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<mods:title id="F640FA3F0C16A6CDC4A5A884C958548F">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="E0BBA084B199C59FB5345F6B0258DDB1">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="359AF721A658E5C8042EE174E1A007E3">Bai, Ming</mods:namePart>
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<mods:namePart id="B0E1B8F2970C7FDCB7BB9C6EF5404627">Ahrens, Dirk</mods:namePart>
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<heading id="D0E3819C6B40FFE4AF89FEACFD113BAC" bold="true" box="[151,719,1734,1761]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<emphasis id="B960EAE26B40FFE4AF89FEACFD113BAC" bold="true" box="[151,719,1734,1761]" pageId="33" pageNumber="34">
<emphasis id="B960EAE26B40FFE4AF89FEACFED23BAD" bold="true" box="[151,268,1734,1761]" italics="true" pageId="33" pageNumber="34">Maladera</emphasis>
(subgen.
<taxonomicName id="4C144D736B40FFE4AE9AFEADFD193BAC" authority="Brenske, 1894" authorityName="Brenske" authorityYear="1894" box="[388,711,1734,1761]" class="Insecta" family="Melolonthidae" genus="Hemiserica" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B40FFE4AE9AFEADFDD13BAD" bold="true" box="[388,527,1735,1761]" italics="true" pageId="33" pageNumber="34">Hemiserica</emphasis>
<bibRefCitation id="EF854B016B40FFE4AD08FEADFD193BAC" author="Brenske, E." box="[534,711,1734,1761]" pageId="33" pageNumber="34" pagination="3 - 87" refId="ref331378" refString="Brenske, E. (1894) Die Melolonthiden der palaearktischen und orientalischen Region im Koniglichen naturhistorischen Museum zu Br ʾ ssel. Beschreibung neuer Arten und Bemerkung zu bekannten. Memoire de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
</taxonomicName>
)
</emphasis>
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<treatmentCitationGroup id="AB0411DE6B40FFE4AF89FF66FAE43A6F" box="[151,1338,1803,1827]" pageId="33" pageNumber="34">
<treatmentCitation id="0AB510E16B40FFE4AF89FF66FE143A6F" author="Brenske, E." box="[151,458,1803,1827]" page="11" pageId="33" pageNumber="34" year="1894">
<taxonomicName id="4C144D736B40FFE4AF89FF66FE143A6F" authority="Brenske, 1894: 11" authorityName="Brenske" authorityPageNumber="11" authorityYear="1894" box="[151,458,1803,1827]" class="Insecta" family="Melolonthidae" genus="Hemiserica" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B40FFE4AF89FF66FED23A6F" box="[151,268,1804,1827]" italics="true" pageId="33" pageNumber="34">Hemiserica</emphasis>
<bibRefCitation id="EF854B016B40FFE4AE0CFF61FE143A6F" author="Brenske, E." box="[274,458,1803,1827]" pageId="33" pageNumber="34" pagination="3 - 87" refId="ref331378" refString="Brenske, E. (1894) Die Melolonthiden der palaearktischen und orientalischen Region im Koniglichen naturhistorischen Museum zu Br ʾ ssel. Beschreibung neuer Arten und Bemerkung zu bekannten. Memoire de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894: 11</bibRefCitation>
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(
<typeStatus id="54AF88526B40FFE4AEC9FF61FDD53A6F" box="[471,523,1803,1827]" pageId="33" pageNumber="34">Type</typeStatus>
species (by monotypy):
<taxonomicName id="4C144D736B40FFE4AC18FF66FBBC3A6F" authority="Brenske, 1894" authorityName="Brenske" authorityYear="1894" box="[774,1122,1803,1827]" class="Insecta" family="Melolonthidae" genus="Hemiserica" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B960EAE26B40FFE4AC18FF66FC1B3A6F" box="[774,965,1804,1827]" italics="true" pageId="33" pageNumber="34">Hemiserica nasuta</emphasis>
<bibRefCitation id="EF854B016B40FFE4ACD5FF61FBBC3A6F" author="Brenske, E." box="[971,1122,1803,1827]" pageId="33" pageNumber="34" pagination="3 - 87" refId="ref331378" refString="Brenske, E. (1894) Die Melolonthiden der palaearktischen und orientalischen Region im Koniglichen naturhistorischen Museum zu Br ʾ ssel. Beschreibung neuer Arten und Bemerkung zu bekannten. Memoire de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="0AB510E16B40FFE4AB6CFF61FAEB3A6F" author="Ahrens, D." box="[1138,1333,1803,1827]" page="342" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="EF854B016B40FFE4AB6CFF61FAEB3A6F" author="Ahrens, D." box="[1138,1333,1803,1827]" pageId="33" pageNumber="34" refId="ref329734" refString="Ahrens, D. (2004 a) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation. de-Verlag im Internet GmbH, Berlin, 534 pp." type="book" year="2004">Ahrens 2004a: 342</bibRefCitation>
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.
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<paragraph id="8BAB36F06B40FFE4AF89FF40FF393A2D" blockId="33.[151,1437,1803,1919]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB0411DE6B40FFE4AF89FF40FF393A2D" pageId="33" pageNumber="34">
<taxonomicName id="4C144D736B40FFE4AF89FF40FE2A3A0E" authority="Frey, 1974: 120" authorityName="Frey" authorityPageNumber="120" authorityYear="1974" box="[151,500,1834,1858]" class="Insecta" family="Melolonthidae" genus="Rhynchosymmela" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B40FFE4AF89FF40FE963A0E" box="[151,328,1834,1858]" italics="true" pageId="33" pageNumber="34">Rhynchosymmela</emphasis>
Frey, 1974: 120
</taxonomicName>
(
<typeStatus id="54AF88526B40FFE4AD1AFF40FDEF3A0E" box="[516,561,1834,1858]" pageId="33" pageNumber="34">type</typeStatus>
species, by monotypy:
<taxonomicName id="4C144D736B40FFE4AC37FF40FB7B3A0E" authority="Frey, 1974" authorityName="Frey" authorityYear="1974" box="[809,1189,1834,1858]" class="Insecta" family="Melolonthidae" genus="Rhynchosymmela" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B960EAE26B40FFE4AC37FF40FBF43A0E" box="[809,1066,1834,1858]" italics="true" pageId="33" pageNumber="34">Rhynchosymmela pallida</emphasis>
Frey, 1974
</taxonomicName>
); syn by
<treatmentCitation id="0AB510E16B40FFE4AA1AFF40FF3D3A2D" author="Ahrens, D." page="32" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="EF854B016B40FFE4AA1AFF40FF3D3A2D" author="Ahrens, D." pageId="33" pageNumber="34" pagination="31 - 35" refId="ref329765" refString="Ahrens, D. (2004 b) Revisional notes on Sericini: The taxonomic status of the &quot; Neotropical &quot; genus Rhynchosymmela Frey, 1974. Beitrage zur Entomologie, 53, 31 - 35." type="journal article" year="2004">Ahrens 2004b: 32</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BAB36F06B40FFE4AF89FF0DFCD23A33" blockId="33.[151,1437,1803,1919]" box="[151,780,1895,1919]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB0411DE6B40FFE4AF89FF0DFCD23A33" box="[151,780,1895,1919]" pageId="33" pageNumber="34">
<taxonomicName id="4C144D736B40FFE4AF89FF0DFF253A33" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[151,251,1895,1919]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B40FFE4AF89FF0DFF253A33" box="[151,251,1895,1919]" italics="true" pageId="33" pageNumber="34">Maladera</emphasis>
</taxonomicName>
(subgen.
<taxonomicName id="4C144D736B40FFE4AE40FF02FE0D3A33" authorityName="Brenske" authorityYear="1894" box="[350,467,1896,1919]" class="Insecta" family="Melolonthidae" genus="Hemiserica" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B40FFE4AE40FF02FE0D3A33" box="[350,467,1896,1919]" italics="true" pageId="33" pageNumber="34">Hemiserica</emphasis>
</taxonomicName>
):
<treatmentCitation id="0AB510E16B40FFE4AEF9FF0DFCD63A33" author="Ahrens, D. &amp; Fabrizi, S." box="[487,776,1895,1919]" page="250" pageId="33" pageNumber="34" year="2016">
<bibRefCitation id="EF854B016B40FFE4AEF9FF0DFCD63A33" author="Ahrens, D. &amp; Fabrizi, S." box="[487,776,1895,1919]" pageId="33" pageNumber="34" pagination="1 - 355" refId="ref330518" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A Monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1 - 355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016: 250</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B43FFE7AF89F8F2FA433F1E" pageId="34" pageNumber="35" type="key">
<paragraph id="8BAB36F06B43FFE7AF89F8F2FBD63DF8" blockId="34.[151,1032,151,180]" box="[151,1032,151,180]" pageId="34" pageNumber="35">
<emphasis id="B960EAE26B43FFE7AF89F8F2FBD63DF8" bold="true" box="[151,1032,151,180]" pageId="34" pageNumber="35">
Key to species of
<taxonomicName id="4C144D736B43FFE7AE70F8FDFE3D3DFE" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[366,483,151,178]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B43FFE7AE70F8FDFE3D3DFE" bold="true" box="[366,483,151,178]" italics="true" pageId="34" pageNumber="35">Maladera</emphasis>
</taxonomicName>
(sbg.
<taxonomicName id="4C144D736B43FFE7AD30F8FDFD663DFD" authorityName="Brenske" authorityYear="1894" box="[558,696,151,177]" class="Insecta" family="Melolonthidae" genus="Hemiserica" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26B43FFE7AD30F8FDFD663DFD" bold="true" box="[558,696,151,177]" italics="true" pageId="34" pageNumber="35">Hemiserica</emphasis>
</taxonomicName>
) occurring in
<collectingCountry id="F30376606B43FFE7AC75F8FDFC663DFD" box="[875,952,151,177]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
(³³):
</emphasis>
</paragraph>
<key id="FF8561C16B43FFE7AF89F8B7FA433F1E" pageId="34" pageNumber="35">
<keyStep id="30E02E556B43FFE7AF89F8B7FA423C60" pageId="34" pageNumber="35">
<paragraph id="8BAB36F06B43FFE7AF89F8B7FA423DBE" blockId="34.[151,1437,221,595]" box="[151,1436,221,243]" pageId="34" pageNumber="35">
<keyLead id="30E595C56B43FFE7AF89F8B7FA423DBE" box="[151,1436,221,243]" pageId="34" pageNumber="35">
1 Body size larger,&gt;
<quantity id="4CEC9B156B43FFE7AE86F8B7FE0F3DBE" box="[408,465,221,242]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="34" pageNumber="35" unit="mm" value="9.0">9 mm</quantity>
. Dorsoapical median emargination of phallobase deep and narrow........................... 3
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B43FFE7AF89F890FA423C60" blockId="34.[151,1437,221,595]" pageId="34" pageNumber="35">
<keyLead id="30E595C56B43FFE7AF89F890FA423C60" pageId="34" pageNumber="35">
1’ Body size smaller, &lt;
<quantity id="4CEC9B156B43FFE7AEB6F890FE3F3C43" box="[424,481,250,271]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="35" unit="mm" value="8.0">8 mm</quantity>
. Labroclypeus finely and sparsely punctate, surface flat. Dorsoapical median emargination of phallobase shallow........................................................................................ 2
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</paragraph>
</keyStep>
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<keyLead id="30E595C56B43FFE7AF89F95FFA433C2B" pageId="34" pageNumber="35">
2 Ventral lobe of left paramere longer, distinctly exceeding the lobiform dorsal lobe................................................................................................
<taxonomicName id="4C144D736B43FFE7ACD8F938FABE3C24" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[966,1376,338,360]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rufopaca" status="sp. nov.">
<emphasis id="B960EAE26B43FFE7ACD8F938FB5C3C2B" box="[966,1154,338,359]" italics="true" pageId="34" pageNumber="35">Maladera rufopaca</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B43FFE7AA77F938FA433C2B" box="[1385,1437,338,359]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B43FFE7AF89F905FA433CEE" blockId="34.[151,1437,221,595]" pageId="34" pageNumber="35">
<keyLead id="30E595C56B43FFE7AF89F905FA433CEE" pageId="34" pageNumber="35">
2’ Ventral lobe of left paramere shorter, only little exceeding the lobiform dorsal lobe...........................................................................................
<taxonomicName id="4C144D736B43FFE7AC97F9E7FABE3CEF" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[905,1376,397,419]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="fangchengensis" status="sp. nov.">
<emphasis id="B960EAE26B43FFE7AC97F9E7FB5C3CEE" box="[905,1154,397,418]" italics="true" pageId="34" pageNumber="35">Maladera fangchengensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B43FFE7AA77F9E7FA433CEE" box="[1385,1437,397,418]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E556B43FFE7AF89F9C0FA433F1E" pageId="34" pageNumber="35">
<paragraph id="8BAB36F06B43FFE7AF89F9C0FA433CB6" blockId="34.[151,1437,221,595]" pageId="34" pageNumber="35">
<keyLead id="30E595C56B43FFE7AF89F9C0FA433CB6" pageId="34" pageNumber="35">
3 Labroclypeus coarsely and rugosely punctate, surface strongly convex. Basal piece of left paramere cylindrical; its narrow apical process fine and weakly bent (
<figureCitation id="132F2A756B43FFE7AD36F9ADFDAA3C91" box="[552,628,455,477]" captionStart="FIGURE 7" captionStartId="38.[151,250,1606,1630]" captionTargetBox="[161,1429,282,1575]" captionTargetId="figure-66@38.[151,1436,273,1583]" captionTargetPageId="38" captionText="FIGURE 7. A–D: Maladera hsui Ahrens, Fabrizi &amp; Liu, sp. n. (paratype); E–H: M. eshanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496330" httpUri="https://zenodo.org/record/4496330/files/figure.png" pageId="34" pageNumber="35">Fig. 7B</figureCitation>
). Right paramere weakly reflexed (lateral view,
<figureCitation id="132F2A756B43FFE7AB5CF9ADFB533C91" box="[1090,1165,455,477]" captionStart="FIGURE 7" captionStartId="38.[151,250,1606,1630]" captionTargetBox="[161,1429,282,1575]" captionTargetId="figure-66@38.[151,1436,273,1583]" captionTargetPageId="38" captionText="FIGURE 7. A–D: Maladera hsui Ahrens, Fabrizi &amp; Liu, sp. n. (paratype); E–H: M. eshanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496330" httpUri="https://zenodo.org/record/4496330/files/figure.png" pageId="34" pageNumber="35">Fig. 7C</figureCitation>
).......................................................................................
<taxonomicName id="4C144D736B43FFE7ACE8F98FFABE3CB7" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[1014,1376,485,507]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hsui" status="sp. nov.">
<emphasis id="B960EAE26B43FFE7ACE8F98FFB5D3CB6" box="[1014,1155,485,506]" italics="true" pageId="34" pageNumber="35">Maladera hsui</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B43FFE7AA77F98FFA433CB6" box="[1385,1437,485,506]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAB36F06B43FFE7AF89FA68FA433F1E" blockId="34.[151,1437,221,595]" pageId="34" pageNumber="35">
<keyLead id="30E595C56B43FFE7AF89FA68FA433F1E" pageId="34" pageNumber="35">
3’ Labroclypeus finely punctate, surface nearly flat. Basal piece of left paramere conically narrowed; its narrow apical process thicker and strongly bent (
<figureCitation id="132F2A756B43FFE7AEC0FA75FDF63F79" box="[478,552,543,565]" captionStart="FIGURE 7" captionStartId="38.[151,250,1606,1630]" captionTargetBox="[161,1429,282,1575]" captionTargetId="figure-66@38.[151,1436,273,1583]" captionTargetPageId="38" captionText="FIGURE 7. A–D: Maladera hsui Ahrens, Fabrizi &amp; Liu, sp. n. (paratype); E–H: M. eshanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496330" httpUri="https://zenodo.org/record/4496330/files/figure.png" pageId="34" pageNumber="35">Fig. 7F</figureCitation>
). Right paramere strongly reflexed (lateral view,
<figureCitation id="132F2A756B43FFE7ACE1FA75FB923F79" box="[1023,1100,543,565]" captionStart="FIGURE 7" captionStartId="38.[151,250,1606,1630]" captionTargetBox="[161,1429,282,1575]" captionTargetId="figure-66@38.[151,1436,273,1583]" captionTargetPageId="38" captionText="FIGURE 7. A–D: Maladera hsui Ahrens, Fabrizi &amp; Liu, sp. n. (paratype); E–H: M. eshanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, F: parameres, dorsal view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496330" httpUri="https://zenodo.org/record/4496330/files/figure.png" pageId="34" pageNumber="35">Fig. 7G</figureCitation>
)........................................................................................
<taxonomicName id="4C144D736B43FFE7ACABFA57FABE3F1F" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[949,1376,573,595]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="eshanensis" status="sp. nov.">
<emphasis id="B960EAE26B43FFE7ACABFA57FB5C3F1E" box="[949,1154,573,594]" italics="true" pageId="34" pageNumber="35">Maladera eshanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B43FFE7AA77FA57FA433F1E" box="[1385,1437,573,594]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
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</subSubSection>
</treatment>
</document>
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