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<document id="EC8CDAE16E1711CAA245DEC9D9427F59" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BAEFF15AF89FB28FD133979" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera pallida Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens 2021, comb.n." docType="treatment" docVersion="10" lastPageNumber="209" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="208" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="BF87DA6208966EC823DFC64B830A0AF8">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="44D6EA4B611916D664A55C29A5760914">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="541517B0B829381AF16C6A158CAEFDAD">Bai, Ming</mods:namePart>
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<mods:namePart id="B5A19986FE01788A92508A4F6BE5D758">Ahrens, Dirk</mods:namePart>
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<mods:date id="D5D60B8D734BE2DEF6647C0632A801F8">2021</mods:date>
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<treatment id="03BD87E66BAEFF15AF89FB28FD133979" ID-DOI="http://doi.org/10.5281/zenodo.4534697" ID-GBIF-Taxon="177323581" ID-Zenodo-Dep="4534697" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979" httpUri="http://treatment.plazi.org/id/03BD87E66BAEFF15AF89FB28FD133979" lastPageId="208" lastPageNumber="209" pageId="207" pageNumber="208">
<subSubSection id="C30E657B6BAEFF0AAF89FB28FD1F3E11" box="[151,705,834,861]" pageId="207" pageNumber="208" type="nomenclature">
<paragraph id="8BAB36F06BAEFF0AAF89FB28FD1F3E11" blockId="207.[151,705,834,897]" box="[151,705,834,861]" pageId="207" pageNumber="208">
<heading id="D0E3819C6BAEFF0AAF89FB28FD1F3E11" bold="true" box="[151,705,834,861]" fontSize="11" level="2" pageId="207" pageNumber="208" reason="4">
<emphasis id="B960EAE26BAEFF0AAF89FB28FD1F3E11" bold="true" box="[151,705,834,861]" pageId="207" pageNumber="208">
<taxonomicName id="4C144D736BAEFF0AAF89FB28FD883E10" authority="(Burmeister, 1855)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[151,598,834,861]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida" status="comb. nov.">
<emphasis id="B960EAE26BAEFF0AAF89FB28FEB93E11" bold="true" box="[151,359,834,861]" italics="true" pageId="207" pageNumber="208">Maladera pallida</emphasis>
(
<bibRefCitation id="EF854B016BAEFF0AAE66FB28FD903E10" author="Burmeister, H." box="[376,590,834,860]" pageId="207" pageNumber="208" refId="ref331559" refString="Burmeister, H. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. T. C. F. Enslin, Berlin, x + 569 pp." type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25357996BAEFF0AAD40FB29FD1F3E11" box="[606,705,835,861]" pageId="207" pageNumber="208" rank="species">comb.n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C30E657B6BAEFF0AAF89FB0DFEA93ECD" box="[151,375,871,897]" pageId="207" pageNumber="208" type="description">
<paragraph id="8BAB36F06BAEFF0AAF89FB0DFEA93ECD" blockId="207.[151,705,834,897]" box="[151,375,871,897]" pageId="207" pageNumber="208">
<figureCitation id="132F2A756BAEFF0AAF89FB0DFED03ECD" box="[151,270,871,897]" captionStart="FIGURE 47" captionStartId="204.[151,250,1944,1968]" captionTargetBox="[212,1375,197,1908]" captionTargetId="figure-19@204.[198,1389,181,1920]" captionTargetPageId="204" captionText="FIGURE 47. A–E: Maladera bansongchana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. peregoi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. pallida (Burmeister, 1855) (China). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496436" httpUri="https://zenodo.org/record/4496436/files/figure.png" pageId="207" pageNumber="208">Figures 47</figureCitation>
J–M, 112
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BAEFF0AAF89FBC1FC9E394D" pageId="207" pageNumber="208" type="reference_group">
<paragraph id="8BAB36F06BAEFF0AAF89FBC1FC893E8F" blockId="207.[151,855,939,1025]" box="[151,855,939,963]" pageId="207" pageNumber="208">
<treatmentCitationGroup id="AB0411DE6BAEFF0AAF89FBC1FC893E8F" box="[151,855,939,963]" pageId="207" pageNumber="208">
<treatmentCitation id="0AB510E16BAEFF0AAF89FBC1FDCC3E8F" author="Burmeister, H." box="[151,530,939,963]" page="167" pageId="207" pageNumber="208" year="1855">
<taxonomicName id="4C144D736BAEFF0AAF89FBC1FDCC3E8F" authority="Burmeister, 1855: 167" authorityName="Burmeister" authorityPageNumber="167" authorityYear="1855" box="[151,530,939,963]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26BAEFF0AAF89FBC1FEF83E8F" box="[151,294,939,963]" italics="true" pageId="207" pageNumber="208">Serica pallida</emphasis>
<bibRefCitation id="EF854B016BAEFF0AAE32FBC1FDCC3E8F" author="Burmeister, H." box="[300,530,939,963]" pageId="207" pageNumber="208" refId="ref331559" refString="Burmeister, H. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. T. C. F. Enslin, Berlin, x + 569 pp." type="book" year="1855">Burmeister, 1855: 167</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16BAEFF0AAD05FBC1FD3E3E8F" author="Ahrens, D." box="[539,736,939,963]" page="245" pageId="207" pageNumber="208" year="2006">
<bibRefCitation id="EF854B016BAEFF0AAD05FBC1FD3E3E8F" author="Ahrens, D." box="[539,736,939,963]" pageId="207" pageNumber="208" refId="ref329946" refString="Ahrens, D. (2006 b) Sericinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 229 - 248." type="book" year="2006">Ahrens 2006b: 245</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB510E16BAEFF0AADF4FBC1FC8D3E8F" author="Ahrens, D." box="[746,851,939,963]" page="36" pageId="207" pageNumber="208" year="2007">
<bibRefCitation id="EF854B016BAEFF0AADF4FBC1FC8D3E8F" author="Ahrens, D." box="[746,851,939,963]" pageId="207" pageNumber="208" pagination="1 - 51" refId="ref330155" refString="Ahrens, D. (2007 b) Taxonomic changes and an updated Catalogue of Palearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 1504 (1), 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1504.1.1" type="journal article" year="2007">2007b: 36</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAF89FBA0FD9C3EAE" blockId="207.[151,855,939,1025]" box="[151,578,970,994]" pageId="207" pageNumber="208">
<treatmentCitationGroup id="AB0411DE6BAEFF0AAF89FBA0FD9C3EAE" box="[151,578,970,994]" pageId="207" pageNumber="208">
<treatmentCitation id="0AB510E16BAEFF0AAF89FBA0FDE03EAE" author="Medvedev, S. I." box="[151,574,970,994]" page="144" pageId="207" pageNumber="208" year="1952">
<taxonomicName id="4C144D736BAEFF0AAF89FBA0FDE03EAE" authority=": Medvedev 1952: 144" authorityName=": Medvedev" authorityPageNumber="144" authorityYear="1952" box="[151,574,970,994]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26BAEFF0AAF89FBA0FE883EAE" box="[151,342,970,994]" italics="true" pageId="207" pageNumber="208">Amaladera pallida</emphasis>
:
<bibRefCitation id="EF854B016BAEFF0AAE7DFBA0FDE03EAE" author="Medvedev, S. I." box="[355,574,970,994]" pageId="207" pageNumber="208" refId="ref333942" refString="Medvedev, S. I. (1952) Fauna USSR: Coleoptera 10 Nr. 1: Lamellicornia (Scarabaeidae), subfam. Melolonthinae. Part 2. Izdatelstvo Akademii Nauk SSSR, Moskva, Leningrad, 276 pp." type="book" year="1952">Medvedev 1952: 144</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAF89FB82FC9E394D" blockId="207.[151,855,939,1025]" box="[151,832,1000,1025]" pageId="207" pageNumber="208">
<taxonomicName id="4C144D736BAEFF0AAF89FB82FE1F394C" authority="Miyake" authorityName="Miyake" box="[151,449,1000,1024]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="ludipennis">
<emphasis id="B960EAE26BAEFF0AAF89FB82FEB5394C" box="[151,363,1000,1024]" italics="true" pageId="207" pageNumber="208">Maladera ludipennis</emphasis>
Miyake
</taxonomicName>
,
<collectingRegion id="49D0F8126BAEFF0AAED4FB82FDE2394C" box="[458,572,1000,1024]" country="Japan" name="Yamaguchi" pageId="207" pageNumber="208">Yamaguchi</collectingRegion>
&amp; Aoki 2002: 52,
<emphasis id="B960EAE26BAEFF0AADE5FB83FC9E394D" bold="true" box="[763,832,1001,1025]" pageId="207" pageNumber="208">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BAEFF0AAF89FC45FBFA38C1" pageId="207" pageNumber="208" type="materials_examined">
<paragraph id="8BAB36F06BAEFF0AAF89FC45FC8839F9" blockId="207.[151,1437,1070,2033]" pageId="207" pageNumber="208">
<emphasis id="B960EAE26BAEFF0AAF89FC45FE1A3904" bold="true" box="[151,452,1070,1097]" pageId="207" pageNumber="208">Type material examined:</emphasis>
<materialsCitation id="3B7C3CAD6BAEFF0AAED3FC45FBB53921" ID-GBIF-Occurrence="3034556556" collectionCode="ZMHB, MLUH" country="Hong Kong" location="Hongkong" pageId="207" pageNumber="208" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="54AF88526BAEFF0AAED3FC45FDE33905" box="[461,573,1071,1097]" pageId="207" pageNumber="208" type="lectotype">Lectotype</typeStatus>
(
<taxonomicName id="4C144D736BAEFF0AAD51FC45FD403905" authorityName=": Medvedev" authorityYear="1952" box="[591,670,1071,1097]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26BAEFF0AAD51FC45FD403905" box="[591,670,1071,1097]" italics="true" pageId="207" pageNumber="208">pallida</emphasis>
</taxonomicName>
, here designated):
<specimenCount id="9D12FD796BAEFF0AAC66FC45FC533905" box="[888,909,1071,1097]" pageId="207" pageNumber="208" type="male">♂</specimenCount>
“703 5/9/51/
<collectingCountry id="F30376606BAEFF0AAB32FC45FB7B3905" box="[1068,1189,1071,1097]" name="Hong Kong" pageId="207" pageNumber="208">Hongkong</collectingCountry>
” (
<collectionCode id="ED05AE356BAEFF0AABDEFC45FAC93905" box="[1216,1303,1071,1097]" pageId="207" pageNumber="208">ZMHB</collectionCode>
). Paralectotype (
<taxonomicName id="4C144D736BAEFF0AAFC4FC39FEF73921" authorityName=": Medvedev" authorityYear="1952" box="[218,297,1107,1133]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26BAEFF0AAFC4FC39FEF73921" box="[218,297,1107,1133]" italics="true" pageId="207" pageNumber="208">pallida</emphasis>
</taxonomicName>
, here designated):
<specimenCount id="9D12FD796BAEFF0AAD18FC39FDED3921" box="[518,563,1107,1133]" pageId="207" pageNumber="208" type="female">1 ♀</specimenCount>
“703 9/5/51/
<taxonomicName id="4C144D736BAEFF0AADCBFC39FC313921" authority="Br. Hongk. Dhn." authorityName="Br. Hongk. Dhn." box="[725,1007,1107,1133]" class="Insecta" family="Melolonthidae" genus="Amaladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26BAEFF0AADCBFC39FCFA3921" box="[725,804,1107,1133]" italics="true" pageId="207" pageNumber="208">pallida</emphasis>
Br. Hongk. Dhn.
</taxonomicName>
” (
<collectionCode id="ED05AE356BAEFF0AAB14FC39FBBD3921" box="[1034,1123,1107,1133]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="207" pageNumber="208">MLUH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BAEFF0AAB66FC39FC8C39F9" ID-GBIF-Occurrence="3034555305" collectingDate="1999-02-23" collectionCode="RIEBT" collectorName="S. Yamaguchi &amp; T. Aoki" country="Hong Kong" pageId="207" pageNumber="208" specimenCount="2" typeStatus="holotype">
<typeStatus id="54AF88526BAEFF0AAB66FC39FB3E3921" box="[1144,1248,1107,1133]" pageId="207" pageNumber="208" type="holotype">Holotype</typeStatus>
(
<taxonomicName id="4C144D736BAEFF0AABEDFC39FAB63921" box="[1267,1384,1107,1133]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="ludipennis">
<emphasis id="B960EAE26BAEFF0AABEDFC39FAB63921" box="[1267,1384,1107,1133]" italics="true" pageId="207" pageNumber="208">ludipennis</emphasis>
</taxonomicName>
): @ “Ban Lao Kram Mukdahan (light) E-Thailand
<date id="FFAA10306BAEFF0AADAAFC1DFC8A39DD" box="[692,852,1143,1169]" pageId="207" pageNumber="208" value="1999-02-23">
<collectingDate id="EFEEE9D86BAEFF0AADAAFC1DFC8A39DD" box="[692,852,1143,1169]" pageId="207" pageNumber="208" value="1999-02-23">23. Feb. 1999</collectingDate>
</date>
<collectorName id="26E153266BAEFF0AAC40FC1DFC2539DD" box="[862,1019,1143,1169]" pageId="207" pageNumber="208">S. Yamaguchi</collectorName>
&amp;
<collectorName id="26E153266BAEFF0AAB3BFC1DFBA539DD" box="[1061,1147,1143,1169]" pageId="207" pageNumber="208">T. Aoki</collectorName>
leg./
<typeStatus id="54AF88526BAEFF0AABDEFC1DFAF639DD" box="[1216,1320,1143,1169]" pageId="207" pageNumber="208" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C144D736BAEFF0AAA2CFC1DFE5639F9" authority="Y. Miyake" authorityName="Y. Miyake" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="species" species="ludipennis">Maladera ludipennis Y. Miyake</taxonomicName>
, Yamaguchi et Akiyama 2002” (
<collectionCode id="ED05AE356BAEFF0AADEDFCF1FC9439F9" box="[755,842,1179,1205]" pageId="207" pageNumber="208">RIEBT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAFD9FCD4FAB53825" blockId="207.[151,1437,1070,2033]" pageId="207" pageNumber="208">
<materialsCitation id="3B7C3CAD6BAEFF0AAFD9FCD4FA863995" ID-GBIF-Occurrence="3034556077" box="[199,1368,1214,1241]" collectionCode="MLUH" country="China" location="China" pageId="207" pageNumber="208" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<emphasis id="B960EAE26BAEFF0AAFD9FCD4FD4F3994" bold="true" box="[199,657,1214,1240]" pageId="207" pageNumber="208">
Additional material examined.
<collectingCountry id="F30376606BAEFF0AAD21FCD4FD523994" box="[575,652,1214,1240]" name="China" pageId="207" pageNumber="208">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BAEFF0AAD87FCD5FD1B3995" box="[665,709,1215,1241]" pageId="207" pageNumber="208" type="female">1 ♀</specimenCount>
[
<typeStatus id="54AF88526BAEFF0AADCAFCD5FCE93995" box="[724,823,1215,1241]" pageId="207" pageNumber="208" type="syntype">Syntype</typeStatus>
?; without abdomen and proper labels] (
<collectionCode id="ED05AE356BAEFF0AABE6FCD5FA8E3995" box="[1272,1360,1215,1241]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="207" pageNumber="208">MLUH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BAEFF0AAA7BFCD5FF2B39B1" ID-GBIF-Occurrence="3034556148" country="China" pageId="207" pageNumber="208" specimenCount="1">
<specimenCount id="9D12FD796BAEFF0AAA7BFCD5FA433995" box="[1381,1437,1215,1241]" pageId="207" pageNumber="208" type="generic">1 ex.</specimenCount>
“China”
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BAEFF0AAE1AFC89FD9939B1" ID-GBIF-Occurrence="3034556515" box="[260,583,1251,1277]" collectionCode="ZMHB" country="China" location="Hongkong" pageId="207" pageNumber="208" specimenCount="1">
<specimenCount id="9D12FD796BAEFF0AAE1AFC89FEE039B1" box="[260,318,1251,1277]" pageId="207" pageNumber="208" type="generic">1 ex.</specimenCount>
“
<location id="8ECB602B6BAEFF0AAE4CFC89FE1039B1" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979:8ECB602B6BAEFF0AAE4CFC89FE1039B1" box="[338,462,1251,1277]" country="China" name="Hongkong" pageId="207" pageNumber="208">Hongkong</location>
” (
<collectionCode id="ED05AE356BAEFF0AAEF7FC89FDE139B1" box="[489,575,1251,1277]" pageId="207" pageNumber="208">ZMHB</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BAEFF0AAD48FC89FB5739B1" ID-GBIF-Occurrence="3034555663" box="[598,1161,1251,1277]" collectionCode="ZMUC" country="China" pageId="207" pageNumber="208" specimenCount="2" specimenCount-female="1">
<specimenCount id="9D12FD796BAEFF0AAD48FC89FD4E39B1" box="[598,656,1251,1277]" pageId="207" pageNumber="208" type="generic">1 ex.</specimenCount>
(
<specimenCount id="9D12FD796BAEFF0AADBAFC89FD6839B1" box="[676,694,1251,1277]" pageId="207" pageNumber="208" type="female">♀</specimenCount>
) “Mus. Westerm./ Hongkong” (
<collectionCode id="ED05AE356BAEFF0AAB35FC89FB5C39B1" box="[1067,1154,1251,1277]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="207" pageNumber="208">ZMUC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BAEFF0AAB86FC89FEDC386D" ID-GBIF-Occurrence="3034555972" collectionCode="HNHM" country="China" location="China Tsonglok" pageId="207" pageNumber="208" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD796BAEFF0AAB86FC89FB1B39B1" box="[1176,1221,1251,1277]" pageId="207" pageNumber="208" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6BAEFF0AABC7FC89FA4D39B1" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979:8ECB602B6BAEFF0AABC7FC89FA4D39B1" box="[1241,1427,1251,1277]" country="China" name="China Tsonglok" pageId="207" pageNumber="208">China Tsonglok</location>
” (
<collectionCode id="ED05AE356BAEFF0AAF81FD6DFF24386D" box="[159,250,1287,1313]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="207" pageNumber="208">HNHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CAD6BAEFF0AAE0EFD6DFC95386D" ID-GBIF-Occurrence="3034555934" box="[272,843,1287,1313]" collectionCode="HNHM" collectorName="Coll. Reitter" country="China" location="Hongkong" municipality="China" pageId="207" pageNumber="208" specimenCount="3">
<specimenCount id="9D12FD796BAEFF0AAE0EFD6DFE94386D" box="[272,330,1287,1313]" pageId="207" pageNumber="208" type="generic">3 ex.</specimenCount>
“
<location id="8ECB602B6BAEFF0AAE43FD6DFE0B386D" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979:8ECB602B6BAEFF0AAE43FD6DFE0B386D" box="[349,469,1287,1313]" country="China" municipality="China" name="Hongkong" pageId="207" pageNumber="208">Hongkong</location>
/
<collectingMunicipality id="6BCFAC8A6BAEFF0AAEF9FD6DFDF7386D" box="[487,553,1287,1313]" pageId="207" pageNumber="208">China</collectingMunicipality>
/
<collectorName id="26E153266BAEFF0AAD22FD6DFD12386D" box="[572,716,1287,1313]" pageId="207" pageNumber="208">Coll. Reitter</collectorName>
” (
<collectionCode id="ED05AE356BAEFF0AADF9FD6DFC9D386D" box="[743,835,1287,1313]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="207" pageNumber="208">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BAEFF0AAC46FD6CFE323809" ID-GBIF-Occurrence="3034556527" collectionCode="ZMHB" country="Vietnam" location="Tonkin Dap" pageId="207" pageNumber="208" specimenCount="8">
<emphasis id="B960EAE26BAEFF0AAC46FD6CFC19386C" bold="true" box="[856,967,1286,1312]" pageId="207" pageNumber="208">
<collectingCountry id="F30376606BAEFF0AAC46FD6CFC1C386C" box="[856,962,1286,1312]" name="Vietnam" pageId="207" pageNumber="208">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796BAEFF0AACCFFD6DFBD4386D" box="[977,1034,1287,1313]" pageId="207" pageNumber="208" type="generic">8 ex.</specimenCount>
“
<location id="8ECB602B6BAEFF0AAB03FD6DFB76386D" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979:8ECB602B6BAEFF0AAB03FD6DFB76386D" box="[1053,1192,1287,1313]" country="Vietnam" name="Tonkin Dap" pageId="207" pageNumber="208">Tonkin Dap</location>
ban/
<emphasis id="B960EAE26BAEFF0AABF5FD6DFA88386C" box="[1259,1366,1287,1312]" italics="true" pageId="207" pageNumber="208">contracta</emphasis>
Brsk. [Handschr. Moser]” (
<collectionCode id="ED05AE356BAEFF0AAE93FD41FE3A3809" box="[397,484,1323,1349]" pageId="207" pageNumber="208">ZMHB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BAEFF0AAEE4FD41FAB63825" ID-GBIF-Occurrence="3034556555" collectingDate="1997-04-27" collectingDateMax="1997-05-01" collectingDateMin="1997-04-27" collectionCode="CP" collectorName="P. Pacholatko" country="Laos" elevation="150" location="Laos" longitude="103.18166" municipality="Ban Phabat" pageId="207" pageNumber="208" specimenCount="1" stateProvince="Viangchan">
<emphasis id="B960EAE26BAEFF0AAEE4FD41FDE33809" bold="true" box="[506,573,1323,1349]" pageId="207" pageNumber="208">
<collectingCountry id="F30376606BAEFF0AAEE4FD41FDE63809" box="[506,568,1323,1349]" name="Laos" pageId="207" pageNumber="208">Laos</collectingCountry>
:
</emphasis>
1 @ “
<location id="8ECB602B6BAEFF0AAD8EFD41FD173809" LSID="urn:lsid:plazi:treatment:03BD87E66BAEFF15AF89FB28FD133979:8ECB602B6BAEFF0AAD8EFD41FD173809" box="[656,713,1323,1349]" country="Laos" longitude="103.18166" municipality="Ban Phabat" name="Laos" pageId="207" pageNumber="208" stateProvince="Viangchan">Laos</location>
centr.,
<date id="FFAA10306BAEFF0AAC3DFD41FC0B3809" box="[803,981,1323,1349]" pageId="207" pageNumber="208" value="1997-04-27" valueMax="1997-05-01" valueMin="1997-04-27">
<collectingDate id="EFEEE9D86BAEFF0AAC3DFD41FC0B3809" box="[803,981,1323,1349]" pageId="207" pageNumber="208" value="1997-04-27" valueMax="1997-05-01" valueMin="1997-04-27">27.IV.-1.V.1997</collectingDate>
</date>
<locationDeviation id="9A192FD26BAEFF0AACFEFD41FB173809" box="[992,1225,1323,1349]" location="Vientiane" pageId="207" pageNumber="208">
<quantity id="4CEC9B156BAEFF0AACFEFD41FBFE3809" box="[992,1056,1323,1349]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="207" pageNumber="208" unit="km" value="70.0">70km</quantity>
NE
<collectingRegion id="49D0F8126BAEFF0AAB45FD41FB173809" box="[1115,1225,1323,1349]" country="Laos" name="Viangchan" pageId="207" pageNumber="208">Vientiane</collectingRegion>
</locationDeviation>
,
<collectingMunicipality id="6BCFAC8A6BAEFF0AABC6FD41FA823809" box="[1240,1372,1323,1349]" pageId="207" pageNumber="208">Ban Phabat</collectingMunicipality>
env.,
<quantity id="4CEC9B156BAEFF0AAF89FD24FF043825" box="[151,218,1358,1385]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="207" pageNumber="208" unit="m" value="150.0">
<elevation id="0039D1C36BAEFF0AAF89FD24FF043825" box="[151,218,1358,1385]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="207" pageNumber="208" unit="m" value="150.0">150m</elevation>
</quantity>
,
<geoCoordinate id="EE2050376BAEFF0AAFFBFD25FEBA3825" box="[229,356,1359,1385]" pageId="207" pageNumber="208">N 18°16.1’</geoCoordinate>
,
<geoCoordinate id="EE2050376BAEFF0AAE6EFD25FE283825" box="[368,502,1359,1385]" pageId="207" pageNumber="208">E 103°10.9’</geoCoordinate>
E.
<collectorName id="26E153266BAEFF0AAD05FD25FDB73825" box="[539,617,1359,1385]" pageId="207" pageNumber="208">Jendek</collectorName>
&amp;
<collectorName id="26E153266BAEFF0AAD93FD25FD2E3827" box="[653,752,1359,1387]" pageId="207" pageNumber="208">O. Šauša</collectorName>
leg./
<collectorName id="26E153266BAEFF0AAC31FD25FBDA3825" box="[815,1028,1359,1385]" pageId="207" pageNumber="208">Coll. P. Pacholatko</collectorName>
/ 360
<taxonomicName id="4C144D736BAEFF0AAB5AFD25FB7E3825" box="[1092,1184,1359,1385]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
: Asia spec.” (
<collectionCode id="ED05AE356BAEFF0AAA26FD25FA813825" box="[1336,1375,1359,1385]" pageId="207" pageNumber="208">CP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAFD9FD19FBFA38C1" blockId="207.[151,1437,1070,2033]" box="[199,1060,1395,1421]" pageId="207" pageNumber="208">
<materialsCitation id="3B7C3CAD6BAEFF0AAFD9FD19FBFE38C1" ID-GBIF-Occurrence="3034555405" box="[199,1056,1395,1421]" collectionCode="SMTD" collectorName="Coll. C. Felsche Kauf" country="Bangladesh" location="Bengalen" pageId="207" pageNumber="208" specimenCount="8">
Doubtful record:
<specimenCount id="9D12FD796BAEFF0AAE94FD19FE1E38C1" box="[394,448,1395,1421]" pageId="207" pageNumber="208" type="generic">8 ex.</specimenCount>
“
<collectingCountry id="F30376606BAEFF0AAECFFD19FDE338C1" box="[465,573,1395,1421]" name="Bangladesh" pageId="207" pageNumber="208">Bengalen</collectingCountry>
/
<collectorName id="26E153266BAEFF0AAD52FD19FCE138C1" box="[588,831,1395,1421]" pageId="207" pageNumber="208">Coll. C. Felsche Kauf</collectorName>
20, 1918” (
<collectionCode id="ED05AE356BAEFF0AACDAFD19FBC638C1" box="[964,1048,1395,1421]" pageId="207" pageNumber="208">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BAEFF15AFD9FDFCFEFB3E85" lastPageId="208" lastPageNumber="209" pageId="207" pageNumber="208" type="description">
<paragraph id="8BAB36F06BAEFF0AAFD9FDFCFB003899" blockId="207.[151,1437,1070,2033]" pageId="207" pageNumber="208">
<emphasis id="B960EAE26BAEFF0AAFD9FDFCFEAB38FC" bold="true" box="[199,373,1430,1456]" pageId="207" pageNumber="208">Redescription.</emphasis>
Length:
<quantity id="4CEC9B156BAEFF0AAEC6FDFDFDF138FD" box="[472,559,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="207" pageNumber="208" unit="mm" value="4.9">4.9 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BAEFF0AADECFDFDFC9738FD" box="[754,841,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="207" pageNumber="208" unit="mm" value="3.8">3.8 mm</quantity>
, width: 3.0 mm. Body oblong-oval, yellowish brown, dull, antenna yellow, head shiny, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAFD9FDB5FC9A3A55" blockId="207.[151,1437,1070,2033]" pageId="207" pageNumber="208">
Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and distinctly convergent anteriorly, anterior angles moderately rounded, anterior margin emarginate medially, strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, finely and densely punctate, with a few larger punctures each bearing an erect seta anteriorly; frontoclypeal suture indistinct, evenly curved; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely and densely punctate, with a terminal seta. Frons completely shiny, with very dense, fine punctures, behind frontoclypeal suture and beside eyes with a few long setae in large sparse punctures. Eyes small, ratio diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three antennomeres and straight, distinctly shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126BAEFF0AAE0AFE95FEAD3A55" box="[276,371,1791,1817]" country="Myanmar" name="Chin" pageId="207" pageNumber="208">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BAEFF0AAFD9FF49FD683A81" blockId="207.[151,1437,1070,2033]" pageId="207" pageNumber="208">Pronotum transverse, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles strongly rounded; anterior margin straight, with complete and fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures; lateral anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures.</paragraph>
<paragraph id="8BAB36F06BAEFF15AFD9FFBDFE603C52" blockId="207.[151,1437,1070,2033]" lastBlockId="208.[151,1437,151,1077]" lastPageId="208" lastPageNumber="209" pageId="207" pageNumber="208">Elytra widest shortly behind middle, striae finely impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, penultimate interval with a very few single, short setae; epipleural margin robust, ending at weakly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06BB1FF15AFD9F94DFAED3C9D" blockId="208.[151,1437,151,1077]" pageId="208" pageNumber="209">Ventral surface dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.35. Pygidium moderately convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few short setae along apical margin.</paragraph>
<paragraph id="8BAB36F06BB1FF15AFD9F9B1FA453E11" blockId="208.[151,1437,151,1077]" pageId="208" pageNumber="209">Legs short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae complete; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at one quarter, apical group at two thirds of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and glabrous; ventral margin finely serrate, with four equidistant, short and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally and dorsally not carinate; first metatarsomere little shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06BB1FF15AFD9FB0DFD7F3ECD" blockId="208.[151,1437,151,1077]" box="[199,673,871,897]" pageId="208" pageNumber="209">
Aedeagus:
<figureCitation id="132F2A756BB1FF15AE5AFB0DFE4B3ECD" box="[324,405,871,897]" captionStart="FIGURE 47" captionStartId="204.[151,250,1944,1968]" captionTargetBox="[212,1375,197,1908]" captionTargetId="figure-19@204.[198,1389,181,1920]" captionTargetPageId="204" captionText="FIGURE 47. A–E: Maladera bansongchana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. peregoi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. pallida (Burmeister, 1855) (China). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496436" httpUri="https://zenodo.org/record/4496436/files/figure.png" pageId="208" pageNumber="209">Fig. 47</figureCitation>
J–L. Habitus:
<figureCitation id="132F2A756BB1FF15AD2CFB0DFD403ECD" box="[562,670,871,897]" captionStart="FIGURE 47" captionStartId="204.[151,250,1944,1968]" captionTargetBox="[212,1375,197,1908]" captionTargetId="figure-19@204.[198,1389,181,1920]" captionTargetPageId="204" captionText="FIGURE 47. A–E: Maladera bansongchana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. peregoi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. pallida (Burmeister, 1855) (China). A, F, J: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496436" httpUri="https://zenodo.org/record/4496436/files/figure.png" pageId="208" pageNumber="209">
Fig.
<specimenCount id="9D12FD796BB1FF15AD7BFB0DFD403ECD" box="[613,670,871,897]" pageId="208" pageNumber="209" type="generic">47M</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="8BAB36F06BB1FF15AFD9FBE0FEFB3E85" blockId="208.[151,1437,151,1077]" pageId="208" pageNumber="209">
<emphasis id="B960EAE26BB1FF15AFD9FBE0FEE13EE8" bold="true" box="[199,319,906,932]" pageId="208" pageNumber="209">Variation.</emphasis>
Pronotum in male slightly wider, pygidium wider and more convex; antennal club in both sexes of same length.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BB1FF15AFD9FBB9FE59395D" pageId="208" pageNumber="209" type="discussion">
<paragraph id="8BAB36F06BB1FF15AFD9FBB9FE59395D" blockId="208.[151,1437,151,1077]" pageId="208" pageNumber="209">
<emphasis id="B960EAE26BB1FF15AFD9FBB9FEE53EA1" bold="true" box="[199,315,979,1005]" pageId="208" pageNumber="209">Remarks.</emphasis>
The shape of the aedeagus in this species is very similar that of
<taxonomicName id="4C144D736BB1FF15AB35FBB9FF11395D" authority="Arrow, 1946" authorityName="Arrow" authorityYear="1946" class="Insecta" family="Melolonthidae" genus="Cephaloserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="208" pageNumber="209" phylum="Arthropoda" rank="species" species="vulpecula">
<emphasis id="B960EAE26BB1FF15AB35FBB9FA9B3EA1" box="[1067,1349,979,1005]" italics="true" pageId="208" pageNumber="209">Cephaloserica vulpecula</emphasis>
Arrow, 1946
</taxonomicName>
from
<collectingCountry id="F30376606BB1FF15AE0AFB9DFE5D395D" box="[276,387,1015,1041]" name="Myanmar" pageId="208" pageNumber="209">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BB1FF15AFD9FC70FD133979" box="[199,717,1050,1077]" pageId="208" pageNumber="209" type="distribution">
<paragraph id="8BAB36F06BB1FF15AFD9FC70FD133979" blockId="208.[151,1437,151,1077]" box="[199,717,1050,1077]" pageId="208" pageNumber="209">
<emphasis id="B960EAE26BB1FF15AFD9FC70FEBF3978" bold="true" box="[199,353,1050,1076]" pageId="208" pageNumber="209">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BB1FF15AEC8FC71FDEB3979" box="[470,565,1051,1077]" captionStart="FIGURE 112" captionStartId="357.[151,250,1981,2005]" captionTargetBox="[202,1384,1127,1951]" captionTargetId="figure-59@357.[196,1392,1118,1957]" captionTargetPageId="357" captionText="FIGURE 112. Distribution of the Maladera indica group: Maladera lushuiensis sp. n., M. pallida (Burmeister, 1855), M. peregoi sp. n., M. pseudoconsularis sp. n., M. sinobiloba sp. n., M. yongrenensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496622" httpUri="https://zenodo.org/record/4496622/files/figure.png" pageId="208" pageNumber="209">Fig. 112</figureCitation>
) and
<tableCitation id="C696034B6BB1FF15AD6DFC71FD193979" box="[627,711,1051,1077]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="208" pageNumber="209">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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