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<document id="7DC90CDE89BE46EB3D98768A3AA8E605" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B85FF20AF89FAAFFD133F71" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera liwenzhui Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="230" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="229" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="A803531FF50D72CAF1BAE399CE866629" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DA5DFBB6F008C0B6341044D3BA76AB1D">
<mods:title id="F0AAF125DA2102E0FDA153AEA688EA34">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="25FEE25808ECABFC84159AC522A02793" type="personal">
<mods:role id="529515FC4D36CC0C691DED42E7737B94">
<mods:roleTerm id="9B0E7187DBF98E54F2FD491666EE6C8D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4173DABC67ABB2073D67B4F168509DB4">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="0FE070B56FDA57774D132508F967B80A" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="43DEF7A9EE2423EC13C3F418F3BA5E83" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="F89C38F390E32E59BD8C0849B18032DA" type="personal">
<mods:role id="CDBC071FA5305F9913A14DC653BD6CC7">
<mods:roleTerm id="EF560C00AB06F1005BFBCB3B27E3498A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B353850E4FBC7C45E09A369350C8A5F0">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="B7639887340CA02A34319A99D3D2695F" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="CA12FC2C745E95645D79505E90B20474" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="F0D53C7863272FA74D954987770D2A51" type="personal">
<mods:role id="8C0A11BF466DAECB69226AE982D617A9">
<mods:roleTerm id="F138D68932DD748476F18104A55D2F0D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D32361114FF82B17FA802A5DC63C6E2C">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="767437D7EA4129F7DBB0EDBA74AA43F4" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="00FE5785B984ED5B405794B4B6E19BC2" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="28F486B5AA80460AA792CD3DA2A58623" type="personal">
<mods:role id="288D82504CAB7F11AA1606FE0F09C3C3">
<mods:roleTerm id="21F12390BDC7170F3DD27BA5BD0A9B86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E60FEDB8E85D34DFE995395C9A7D107">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="EAFCE106A475E2D7C8C2E28E197344DB" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="14A4FEBF8EEB8039E5D90A2FB2E1C45A" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="ABBAA0151DAE00AC26DBEA80A9BFD2D3" type="personal">
<mods:role id="67DA9071498B87CB8A9A2AD5539CC26E">
<mods:roleTerm id="7ACDA175A82FA7117A3F4FB7FA10471E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="013D1D88B5D008CC3908C4BA2524CA58">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="06FB1D14388CE640DFB37B4FE5572A8E" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="53DE4FAFE29E711A1F45420339E83D8B" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0D83F0B821AD8879120741D4CD7D6135">text</mods:typeOfResource>
<mods:relatedItem id="B9784D31BAE70A3094F36162C843A25A" type="host">
<mods:titleInfo id="52A676BF6CB0D102FEF5A83D64168585">
<mods:title id="F3F07E36500BF8F48EA81E9F446149FF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1167D60B1D135300927A103C50B0937C">
<mods:date id="588E4B97C71509C622620B06F634C322">2021</mods:date>
<mods:detail id="177C5C0A1E3B59013D956D0541B0ECCF" type="pubDate">
<mods:number id="79EB886245C4E1D741E0D6B2388141EA">2021-02-03</mods:number>
</mods:detail>
<mods:detail id="0276B1EAD7469658FD8B616534CBFF16" type="volume">
<mods:number id="659276F67C6E6EBD10000BC4CDF7668C">4922</mods:number>
</mods:detail>
<mods:detail id="5318FDECA23B1B34E88E685004051504" type="issue">
<mods:number id="CA529ACEA9B5AD5576003C77F9660B61">1</mods:number>
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<mods:start id="9C46C7E8350D20E1DDC4251D46040381">1</mods:start>
<mods:end id="CFFB87A5BEC8A0916F88561AC2D86469">400</mods:end>
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<mods:classification id="0B9420D33D1F936B35FE82FB2CC7F2DC">journal article</mods:classification>
<mods:identifier id="B6CDAD25A604016C171DBDB32D82ADC9" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="E2DBF465B6F1F871A55CA11103D0A8E4" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="4A90DFFC4017361690866310B48AFA26" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="A259AC0CFAF38ACD919CDBBCE4F0A6A9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E5721AA1E384236FC9CD4B4B314FFF3F" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="ECC23BCADCD37EEDDF989C47783DB989" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
</mods:mods>
<treatment id="03BD87E66B85FF20AF89FAAFFD133F71" ID-DOI="http://doi.org/10.5281/zenodo.4534756" ID-GBIF-Taxon="177323614" ID-Zenodo-Dep="4534756" LSID="urn:lsid:plazi:treatment:03BD87E66B85FF20AF89FAAFFD133F71" httpUri="http://treatment.plazi.org/id/03BD87E66B85FF20AF89FAAFFD133F71" lastPageId="229" lastPageNumber="230" pageId="228" pageNumber="229">
<subSubSection id="C30E657B6B85FF21AF89FAAFFD273F93" box="[151,761,709,736]" pageId="228" pageNumber="229" type="nomenclature">
<paragraph id="8BAB36F06B85FF21AF89FAAFFD273F93" blockId="228.[151,761,709,772]" box="[151,761,709,736]" pageId="228" pageNumber="229">
<heading id="D0E3819C6B85FF21AF89FAAFFD273F93" bold="true" box="[151,761,709,736]" fontSize="11" level="2" pageId="228" pageNumber="229" reason="4">
<emphasis id="B960EAE26B85FF21AF89FAAFFD273F93" bold="true" box="[151,761,709,736]" pageId="228" pageNumber="229">
<taxonomicName id="4C144D736B85FF21AF89FAAFFD743F93" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,682,709,736]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="liwenzhui" status="sp. nov.">
<emphasis id="B960EAE26B85FF21AF89FAAFFE573FAC" bold="true" box="[151,393,709,736]" italics="true" pageId="228" pageNumber="229">Maladera liwenzhui</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B85FF21ADA9FAAFFD273F93" box="[695,761,709,735]" pageId="228" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B85FF21AF89FA83FEB03E4F" box="[151,366,745,772]" pageId="228" pageNumber="229" type="description">
<paragraph id="8BAB36F06B85FF21AF89FA83FEB03E4F" blockId="228.[151,761,709,772]" box="[151,366,745,772]" pageId="228" pageNumber="229">
<figureCitation id="132F2A756B85FF21AF89FA83FED03E48" box="[151,270,745,772]" captionStart="FIGURE 52" captionStartId="224.[151,250,1933,1957]" captionTargetBox="[225,1362,288,1896]" captionTargetId="figure-64@224.[213,1375,273,1909]" captionTargetPageId="224" captionText="FIGURE 52. A–D: Maladera filigraniforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. flavipennis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. liwenzhui Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496448" httpUri="https://zenodo.org/record/4496448/files/figure.png" pageId="228" pageNumber="229">Figures 52</figureCitation>
I–L, 117
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B85FF21AF89FB5BFAD13E23" pageId="228" pageNumber="229" type="materials_examined">
<paragraph id="8BAB36F06B85FF21AF89FB5BFAD13E23" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">
<emphasis id="B960EAE26B85FF21AF89FB5BFE163E07" bold="true" box="[151,456,817,843]" pageId="228" pageNumber="229">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B85FF21AECAFB5BFF353E23" ID-GBIF-Occurrence="3034556901" collectingDate="2000-06-19" collectionCode="IZAS" collectorName="Li Wenzhu" country="China" location="Napo" municipality="Defu" pageId="228" pageNumber="229" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="54AF88526B85FF21AECAFB5BFD9F3E07" box="[468,577,817,843]" pageId="228" pageNumber="229" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796B85FF21AD51FB58FDBA3E00" box="[591,612,818,844]" pageId="228" pageNumber="229" type="male">♂</specimenCount>
“[
<collectingCountry id="F30376606B85FF21AD9FFB5BFD153E00" box="[641,715,817,844]" name="China" pageId="228" pageNumber="229">China</collectingCountry>
]
<collectingMunicipality id="6BCFAC8A6B85FF21ADC3FB5BFCC63E07" box="[733,792,817,843]" pageId="228" pageNumber="229">Defu</collectingMunicipality>
,
<location id="8ECB602B6B85FF21AC36FB58FCB93E00" LSID="urn:lsid:plazi:treatment:03BD87E66B85FF20AF89FAAFFD133F71:8ECB602B6B85FF21AC36FB58FCB93E00" box="[808,871,818,844]" country="China" municipality="Defu" name="Napo" pageId="228" pageNumber="229" stateProvince="Guangxi">Napo</location>
,
<collectingRegion id="49D0F8126B85FF21AC69FB5BFC043E07" box="[887,986,817,843]" country="China" name="Guangxi" pageId="228" pageNumber="229">Guangxi</collectingRegion>
,
<date id="FFAA10306B85FF21ACF4FB5BFBB23E07" box="[1002,1132,817,843]" pageId="228" pageNumber="229" value="2000-06-19">
<collectingDate id="EFEEE9D86B85FF21ACF4FB5BFBB23E07" box="[1002,1132,817,843]" pageId="228" pageNumber="229" value="2000-06-19">19.VI.2000</collectingDate>
</date>
, 1350m, leg.
<collectorName id="26E153266B85FF21AA0FFB5BFA4D3E07" box="[1297,1427,817,843]" pageId="228" pageNumber="229">Li Wenzhu</collectorName>
” (
<collectionCode id="ED05AE356B85FF21AF81FB3FFF3D3E23" box="[159,227,853,879]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="228" pageNumber="229">IZAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6B85FF21AFE8FB3CFAD53E23" ID-GBIF-Occurrence="3034555433" box="[246,1291,853,880]" collectingDate="2000-06-19" collectionCode="ZFMK" collectorName="Li Wenzhu" country="China" location="Napo" municipality="Defu" pageId="228" pageNumber="229" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus id="54AF88526B85FF21AFE8FB3CFE823E3C" box="[246,348,854,880]" pageId="228" pageNumber="229" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D12FD796B85FF21AE79FB3FFE4F3E3C" box="[359,401,853,880]" pageId="228" pageNumber="229" type="male">1 ♂</specimenCount>
“[
<collectingCountry id="F30376606B85FF21AEB7FB3FFE2D3E3C" box="[425,499,853,880]" name="China" pageId="228" pageNumber="229">China</collectingCountry>
]
<collectingMunicipality id="6BCFAC8A6B85FF21AD1FFB3FFDE23E23" box="[513,572,853,879]" pageId="228" pageNumber="229">Defu</collectingMunicipality>
,
<location id="8ECB602B6B85FF21AD59FB3CFD583E3C" LSID="urn:lsid:plazi:treatment:03BD87E66B85FF20AF89FAAFFD133F71:8ECB602B6B85FF21AD59FB3CFD583E3C" box="[583,646,854,880]" country="China" municipality="Defu" name="Napo" pageId="228" pageNumber="229" stateProvince="Guangxi">Napo</location>
,
<collectingRegion id="49D0F8126B85FF21AD8FFB3FFD2A3E23" box="[657,756,853,879]" country="China" name="Guangxi" pageId="228" pageNumber="229">Guangxi</collectingRegion>
,
<date id="FFAA10306B85FF21AC1EFB3FFC5C3E23" box="[768,898,853,879]" pageId="228" pageNumber="229" value="2000-06-19">
<collectingDate id="EFEEE9D86B85FF21AC1EFB3FFC5C3E23" box="[768,898,853,879]" pageId="228" pageNumber="229" value="2000-06-19">19.VI.2000</collectingDate>
</date>
, 1350m, leg.
<collectorName id="26E153266B85FF21AB07FB3FFB493E23" box="[1049,1175,853,879]" pageId="228" pageNumber="229">Li Wenzhu</collectorName>
” (
<collectionCode id="ED05AE356B85FF21ABB1FB3FFADD3E23" box="[1199,1283,853,879]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="228" pageNumber="229" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B85FF20AFD9FB13FCB83C51" lastPageId="229" lastPageNumber="230" pageId="228" pageNumber="229" type="description">
<paragraph id="8BAB36F06B85FF21AFD9FB13FB593EFB" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">
<emphasis id="B960EAE26B85FF21AFD9FB13FE873EDF" bold="true" box="[199,345,889,915]" pageId="228" pageNumber="229">Description.</emphasis>
Length:
<quantity id="4CEC9B156B85FF21AEA2FB13FDFE3ED8" box="[444,544,889,916]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="228" pageNumber="229" unit="mm" value="11.3">11.3 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B85FF21ADFDFB13FCE43ED8" box="[739,826,889,916]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="228" pageNumber="229" unit="mm" value="8.9">8.9 mm</quantity>
, width:
<quantity id="4CEC9B156B85FF21AC8CFB13FC353ED8" box="[914,1003,889,916]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="228" pageNumber="229" unit="mm" value="6.2">6.2 mm</quantity>
. Body wide, oval, dark brown, antenna yellowish brown, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous.
</paragraph>
<paragraph id="8BAB36F06B85FF21AFD9FBABFAA639B7" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">
Labroclypeus wide, lateral margins weakly convex and moderately convergent, lateral margins produce with ocular canthus a blunt angle, anterior angles convex, anterior margin shallowly emarginate medially, margins weakly reflexed; surface moderately shiny, at base narrowly dull, surface weakly convex, finely and densely punctate, distance between punctures smaller their diameter, with numerous long erect setae; smooth area in front of eyes twice as wide as long; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and short (1/3 of ocular diameter), finely and densely punctate, with one or two short terminal setae. Eyes moderately large, ratio diameter/ interocular width: 0.54. Frons with fine and moderately dense punctures, with a few single setae beside eyes and behind frontoclypeal suture. Antenna with ten antennomeres, club with three antennomeres and straight, as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126B85FF21AC28FC88FC4B39B0" box="[822,917,1250,1276]" country="Myanmar" name="Chin" pageId="228" pageNumber="229">Mentum</collectingRegion>
convexly elevated and flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B85FF21AFD9FD6CFD8838E3" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">Pronotum widest shortly before base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles strongly rounded; anterior margin convex, with complete marginal line; basal margin without marginal line; surface finely and densely punctate, punctures with microscopic setae only, setae of lateral and anterior margin sparse; hypomeron carinate. Scutellum wide, triangular, punctation as in pronotum, impunctate on basal midline.</paragraph>
<paragraph id="8BAB36F06B85FF21AFD9FDD3FDA03B73" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">Elytra convex, widest at posterior third, striae finely impressed, finely punctate, intervals flat, finely and densely punctate, punctures concentrated along striae, with minute setae in punctures, odd intervals with a few erect short setae; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06B85FF21AFD9FE23FBF33BB8" blockId="228.[151,1437,817,2032]" pageId="228" pageNumber="229">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Distance between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta; penultimate sternite medially retracted, ultimate sternite twice as long as penultimate. Pygidium moderately convex, coarsely and densely punctate, on apex with a few long setae.</paragraph>
<paragraph id="8BAB36F06B85FF20AFD9FE94FE083DB5" blockId="228.[151,1437,817,2032]" lastBlockId="229.[151,1437,151,573]" lastPageId="229" lastPageNumber="230" pageId="228" pageNumber="229">
Legs moderately long and wide; femora superficially and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior longitudinal row of setae present; posterior ventral margin almost straight, not widened in apical half, posterior margin ventrally in apical half and dorsally completely serrated, glabrous. Metatibia moderately long and wide, dorsal and ventral margin subparallel, ratio width/length: 1/3.5, sharply carinate dorsally, with two groups of spines, basal one at basal third, apical one at three quarters of metatibial length, basally with a few robust and short setae; lateral face weakly longitudinally convex, with coarse and moderately dense punctures; ventral margin with two widely separated spines; medial face impunctate, apex shallowly and concavely emarginate interiorly near tarsal articulation. Tarsomeres sparsely finely punctate dorsally, with dense, fine setae ventrally; metatarsomeres lacking in
<typeStatus id="54AF88526B84FF20AB0CF8D1FBA63D99" box="[1042,1144,187,213]" pageId="229" pageNumber="230" type="holotype">holotype</typeStatus>
. Protibia short, bidentate. Anterior claws symmetrical.
</paragraph>
<paragraph id="8BAB36F06B84FF20AFD9F969FCB83C51" blockId="229.[151,1437,151,573]" box="[199,870,258,285]" pageId="229" pageNumber="230">
Aedeagus:
<figureCitation id="132F2A756B84FF20AE5AF969FE4B3C51" box="[324,405,258,285]" captionStart="FIGURE 52" captionStartId="224.[151,250,1933,1957]" captionTargetBox="[225,1362,288,1896]" captionTargetId="figure-64@224.[213,1375,273,1909]" captionTargetPageId="224" captionText="FIGURE 52. A–D: Maladera filigraniforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. flavipennis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. liwenzhui Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496448" httpUri="https://zenodo.org/record/4496448/files/figure.png" pageId="229" pageNumber="230">Fig. 52</figureCitation>
I-K. Habitus:
<figureCitation id="132F2A756B84FF20AD30F969FD4D3C51" box="[558,659,258,285]" captionStart="FIGURE 52" captionStartId="224.[151,250,1933,1957]" captionTargetBox="[225,1362,288,1896]" captionTargetId="figure-64@224.[213,1375,273,1909]" captionTargetPageId="224" captionText="FIGURE 52. A–D: Maladera filigraniforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. flavipennis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. liwenzhui Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496448" httpUri="https://zenodo.org/record/4496448/files/figure.png" pageId="229" pageNumber="230">
Fig.
<specimenCount id="9D12FD796B84FF20AD7CF968FD4D3C51" box="[610,659,258,285]" pageId="229" pageNumber="230" type="generic">52L</specimenCount>
</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B84FF20AFD9F94CFAD03C2A" pageId="229" pageNumber="230" type="diagnosis">
<paragraph id="8BAB36F06B84FF20AFD9F94CFAD03C2A" blockId="229.[151,1437,151,573]" pageId="229" pageNumber="230">
<emphasis id="B960EAE26B84FF20AFD9F94CFE9C3C0C" bold="true" box="[199,322,294,320]" pageId="229" pageNumber="230">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B84FF20AE57F94DFCEE3C0D" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[329,816,295,321]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="liwenzhui" status="sp. nov.">
<emphasis id="B960EAE26B84FF20AE57F94DFDF43C0D" box="[329,554,295,321]" italics="true" pageId="229" pageNumber="230">Maladera liwenzhui</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B84FF20AC22F94DFCA63C0D" box="[828,888,295,321]" pageId="229" pageNumber="230" rank="species">sp. n.</taxonomicNameLabel>
differs from all other Chinese
<taxonomicName id="4C144D736B84FF20ABCBF94DFA9D3C0D" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[1237,1347,295,321]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B84FF20ABCBF94DFA9D3C0D" box="[1237,1347,295,321]" italics="true" pageId="229" pageNumber="230">Maladera</emphasis>
</taxonomicName>
species in having the dorsal apex of the phallobase deeply incised (over more than half of phallobasal length).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B84FF20AFD9F905FF063CE1" pageId="229" pageNumber="230" type="etymology">
<paragraph id="8BAB36F06B84FF20AFD9F905FF063CE1" blockId="229.[151,1437,151,573]" pageId="229" pageNumber="230">
<emphasis id="B960EAE26B84FF20AFD9F905FE903CC5" bold="true" box="[199,334,367,393]" pageId="229" pageNumber="230">Etymology.</emphasis>
The new species is named after the Chinese entomologist Li Wenzhu (noun in genitive singular case).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B84FF20AFD9F9DCFBB73F55" pageId="229" pageNumber="230" type="description">
<paragraph id="8BAB36F06B84FF20AFD9F9DCFBB73F55" blockId="229.[151,1437,151,573]" pageId="229" pageNumber="230">
<emphasis id="B960EAE26B84FF20AFD9F9DCFEE13C9C" bold="true" box="[199,319,438,464]" pageId="229" pageNumber="230">Variation.</emphasis>
Length:
<quantity id="4CEC9B156B84FF20AEBFF9DDFD9D3C9D" box="[417,579,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.13" metricValueMin="1.1199999999999999" pageId="229" pageNumber="230" unit="mm" value="11.25" valueMax="11.3" valueMin="11.2">11.2–11.3 mm</quantity>
, length of elytra: 8.9–9.0 mm, width:
<quantity id="4CEC9B156B84FF20ACFDF9DDFBB23C9D" box="[995,1132,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.9" metricValueMin="6.2" pageId="229" pageNumber="230" unit="mm" value="6.55" valueMax="6.9" valueMin="6.2">6.2–6.9 mm</quantity>
. Metatarsomeres (only first three present) ventrally with a strongly serrated ridge, beside which is not a strong longitudinal carina, first metatarsomere little shorter than following two tarsomeres combined, dorsal tibial spur absent.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B84FF20AFD9FA48FD133F71" box="[199,717,546,573]" pageId="229" pageNumber="230" type="distribution">
<paragraph id="8BAB36F06B84FF20AFD9FA48FD133F71" blockId="229.[151,1437,151,573]" box="[199,717,546,573]" pageId="229" pageNumber="230">
<emphasis id="B960EAE26B84FF20AFD9FA48FEBF3F70" bold="true" box="[199,353,546,572]" pageId="229" pageNumber="230">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B84FF20AEC8FA49FDEB3F71" box="[470,565,547,573]" captionStart="FIGURE 117" captionStartId="360.[151,250,1034,1058]" captionTargetBox="[210,1379,187,1002]" captionTargetId="figure-19@360.[202,1385,181,1010]" captionTargetPageId="360" captionText="FIGURE 117. Distribution of the Maladera liwenzhui group: Maladera bikouensis sp. n., M. liwenzhui sp. n., M. shiwandashanensis sp. n., M. wupingensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496636" httpUri="https://zenodo.org/record/4496636/files/figure.png" pageId="229" pageNumber="230">Fig. 117</figureCitation>
) and
<tableCitation id="C696034B6B84FF20AD6DFA49FD193F71" box="[627,711,547,573]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="229" pageNumber="230">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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