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<document id="6F844048F562C52BC58102E4AA678B23" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66A31FE97AF89FDA0FD103931" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera nabanensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="339" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="337" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="BE4DC650ED58775626ADCB27CBF28B68" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="55BB854AB9094BE45E05AEFFE197D4F5">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="D6DA1CB53C69B28155EA3C610A8A4C9D">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="DA77086F8CA023CE0210656479C85693" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="CE7F74ACE4F5C49AD84967D64158FB82" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:name id="7165910B9B80D6CDEE99EA338620A117" type="personal">
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</mods:role>
<mods:namePart id="CB47169891C904650C1EFD5A69AC9668">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="5E65E403F615E0389F36C91C2BB45705" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="701B47E1F629542F351977464C1AC538" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="C72F2E015028C6B61C47A97DF4C60988" type="personal">
<mods:role id="9FF645D0C3104C742718E0714C3C7D85">
<mods:roleTerm id="516F3D2CEA565E3A0C1A0E8B95ACAFF6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6CB13720197CE3AAAA7739F1117F346D">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="E8DE43F81BF6A8EC5F4389842E6527A9" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="2D410E872F0C0E068BEE2AC3E01A4E6C" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="B245E887E47B3E9A893598A09E52F8D5" type="personal">
<mods:role id="8169DE65DE0B8939853BEDC4A3A16050">
<mods:roleTerm id="136781007B537D7536C916D381310AA0">Author</mods:roleTerm>
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<mods:namePart id="EB23A207A4625071E2BEEB8C01ACD8B6">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="D327560ED519A795EC387001A2ADCC39" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="D2BB2D976E4E519D0B0BC129A18F3B04" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:name id="C6DBBC7E1B1DA63D4F807A8AB5821BFB" type="personal">
<mods:role id="0AA035B0C9F33CF54D71B4C6162954EF">
<mods:roleTerm id="C1DA1702523D6BE9244917415B36758F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27EB9F9360C387C99C71D93116C5177D">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="BC35111AAE172E1D3E35F8C6C9643016" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="44928AFFD3B2FB8D192C6146F411FC8D" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:typeOfResource id="65B1A4224D89EFF1EEA83340999BD551">text</mods:typeOfResource>
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<mods:title id="D573A08DAC8260C06CB593DF90AA82DD">Zootaxa</mods:title>
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<mods:part id="6A9D6CF6DF4BFFAC285C0C8D6B724637">
<mods:date id="15CD4250537DF12A6715A59C99F9F937">2021</mods:date>
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<mods:number id="CBB89D40400F26217814FA3148D9E968">4922</mods:number>
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<mods:classification id="3852EA773108FFC8D51A0E4207BBFA98">journal article</mods:classification>
<mods:identifier id="B0063ACE47A66B91CF26CE2A457E9DAB" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="0DE54BFCCD9B1897E784BB27C0E36C01" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="5E08D9472A6956D1F6DA49E5B1AAFFFE" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
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<treatment id="03BD87E66A31FE97AF89FDA0FD103931" ID-DOI="http://doi.org/10.5281/zenodo.4534987" ID-GBIF-Taxon="177323741" ID-Zenodo-Dep="4534987" LSID="urn:lsid:plazi:treatment:03BD87E66A31FE97AF89FDA0FD103931" httpUri="http://treatment.plazi.org/id/03BD87E66A31FE97AF89FDA0FD103931" lastPageId="338" lastPageNumber="339" pageId="336" pageNumber="337">
<subSubSection id="C30E657B6A31FE95AF89FDA0FCD538A9" box="[151,779,1482,1509]" pageId="336" pageNumber="337" type="nomenclature">
<paragraph id="8BAB36F06A31FE95AF89FDA0FCD538A9" blockId="336.[151,779,1482,1545]" box="[151,779,1482,1509]" pageId="336" pageNumber="337">
<heading id="D0E3819C6A31FE95AF89FDA0FCD538A9" bold="true" box="[151,779,1482,1509]" fontSize="11" level="2" pageId="336" pageNumber="337" reason="4">
<emphasis id="B960EAE26A31FE95AF89FDA0FCD538A9" bold="true" box="[151,779,1482,1509]" pageId="336" pageNumber="337">
<taxonomicName id="4C144D736A31FE95AF89FDA0FD6238A8" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,700,1482,1509]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="336" pageNumber="337" phylum="Arthropoda" rank="species" species="nabanensis" status="sp. nov.">
<emphasis id="B960EAE26A31FE95AF89FDA0FE4338A9" bold="true" box="[151,413,1482,1509]" italics="true" pageId="336" pageNumber="337">Maladera nabanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A31FE95ADD7FDA1FCD538A9" box="[713,779,1483,1509]" pageId="336" pageNumber="337" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A31FE95AF89FD85FEA33B45" box="[151,381,1519,1545]" pageId="336" pageNumber="337" type="description">
<paragraph id="8BAB36F06A31FE95AF89FD85FEA33B45" blockId="336.[151,779,1482,1545]" box="[151,381,1519,1545]" pageId="336" pageNumber="337">
<figureCitation id="132F2A756A31FE95AF89FD85FED23B45" box="[151,268,1519,1545]" captionStart="FIGURE 81" captionStartId="337.[151,250,1744,1768]" captionTargetBox="[162,1425,283,1712]" captionTargetId="figure-50@337.[151,1436,273,1721]" captionTargetPageId="337" captionText="FIGURE 81. A–D: Maladera nabanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. luoxiangensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, E: parameres, dorsal view; F: aedeagus, ventral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496534" httpUri="https://zenodo.org/record/4496534/files/figure.png" pageId="336" pageNumber="337">Figures 81</figureCitation>
A–D, 129
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A31FE95AF89FE5AFD4A3B96" pageId="336" pageNumber="337" type="materials_examined">
<paragraph id="8BAB36F06A31FE95AF89FE5AFD4A3B96" blockId="336.[151,1437,1584,2043]" pageId="336" pageNumber="337">
<emphasis id="B960EAE26A31FE95AF89FE5AFE1F3B06" bold="true" box="[151,449,1584,1610]" pageId="336" pageNumber="337">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6A31FE95AED7FE5AFAC43B22" ID-GBIF-Occurrence="3034556806" collectingDate="2008-05-20" collectionCode="NME" collectorName="A. Weigel" elevation="680" latitude="22.167334" location="China" longLatPrecision="12" longitude="100.65867" pageId="336" pageNumber="337" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54AF88526A31FE95AED7FE5AFDE83B06" box="[457,566,1584,1610]" pageId="336" pageNumber="337" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796A31FE95AD5FFE5BFD883B07" box="[577,598,1585,1611]" pageId="336" pageNumber="337" type="male">♂</specimenCount>
“
<location id="8ECB602B6A31FE95AD76FE5AFD6E3B06" LSID="urn:lsid:plazi:treatment:03BD87E66A31FE97AF89FDA0FD103931:8ECB602B6A31FE95AD76FE5AFD6E3B06" box="[616,688,1584,1610]" latitude="22.167334" longLatPrecision="12" longitude="100.65867" name="China" pageId="336" pageNumber="337">China</location>
, S-Yunnan Prov. Xishuangbanna
<locationDeviation id="9A192FD26A31FE95AB35FE5AFAB33B07" box="[1067,1389,1584,1611]" location="Jinghong Na Ban" pageId="336" pageNumber="337">
<quantity id="4CEC9B156A31FE95AB35FE5AFBB43B06" box="[1067,1130,1584,1610]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="336" pageNumber="337" unit="km" value="23.0">23km</quantity>
NW Jinghong Na Ban
</locationDeviation>
village
<quantity id="4CEC9B156A31FE95AFCEFE3EFED13B22" box="[208,271,1620,1646]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="336" pageNumber="337" unit="m" value="680.0">
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</quantity>
<geoCoordinate id="EE2050376A31FE95AE07FE3EFE433B22" box="[281,413,1620,1646]" degrees="22" direction="north" minutes="10.04" orientation="latitude" pageId="336" pageNumber="337" precision="9" value="22.167334">22°10.04’N</geoCoordinate>
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,
<date id="FFAA10306A31FE95AD58FE3EFD623B22" box="[582,700,1620,1646]" pageId="336" pageNumber="337" value="2008-05-20">
<collectingDate id="EFEEE9D86A31FE95AD58FE3EFD623B22" box="[582,700,1620,1646]" pageId="336" pageNumber="337" value="2008-05-20">20.V.2008</collectingDate>
</date>
, leg.
<collectorName id="26E153266A31FE95ADE5FE3EFCAF3B22" box="[763,881,1620,1646]" pageId="336" pageNumber="337">A. Weigel</collectorName>
, LF/ 785
<taxonomicName id="4C144D736A31FE95ACC4FE3EFBEC3B23" box="[986,1074,1620,1647]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="336" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356A31FE95ABD0FE3EFACC3B22" box="[1230,1298,1620,1646]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="336" pageNumber="337">NME</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A31FE95AA36FE3FFA903BDE" ID-GBIF-Occurrence="3034555767" collectingDate="1956-05-12" collectionCode="IZAS" collectorName="Huang Keren" country="China" county="China" elevation="1650" location="Jinping" municipality="Hetouzhai" pageId="336" pageNumber="337" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54AF88526A31FE95AA36FE3FFA473B23" box="[1320,1433,1621,1647]" pageId="336" pageNumber="337" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B960EAE26A31FE95AF89FE12FF343BDE" bold="true" box="[151,234,1656,1682]" pageId="336" pageNumber="337">
<collectingCountry id="F30376606A31FE95AF89FE12FF3A3BDE" box="[151,228,1656,1682]" name="China" pageId="336" pageNumber="337">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A31FE95AFEFFE12FEC53BDF" box="[241,283,1656,1683]" pageId="336" pageNumber="337" type="male">1 ♂</specimenCount>
“[
<collectingCounty id="62CA4E7C6A31FE95AE2AFE1DFEA03BDE" box="[308,382,1655,1682]" pageId="336" pageNumber="337">China</collectingCounty>
]
<collectingMunicipality id="6BCFAC8A6A31FE95AE92FE12FDDC3BDE" box="[396,514,1656,1682]" pageId="336" pageNumber="337">Hetouzhai</collectingMunicipality>
,
<location id="8ECB602B6A31FE95AD10FE12FDBD3BDE" LSID="urn:lsid:plazi:treatment:03BD87E66A31FE97AF89FDA0FD103931:8ECB602B6A31FE95AD10FE12FDBD3BDE" box="[526,611,1656,1682]" country="China" county="China" municipality="Hetouzhai" name="Jinping" pageId="336" pageNumber="337" stateProvince="Yunnan">Jinping</location>
,
<collectingRegion id="49D0F8126A31FE95AD70FE13FD193BDF" box="[622,711,1657,1683]" country="China" name="Yunnan" pageId="336" pageNumber="337">Yunnan</collectingRegion>
,
<date id="FFAA10306A31FE95ADCCFE12FC993BDF" box="[722,839,1656,1683]" pageId="336" pageNumber="337" value="1956-05-12">
<collectingDate id="EFEEE9D86A31FE95ADCCFE12FC993BDF" box="[722,839,1656,1683]" pageId="336" pageNumber="337" value="1956-05-12">12.V.1956</collectingDate>
</date>
,
<quantity id="4CEC9B156A31FE95AC4DFE12FC3A3BDE" box="[851,996,1656,1682]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="336" pageNumber="337" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="0039D1C36A31FE95AC4DFE12FC3A3BDE" box="[851,996,1656,1682]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="336" pageNumber="337" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600-1700m</elevation>
</quantity>
, leg.
<collectorName id="26E153266A31FE95AB3FFE13FB693BDF" box="[1057,1207,1657,1683]" pageId="336" pageNumber="337">Huang Keren</collectorName>
etc.” (
<collectionCode id="ED05AE356A31FE95AA1CFE12FA983BDE" box="[1282,1350,1656,1682]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="336" pageNumber="337">IZAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A31FE95AA47FE12FD4E3B96" ID-GBIF-Occurrence="3034555390" collectingDate="1997-05-13" collectingDateMax="1997-05-24" collectingDateMin="1997-05-13" collectionCode="ZFMK" collectorName="E. Jendek &amp; O. Sausa" country="Laos" elevation="100" location="Laos" pageId="336" pageNumber="337" specimenCount="1" specimenCount-male="1" stateProvince="Louangnamtha" typeStatus="paratype">
<emphasis id="B960EAE26A31FE95AA47FE12FA423BDE" bold="true" box="[1369,1436,1656,1682]" pageId="336" pageNumber="337">
<collectingCountry id="F30376606A31FE95AA47FE12FA493BDE" box="[1369,1431,1656,1682]" name="Laos" pageId="336" pageNumber="337">Laos</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796A31FE95AF89FEF6FF1C3BFB" box="[151,194,1692,1719]" pageId="336" pageNumber="337" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6A31FE95AFCAFEF6FED33BFA" LSID="urn:lsid:plazi:treatment:03BD87E66A31FE97AF89FDA0FD103931:8ECB602B6A31FE95AFCAFEF6FED33BFA" box="[212,269,1692,1718]" country="Laos" name="Laos" pageId="336" pageNumber="337" stateProvince="Louangnamtha">Laos</location>
north,
<date id="FFAA10306A31FE95AE40FEF6FE233BFA" box="[350,509,1692,1718]" pageId="336" pageNumber="337" value="1997-05-13" valueMax="1997-05-24" valueMin="1997-05-13">
<collectingDate id="EFEEE9D86A31FE95AE40FEF6FE233BFA" box="[350,509,1692,1718]" pageId="336" pageNumber="337" value="1997-05-13" valueMax="1997-05-24" valueMin="1997-05-13">13.-24.V.1997</collectingDate>
</date>
<quantity id="4CEC9B156A31FE95AD1BFEF6FD933BFA" box="[517,589,1692,1719]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="336" pageNumber="337" unit="km" value="15.0">15 km</quantity>
NW
<collectingRegion id="49D0F8126A31FE95AD94FEF7FC9E3BFA" box="[650,832,1692,1719]" country="Laos" name="Louangnamtha" pageId="336" pageNumber="337">Louang Namtha</collectingRegion>
N21°07.5, E101°21.0 alt. 750+
<quantity id="4CEC9B156A31FE95ABB9FEF6FB393BFB" box="[1191,1255,1692,1719]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="336" pageNumber="337" unit="m" value="100.0">
<elevation id="0039D1C36A31FE95ABB9FEF6FB393BFB" box="[1191,1255,1692,1719]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="336" pageNumber="337" unit="m" value="100.0">100m</elevation>
</quantity>
<collectorName id="26E153266A31FE95ABF0FEF7FA823BFA" box="[1262,1372,1692,1719]" pageId="336" pageNumber="337">E. Jendek</collectorName>
&amp;
<collectorName id="26E153266A31FE95AA9FFEF6FF063B90" pageId="336" pageNumber="337">O. Šauša</collectorName>
leg./ 367
<taxonomicName id="4C144D736A31FE95AE57FEAAFE7E3B96" box="[329,416,1728,1754]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="336" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356A31FE95AD2DFEAAFD593B96" box="[563,647,1728,1754]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="336" pageNumber="337" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A31FE97AFD9FE8EFCA93EE9" lastPageId="338" lastPageNumber="339" pageId="336" pageNumber="337" type="description">
<paragraph id="8BAB36F06A31FE95AFD9FE8EFCA33A6F" blockId="336.[151,1437,1584,2043]" pageId="336" pageNumber="337">
<emphasis id="B960EAE26A31FE95AFD9FE8EFE873BB2" bold="true" box="[199,345,1764,1790]" pageId="336" pageNumber="337">Description.</emphasis>
Length:
<quantity id="4CEC9B156A31FE95AEA5FE8EFDCC3BB3" box="[443,530,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="336" pageNumber="337" unit="mm" value="5.4">5.4 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156A31FE95ADCDFE8EFCF43BB3" box="[723,810,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="336" pageNumber="337" unit="mm" value="3.6">3.6 mm</quantity>
, width:
<quantity id="4CEC9B156A31FE95AC9FFE8EFC073BB3" box="[897,985,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="336" pageNumber="337" unit="mm" value="3.1">3.1 mm</quantity>
. Body oblong-oval, dark brown, antenna yellowish, dorsal surface shiny, dorsal face shortly densely setose.
</paragraph>
<paragraph id="8BAB36F06A31FE94AFD9FF46FF213DB5" blockId="336.[151,1437,1584,2043]" lastBlockId="337.[151,1436,151,249]" lastPageId="337" lastPageNumber="338" pageId="336" pageNumber="337">
Labroclypeus narrow and subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles, lateral margin and ocular canthus producing a blunt angle; margins weakly reflexed; anterior margin weakly convex medially; surface moderately convex, shiny, finely and moderately densely punctate, glabrous; frontoclypeal suture weakly impressed and curved medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, finely punctate and with minute setae, without terminal seta. Frons with fine, very dense punctures, glabrous except for minute setae in punctures. Eyes extremely large, ratio of diameter/interocular width: 1.12. Antenna with ten antennomeres; club with three antennomeres, twice as long as remaining antennomeres combined, reflexed.
<collectingRegion id="49D0F8126A30FE94AB29F8D1FB483D99" box="[1079,1174,187,213]" country="Myanmar" name="Chin" pageId="337" pageNumber="338">Mentum</collectingRegion>
anteriorly elevated and flattened.
</paragraph>
<caption id="DF6B66786A30FE94AF89FEBAFC6E3A7C" ID-DOI="http://doi.org/10.5281/zenodo.4496534" ID-Zenodo-Dep="4496534" httpUri="https://zenodo.org/record/4496534/files/figure.png" pageId="337" pageNumber="338" startId="337.[151,250,1744,1768]" targetBox="[162,1425,283,1712]" targetPageId="337">
<paragraph id="8BAB36F06A30FE94AF89FEBAFC6E3A7C" blockId="337.[151,1437,1744,1840]" pageId="337" pageNumber="338">
<emphasis id="B960EAE26A30FE94AF89FEBAFE813BA4" bold="true" box="[151,351,1744,1768]" pageId="337" pageNumber="338">FIGURE 81. A–D:</emphasis>
<taxonomicName id="4C144D736A30FE94AE78FEBAFCF13BA4" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[358,815,1744,1768]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="337" pageNumber="338" phylum="Arthropoda" rank="species" species="nabanensis" status="sp. nov.">
<emphasis id="B960EAE26A30FE94AE78FEBAFD9D3BA4" box="[358,579,1744,1768]" italics="true" pageId="337" pageNumber="338">Maladera nabanensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A30FE94AC24FEBBFCAF3BA4" box="[826,881,1745,1768]" pageId="337" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
(holotype);
<emphasis id="B960EAE26A30FE94ACF1FEBAFBF63BA4" bold="true" box="[1007,1064,1744,1768]" pageId="337" pageNumber="338">E–H:</emphasis>
<taxonomicName id="4C144D736A30FE94AB31FEBBFF633A40" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="337" pageNumber="338" phylum="Arthropoda" rank="species" species="luoxiangensis" status="sp. nov.">
<emphasis id="B960EAE26A30FE94AB31FEBBFB023BA4" box="[1071,1244,1744,1768]" italics="true" pageId="337" pageNumber="338">M. luoxiangensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A30FE94AFD9FE9FFF233A40" box="[199,253,1781,1804]" pageId="337" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
(holotype).
<emphasis id="B960EAE26A30FE94AE66FE9EFE4D3A40" bold="true" box="[376,403,1780,1804]" pageId="337" pageNumber="338">A:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B960EAE26A30FE94ADC3FE9EFCC93A40" bold="true" box="[733,791,1780,1804]" pageId="337" pageNumber="338">C, G:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B960EAE26A30FE94AB71FE9EFB7B3A40" bold="true" box="[1135,1189,1780,1804]" pageId="337" pageNumber="338">B, E:</emphasis>
parameres, dorsal view;
<emphasis id="B960EAE26A30FE94AF89FF72FF713A7C" bold="true" box="[151,175,1816,1840]" pageId="337" pageNumber="338">F:</emphasis>
aedeagus, ventral view;
<emphasis id="B960EAE26A30FE94AEB4FF72FE3B3A7C" bold="true" box="[426,485,1816,1840]" pageId="337" pageNumber="338">D, H:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06A30FE94AFD9FF32FB523A92" blockId="337.[151,1437,1880,2014]" pageId="337" pageNumber="338">Pronotum subrectangular, widest at base, lateral margins weakly convex and weakly convergent anteriorly; anterior angles produced and sharp, posterior angles blunt; anterior margin weakly convexly produced medially, marginal line complete; surface finely and densely punctate, with short dense setae; anterior and lateral margins glabrous. Scutellum wide, triangular, with fine and dense punctures, and short dense setae.</paragraph>
<paragraph id="8BAB36F06A33FE97AFD9F8FDFBB63DB5" blockId="338.[151,1437,151,1149]" pageId="338" pageNumber="339">Elytra oblong, widest at middle, striae weakly impressed, finely and densely punctate, intervals flat, with fine, dense punctures and dense short setae; epipleural margin fine, ending at convex external apical angle of elytra, epipleura densely setose; apical border membraneous, with a rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06A33FE97AFD9F969FB5D3C9D" blockId="338.[151,1437,151,1149]" pageId="338" pageNumber="339">Ventral surface with fine, dense punctures, densely shortly setose; metacoxa also shortly setose including several longer lateral setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/2.0. Abdominal sternites, in addition to fine and dense punctures with minute setae, each sternite with a distinct transverse row of coarse punctures each bearing a short and robust seta; penultimate sternite apically with a shiny smooth chitinous border, which is one quarter as long as sternite. Pygidium weakly convex, finely and densely punctate, without smooth midline; with dense short setae as in the rest of surface.</paragraph>
<paragraph id="8BAB36F06A33FE97AFD9F9B1FE4D3ECD" blockId="338.[151,1437,151,1149]" pageId="338" pageNumber="339">Legs short and wide; femora with two longitudinal rows of setae, finely and densely punctate, densely shortly setose; metafemur dull, anterior margin acute, with complete adjacent serrated line, posterior ventral margin widened in apical half, smooth, not serrate, dorsal posterior margin not serrated, finely setose. Metatibia wide and short, widest at apex, ratio width/length: 1/2.6, dorsal margin sharply carinate, with a continuously serrated line over its entire length; with two groups of spines, basal one at middle, apical one at three quarters of metatibial length; in basal half with a few robust single setae; lateral face weakly convex, with dense and fine punctures, and short setae in punctures; ventral margin serrate, with five equidistant robust and very long spines; medial face finely impunctate and glabrous; apex interiorly near tarsal articulation shallowly concave. Tarsomeres very sparsely punctate and setose dorsally, with dense, short setae ventrally; metatarsomeres ventrally with a strongly serrated ridge, beside which is a fine carina; first metatarsomere a little shorter than two following tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="8BAB36F06A33FE97AFD9FBE1FCA93EE9" blockId="338.[151,1437,151,1149]" box="[199,887,907,933]" pageId="338" pageNumber="339">
Aedeagus:
<figureCitation id="132F2A756A33FE97AE5AFBE1FE4C3EE9" box="[324,402,907,933]" captionStart="FIGURE 81" captionStartId="337.[151,250,1744,1768]" captionTargetBox="[162,1425,283,1712]" captionTargetId="figure-50@337.[151,1436,273,1721]" captionTargetPageId="337" captionText="FIGURE 81. A–D: Maladera nabanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. luoxiangensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, E: parameres, dorsal view; F: aedeagus, ventral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496534" httpUri="https://zenodo.org/record/4496534/files/figure.png" pageId="338" pageNumber="339">Fig. 81</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756A33FE97AD23FBE1FD7D3EE9" box="[573,675,907,933]" captionStart="FIGURE 81" captionStartId="337.[151,250,1744,1768]" captionTargetBox="[162,1425,283,1712]" captionTargetId="figure-50@337.[151,1436,273,1721]" captionTargetPageId="337" captionText="FIGURE 81. A–D: Maladera nabanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. luoxiangensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A: aedeagus, left side lateral view; C, G: aedeagus, right side lateral view; B, E: parameres, dorsal view; F: aedeagus, ventral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496534" httpUri="https://zenodo.org/record/4496534/files/figure.png" pageId="338" pageNumber="339">Fig. 81D</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A33FE97AFD9FBC4FB883EA1" pageId="338" pageNumber="339" type="diagnosis">
<paragraph id="8BAB36F06A33FE97AFD9FBC4FB883EA1" blockId="338.[151,1437,151,1149]" pageId="338" pageNumber="339">
<emphasis id="B960EAE26A33FE97AFD9FBC4FE9C3E84" bold="true" box="[199,322,942,968]" pageId="338" pageNumber="339">Diagnosis.</emphasis>
This new species differs from all other Chinese
<taxonomicName id="4C144D736A33FE97AC74FBC5FC063E85" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[874,984,943,969]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="338" pageNumber="339" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26A33FE97AC74FBC5FC063E85" box="[874,984,943,969]" italics="true" pageId="338" pageNumber="339">Maladera</emphasis>
</taxonomicName>
species in having extremely large eyes, short dense dorsal pilosity, and a serrated line along the entire length of the metatibia.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A33FE97AFD9FB9DFF3C397A" pageId="338" pageNumber="339" type="etymology">
<paragraph id="8BAB36F06A33FE97AFD9FB9DFF3C397A" blockId="338.[151,1437,151,1149]" pageId="338" pageNumber="339">
<emphasis id="B960EAE26A33FE97AFD9FB9DFE90395D" bold="true" box="[199,334,1015,1041]" pageId="338" pageNumber="339">Etymology.</emphasis>
The new species is named after its
<typeStatus id="54AF88526A33FE97ADFDFB9DFCCD395D" box="[739,787,1015,1041]" pageId="338" pageNumber="339">type</typeStatus>
locality, Na Ban village (adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A33FE97AFD9FC54FBBA3915" box="[199,1124,1086,1113]" pageId="338" pageNumber="339" type="description">
<paragraph id="8BAB36F06A33FE97AFD9FC54FBBA3915" blockId="338.[151,1437,151,1149]" box="[199,1124,1086,1113]" pageId="338" pageNumber="339">
<emphasis id="B960EAE26A33FE97AFD9FC54FEE13914" bold="true" box="[199,319,1086,1112]" pageId="338" pageNumber="339">Variation.</emphasis>
Length: 5.4–6.0 mm, length of elytra: 3.6–4.0 mm, width:
<quantity id="4CEC9B156A33FE97ACCBFC55FB813915" box="[981,1119,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="338" pageNumber="339" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A33FE97AFD9FC08FD103931" box="[199,718,1122,1149]" pageId="338" pageNumber="339" type="distribution">
<paragraph id="8BAB36F06A33FE97AFD9FC08FD103931" blockId="338.[151,1437,151,1149]" box="[199,718,1122,1149]" pageId="338" pageNumber="339">
<emphasis id="B960EAE26A33FE97AFD9FC08FEBF3930" bold="true" box="[199,353,1122,1148]" pageId="338" pageNumber="339">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756A33FE97AEC8FC09FDE83931" box="[470,566,1123,1149]" captionStart="FIGURE 129" captionStartId="366.[151,250,1001,1025]" captionTargetBox="[231,1354,187,971]" captionTargetId="figure-19@366.[226,1362,181,977]" captionTargetPageId="366" captionText="FIGURE 129. Distribution of Maladera species with uncertain systematic status: Maladera baishaoensis sp. n., M. hutiaoensis sp. n., M. jinggangshanica sp. n., M. lukjanovitschi (Medvedev, 1966), M. luoxiangensis sp. n., M. nabanensis sp. n., M. nanlingensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496668" httpUri="https://zenodo.org/record/4496668/files/figure.png" pageId="338" pageNumber="339">Fig. 129</figureCitation>
) and
<tableCitation id="C696034B6A33FE97AD6AFC09FD163931" box="[628,712,1123,1149]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="338" pageNumber="339">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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