber="224">Elytra moderately oblong, widest behind the middle, striae strongly impressed and finely and densely punctate, intervals distinctly convex, with fine and sparsely scattered punctures, mostly concentrated along striae, a few punctures with sparse fine, short, erect setae; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setaceous, apical border chitinous, covered with very short microtrichomes.</paragraph>
<paragraph id="8BAB36F06BBEFF1AAFD9F9F9FD613F2D" blockId="223.[151,1437,151,1150]" pageId="223" pageNumber="224">Ventral surface shiny, coarsely and densely punctate, almost glabrous, metacoxal plates laterally with a few long setae; each abdominal sternite with a transversal row of coarse punctures bearing short and fine setae between fine and dense punctation, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as sternite. Mesosternum between mesocoxae a little less wide than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.41. Pygidium moderately convex, finely and densely punctate, without smooth midline, surface apically dull and with numerous fine and long setae.</paragraph>
<paragraph id="8BAB36F06BBEFF1AAFD9FA01FC21395D" blockId="223.[151,1437,151,1150]" pageId="223" pageNumber="224">
Legs moderately slender; femora shiny, with two longitudinal rows of setae, between the rows only sparsely punctate; metafemur anteriorly sharply carinate, behind anterior margin without serrated line, posterior margin in apical half dorsally finely serrated, ventrally completely serrated, in basal half with a few fine setae. Metatibia moderately slender and moderately long, widest at apex, ratio width/ length: 1/ 3.33, dorsally moderately carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length, basally with two single setae with punctures having a serrated margin; lateral face longitudinally convex, with moderately dense, coarse and longitudinally impressed punctures and glabrous; ventral margin with three strong spines, of which the apical two are very distant, inside sparsely scattered, finely punctate, apex interiorly near tarsal articulation shallowly concavely emarginate. Tarsomeres distinctly and densely punctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a moderate longitudinal margin, segments one to four ventrally glabrous, first metatarsomere distinctly shorter than the two following segments combined and about one third longer than dorsal tibial spur. Protibia in
<typeStatus id="54AF88526BBEFF1AAC0DFB9DFCAB395D" box="[787,885,1015,1041]" pageId="223" pageNumber="224" type="holotype">holotype</typeStatus>
are lacking.
</paragraph>
<paragraph id="8BAB36F06BBEFF1AAFD9FC71FD793979" blockId="223.[151,1437,151,1150]" box="[199,679,1050,1077]" pageId="223" pageNumber="224">
Aedeagus:
<figureCitation id="132F2A756BBEFF1AAE5AFC71FE4D3979" box="[324,403,1050,1077]" captionStart="FIGURE 51" captionStartId="222.[151,250,1374,1398]" captionTargetBox="[159,1424,191,1339]" captionTargetId="figure-20@222.[151,1436,181,1351]" captionTargetPageId="222" captionText="FIGURE 51. A–D: Maladera brunnescens (Frey, 1972) (holotype); E–H: M. parabrunnescens Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496446" httpUri="https://zenodo.org/record/4496446/files/figure.png" pageId="223" pageNumber="224">Fig. 51</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BBEFF1AAD23FC71FD7A3979" box="[573,676,1050,1077]" captionStart="FIGURE 51" captionStartId="222.[151,250,1374,1398]" captionTargetBox="[159,1424,191,1339]" captionTargetId="figure-20@222.[151,1436,181,1351]" captionTargetPageId="222" captionText="FIGURE 51. A–D: Maladera brunnescens (Frey, 1972) (holotype); E–H: M. parabrunnescens Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496446" httpUri="https://zenodo.org/record/4496446/files/figure.png" pageId="223" pageNumber="224">Fig. 51D</figureCitation>
.
</paragraph>
<paragraph id="8BAB36F06BBEFF1AAFD9FC54FB663915" blockId="223.[151,1437,151,1150]" box="[199,1208,1086,1113]" pageId="223" pageNumber="224">
<emphasis id="B960EAE26BBEFF1AAFD9FC54FEE13914" bold="true" box="[199,319,1086,1112]" pageId="223" pageNumber="224">Variation.</emphasis>
Antennal club in female a little shorter than remaining antennal joints together.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BBEFF1AAFD9FC08FD133931" box="[199,717,1122,1150]" pageId="223" pageNumber="224" type="distribution">
<paragraph id="8BAB36F06BBEFF1AAFD9FC08FD133931" blockId="223.[151,1437,151,1150]" box="[199,717,1122,1150]" pageId="223" pageNumber="224">
<emphasis id="B960EAE26BBEFF1AAFD9FC08FEBF3930" bold="true" box="[199,353,1122,1148]" pageId="223" pageNumber="224">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BBEFF1AAEC8FC09FDEB3932" box="[470,565,1123,1150]" captionStart="FIGURE 116" captionStartId="359.[151,250,1952,1976]" captionTargetBox="[202,1383,1098,1922]" captionTargetId="figure-42@359.[196,1391,1091,1928]" captionTargetPageId="359" captionText="FIGURE 116. Distribution of the Maladera brunnescens group: Maladera brunnescens (Frey, 1972), M. filigraniforceps sp. n., M. flavipennis sp. n., M. parabrunnescens sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496632" httpUri="https://zenodo.org/record/4496632/files/figure.png" pageId="223" pageNumber="224">Fig. 116</figureCitation>
) and
<tableCitation id="C696034B6BBEFF1AAD6DFC09FD193931" box="[627,711,1123,1149]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="223" pageNumber="224">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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