215 lines
24 KiB
XML
215 lines
24 KiB
XML
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<mods:title id="3B42D97F2826890747535F28585EC7F8">A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="FF3B013853D4BA750BA9A94F8EE80846">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="A34DC59996B516460D165E708BB71373">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="999CEA35A586CA0CDC59A4809F5EB8C9">Bai, Ming</mods:namePart>
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<mods:namePart id="BDFA158EA212094F87B01E3AFC8FCDA5">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="1C3888E19C78BF62F5FF542EF11515AC">Ahrens, Dirk</mods:namePart>
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<mods:title id="1A1D66A84F935C9B761417E445AF2CE6">Zootaxa</mods:title>
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<mods:number id="022016814EA7D99AB7459CF11F129A89">2021-02-03</mods:number>
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<treatment id="03BD87E66A28FE8EAF89FB0CFD103FE5" ID-DOI="http://doi.org/10.5281/zenodo.4534954" ID-GBIF-Taxon="177323737" ID-Zenodo-Dep="4534954" LSID="urn:lsid:plazi:treatment:03BD87E66A28FE8EAF89FB0CFD103FE5" httpUri="http://treatment.plazi.org/id/03BD87E66A28FE8EAF89FB0CFD103FE5" lastPageId="331" lastPageNumber="332" pageId="329" pageNumber="330">
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<emphasis id="B960EAE26A28FE8CAF89FB0CFCC63ECD" bold="true" box="[151,792,870,897]" pageId="329" pageNumber="330">
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<taxonomicName id="4C144D736A28FE8CAF89FB0CFD173ECC" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[151,713,870,897]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="329" pageNumber="330" phylum="Arthropoda" rank="species" species="wandingana" status="sp. nov.">
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<emphasis id="B960EAE26A28FE8CAF89FB0CFE743ECD" bold="true" box="[151,426,870,897]" italics="true" pageId="329" pageNumber="330">Maladera wandingana</emphasis>
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Ahrens, Fabrizi & Liu
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25357996A28FE8CADC8FB0DFCC63ECD" box="[726,792,871,897]" pageId="329" pageNumber="330" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C30E657B6A28FE8CAF89FBE1FEA33EE9" box="[151,381,907,933]" pageId="329" pageNumber="330" type="description">
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<paragraph id="8BAB36F06A28FE8CAF89FBE1FEA33EE9" blockId="329.[151,792,870,933]" box="[151,381,907,933]" pageId="329" pageNumber="330">
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<figureCitation id="132F2A756A28FE8CAF89FBE1FED03EE9" box="[151,270,907,933]" captionStart="FIGURE 79" captionStartId="330.[151,250,1943,1967]" captionTargetBox="[172,1417,198,1910]" captionTargetId="figure-19@330.[156,1431,181,1920]" captionTargetPageId="330" captionText="FIGURE 79. A–D: Maladera wandingana Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. trifidiforceps Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. nanlingensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale" figureDoi="http://doi.org/10.5281/zenodo.4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="329" pageNumber="330">Figures 79</figureCitation>
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A–D, 130
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E657B6A28FE8CAF89FBB9FEE5395D" pageId="329" pageNumber="330" type="materials_examined">
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<paragraph id="8BAB36F06A28FE8CAF89FBB9FEE5395D" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">
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<emphasis id="B960EAE26A28FE8CAF89FBB9FE613EA0" bold="true" box="[151,447,978,1005]" pageId="329" pageNumber="330">Type material examined.</emphasis>
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<materialsCitation id="3B7C3CAD6A28FE8CAED8FBB9FEE5395D" ID-GBIF-Occurrence="3034555987" collectingDate="2011-04-29" collectionCode="ISAZ" country="China" elevation="900" location="Wanding" pageId="329" pageNumber="330" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54AF88526A28FE8CAED8FBB9FDED3EA1" box="[454,563,979,1005]" pageId="329" pageNumber="330" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D12FD796A28FE8CAD23FBB9FD8C3EA1" box="[573,594,979,1005]" pageId="329" pageNumber="330" type="male">♂</specimenCount>
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“[
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<collectingCountry id="F30376606A28FE8CAD77FBB8FD6D3EA1" box="[617,691,978,1005]" name="China" pageId="329" pageNumber="330">China</collectingCountry>
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]
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<collectingRegion id="49D0F8126A28FE8CADDEFBB9FCC73EA1" box="[704,793,979,1005]" country="China" name="Yunnan" pageId="329" pageNumber="330">Yunnan</collectingRegion>
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,
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<location id="8ECB602B6A28FE8CAC3DFBB9FC543EA1" LSID="urn:lsid:plazi:treatment:03BD87E66A28FE8EAF89FB0CFD103FE5:8ECB602B6A28FE8CAC3DFBB9FC543EA1" box="[803,906,979,1005]" country="China" name="Wanding" pageId="329" pageNumber="330" stateProvince="Yunnan">Wanding</location>
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,
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<date id="FFAA10306A28FE8CAC88FBB9FBC73EA1" box="[918,1049,979,1005]" pageId="329" pageNumber="330" value="2011-04-29">
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<collectingDate id="EFEEE9D86A28FE8CAC88FBB9FBC73EA1" box="[918,1049,979,1005]" pageId="329" pageNumber="330" value="2011-04-29">2011-IV-29</collectingDate>
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</date>
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, N: 24.086, E: 98.072,
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<quantity id="4CEC9B156A28FE8CAA03FBB9FA823EA1" box="[1309,1372,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="329" pageNumber="330" unit="m" value="900.0">
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<elevation id="0039D1C36A28FE8CAA03FBB9FA823EA1" box="[1309,1372,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="329" pageNumber="330" unit="m" value="900.0">900m</elevation>
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</quantity>
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/ LW- 1249” (
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<collectionCode id="ED05AE356A28FE8CAFF5FB9DFEF1395D" box="[235,303,1015,1041]" pageId="329" pageNumber="330">ISAZ</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E657B6A28FE8EAFD9FC70FC9A3CB9" lastPageId="331" lastPageNumber="332" pageId="329" pageNumber="330" type="description">
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<paragraph id="8BAB36F06A28FE8CAFD9FC70FDE33931" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">
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<emphasis id="B960EAE26A28FE8CAFD9FC70FE873978" bold="true" box="[199,345,1050,1076]" pageId="329" pageNumber="330">Description.</emphasis>
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Length:
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<quantity id="4CEC9B156A28FE8CAEDFFC71FDC43979" box="[449,538,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="329" pageNumber="330" unit="mm" value="9.8">9.8 mm</quantity>
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, length of elytra:
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<quantity id="4CEC9B156A28FE8CADFBFC71FCE13979" box="[741,831,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="329" pageNumber="330" unit="mm" value="7.1">7.1 mm</quantity>
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, width:
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<quantity id="4CEC9B156A28FE8CAC82FC70FC283979" box="[924,1014,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="329" pageNumber="330" unit="mm" value="5.5">5.5 mm</quantity>
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. Body oval, dark brown, legs brown, antenna yellow, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, except long setae of head, and margins of pronotum and elytra glabrous.
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</paragraph>
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<paragraph id="8BAB36F06A28FE8CAFD9FCEDFDD938D1" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">
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Labroclypeus wide, trapezoidal, lateral margins slightly convex and convergent, producing with the ocular canthus a blunt angle, anterior angles moderately rounded, anterior margin weakly emarginate medially, margins weakly reflexed; surface flat, completely shiny, finely and densely punctate, glabrous; frontoclypeal suture finely incised, moderately curved; ocular canthus moderately wide and long (equals 1/3 of ocular width), finely and sparsely punctate, with a long terminal seta. Frons with fine, superficial, moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres, 1.1 times as long as remaining antennomeres combined. Eyes moderately large, ratio diameter/ interocular width: 0.58.
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<collectingRegion id="49D0F8126A28FE8CABCFFD35FAEE3835" box="[1233,1328,1375,1401]" country="Myanmar" name="Chin" pageId="329" pageNumber="330">Mentum</collectingRegion>
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convexly elevated and flattened anteriorly.
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</paragraph>
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<paragraph id="8BAB36F06A28FE8CAFD9FDCDFE143B1D" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">Pronotum widest at base, lateral margins evenly convex and convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt; anterior margin weakly convex, with a fine complete marginal line; basal margin without marginal line; surface finely and moderately densely punctate, setae of lateral and anterior margin long and dense, punctures with microscopic setae only. Hypomeron carinate. Scutellum wide, triangular, punctation as on pronotum.</paragraph>
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<paragraph id="8BAB36F06A28FE8CAFD9FE31FD233BF1" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">Elytra widest behind middle, striae finely impressed, finely punctate, intervals flat, finely and densely punctate; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes.</paragraph>
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<paragraph id="8BAB36F06A28FE8CAFD9FEADFEA83A3D" blockId="329.[151,1437,978,1977]" pageId="329" pageNumber="330">Ventral face coarsely and densely punctate, nearly glabrous, only metasternal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.84. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium weakly convex, coarsely and densely punctate, with moderately densely long setae on apical half.</paragraph>
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<paragraph id="8BAB36F06A28FE8EAFD9FF11FF673C9D" blockId="329.[151,1437,978,1977]" lastBlockId="331.[151,1437,151,681]" lastPageId="331" lastPageNumber="332" pageId="329" pageNumber="330">Legs short and wide; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur shiny, anterior margin acute, without adjacent serrated line, anterior row of setae reduced; posterior ventral margin, strongly widened in apical half, ventrally and dorsally not serrated, glabrous. Metatibia moderately long and wide, ratio width/ length: 1/ 2.33, sharply carinate dorsally, with two groups of spines, basal one shortly behind middle, apical one at five sixths of metatibial length, basally with a few robust and short setae; lateral face nearly flat, finely and sparsely superficially punctate on sides, along middle smooth; apex concavely emarginate interiorly near tarsal articulation. Tarsomeres impunctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined, and little shorter than dorsal tibial spur. Protibia moderately long, bidentate, with a blunt external extension at middle; anterior claws symmetric, interior tarsal claws sharply pointed.</paragraph>
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<caption id="DF6B66786A2BFE8FAF89FFFDFA4A3AB4" ID-DOI="http://doi.org/10.5281/zenodo.4496528" ID-Zenodo-Dep="4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="330" pageNumber="331" startId="330.[151,250,1943,1967]" targetBox="[172,1417,198,1910]" targetPageId="330">
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<paragraph id="8BAB36F06A2BFE8FAF89FFFDFA4A3AB4" blockId="330.[151,1437,1943,2040]" pageId="330" pageNumber="331">
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||
<emphasis id="B960EAE26A2BFE8FAF89FFFDFE813AFC" bold="true" box="[151,351,1943,1968]" pageId="330" pageNumber="331">FIGURE 79. A–D:</emphasis>
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<taxonomicName id="4C144D736A2BFE8FAE78FFF2FCE73AFC" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[358,825,1944,1968]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="330" pageNumber="331" phylum="Arthropoda" rank="species" species="wandingana" status="sp. nov.">
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<emphasis id="B960EAE26A2BFE8FAE78FFF2FD903AFC" box="[358,590,1944,1968]" italics="true" pageId="330" pageNumber="331">Maladera wandingana</emphasis>
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||
Ahrens, Fabrizi & Liu
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||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996A2BFE8FAC5DFFF2FCA43AE3" box="[835,890,1944,1967]" pageId="330" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype);
|
||
<emphasis id="B960EAE26A2BFE8FACE6FFF2FBEF3AFC" bold="true" box="[1016,1073,1944,1968]" pageId="330" pageNumber="331">E–H:</emphasis>
|
||
<taxonomicName id="4C144D736A2BFE8FAB27FFF2FF633A98" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="330" pageNumber="331" phylum="Arthropoda" rank="species" species="trifidiforceps" status="sp. nov.">
|
||
<emphasis id="B960EAE26A2BFE8FAB27FFF2FB003AFC" box="[1081,1246,1944,1968]" italics="true" pageId="330" pageNumber="331">M. trifidiforceps</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996A2BFE8FAFD8FFD6FF223A9F" box="[198,252,1980,2003]" pageId="330" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype);
|
||
<emphasis id="B960EAE26A2BFE8FAE66FFD6FE793A98" bold="true" box="[376,423,1980,2004]" pageId="330" pageNumber="331">I–L:</emphasis>
|
||
<taxonomicName id="4C144D736A2BFE8FAEB3FFD6FCE83A98" authority="Ahrens, Fabrizi & Liu" authorityName="Ahrens, Fabrizi & Liu" authorityYear="2021" box="[429,822,1980,2004]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="330" pageNumber="331" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
|
||
<emphasis id="B960EAE26A2BFE8FAEB3FFD6FD903A98" box="[429,590,1980,2004]" italics="true" pageId="330" pageNumber="331">M. nanlingensis</emphasis>
|
||
Ahrens, Fabrizi & Liu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25357996A2BFE8FAC21FFD6FCAB3A9F" box="[831,885,1980,2003]" pageId="330" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype).
|
||
<emphasis id="B960EAE26A2BFE8FACF1FFD1FBE23A98" bold="true" box="[1007,1084,1979,2004]" pageId="330" pageNumber="331">A, E, I:</emphasis>
|
||
aedeagus, left side lateral view;
|
||
<emphasis id="B960EAE26A2BFE8FAA9AFFD1FF0D3AB4" bold="true" pageId="330" pageNumber="331">C, G, K:</emphasis>
|
||
aedeagus, right side lateral view;
|
||
<emphasis id="B960EAE26A2BFE8FAD33FF8AFDA43AB4" bold="true" box="[557,634,2016,2040]" pageId="330" pageNumber="331">B, F, J:</emphasis>
|
||
parameres, dorsal view;
|
||
<emphasis id="B960EAE26A2BFE8FAC66FF8AFC0E3AB4" bold="true" box="[888,976,2016,2040]" pageId="330" pageNumber="331">D, H, L:</emphasis>
|
||
habitus. Scale: 0.5 mm. Habitus not to scale
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAB36F06A2AFE8EAFD9F9B1FC9A3CB9" blockId="331.[151,1437,151,681]" box="[199,836,475,501]" pageId="331" pageNumber="332">
|
||
Aedeagus:
|
||
<figureCitation id="132F2A756A2AFE8EAE5AF9B1FE4B3CB9" box="[324,405,475,501]" captionStart="FIGURE 79" captionStartId="330.[151,250,1943,1967]" captionTargetBox="[172,1417,198,1910]" captionTargetId="figure-19@330.[156,1431,181,1920]" captionTargetPageId="330" captionText="FIGURE 79. A–D: Maladera wandingana Ahrens, Fabrizi & Liu, sp. n. (holotype); E–H: M. trifidiforceps Ahrens, Fabrizi & Liu, sp. n. (holotype); I–L: M. nanlingensis Ahrens, Fabrizi & Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale" figureDoi="http://doi.org/10.5281/zenodo.4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="331" pageNumber="332">Fig. 79</figureCitation>
|
||
A–C. Habitus: 79D. Female unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6A2AFE8EAFD9F994FAEA3F71" pageId="331" pageNumber="332" type="diagnosis">
|
||
<paragraph id="8BAB36F06A2AFE8EAFD9F994FAEA3F71" blockId="331.[151,1437,151,681]" pageId="331" pageNumber="332">
|
||
<emphasis id="B960EAE26A2AFE8EAFD9F994FE9C3F54" bold="true" box="[199,322,510,536]" pageId="331" pageNumber="332">Diagnosis.</emphasis>
|
||
The species differs from all other Chinese
|
||
<taxonomicName id="4C144D736A2AFE8EAC3CF995FC4E3F55" authorityName="Mulsant & Rey" authorityYear="1871" box="[802,912,511,537]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="331" pageNumber="332" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B960EAE26A2AFE8EAC3CF995FC4E3F55" box="[802,912,511,537]" italics="true" pageId="331" pageNumber="332">Maladera</emphasis>
|
||
</taxonomicName>
|
||
species by the extended, but separated, ventral lobes fused with the ventral phallobase, and being well separated from the dorsally positioned parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6A2AFE8EAFD9FA2DFE043FC9" pageId="331" pageNumber="332" type="etymology">
|
||
<paragraph id="8BAB36F06A2AFE8EAFD9FA2DFE043FC9" blockId="331.[151,1437,151,681]" pageId="331" pageNumber="332">
|
||
<emphasis id="B960EAE26A2AFE8EAFD9FA2DFE903F2D" bold="true" box="[199,334,583,609]" pageId="331" pageNumber="332">Etymology.</emphasis>
|
||
The name (adjective in the nominative singular case) of the new species is derived from the name of its
|
||
<typeStatus id="54AF88526A2AFE8EAFC9FA01FED93FC9" box="[215,263,619,645]" pageId="331" pageNumber="332">type</typeStatus>
|
||
locality, Wanding.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30E657B6A2AFE8EAFD9FAE4FD103FE5" box="[199,718,654,681]" pageId="331" pageNumber="332" type="distribution">
|
||
<paragraph id="8BAB36F06A2AFE8EAFD9FAE4FD103FE5" blockId="331.[151,1437,151,681]" box="[199,718,654,681]" pageId="331" pageNumber="332">
|
||
<emphasis id="B960EAE26A2AFE8EAFD9FAE4FEBF3FE4" bold="true" box="[199,353,654,680]" pageId="331" pageNumber="332">Distribution.</emphasis>
|
||
See map (
|
||
<figureCitation id="132F2A756A2AFE8EAEC8FAE5FDE83FE5" box="[470,566,655,681]" captionStart="FIGURE 130" captionStartId="366.[151,250,1942,1966]" captionTargetBox="[232,1356,1126,1912]" captionTargetId="figure-64@366.[226,1362,1122,1918]" captionTargetPageId="366" captionText="FIGURE 130. Distribution of Maladera species with uncertain systematic status: Maladera panyuensis sp. n., M. parva (Moser, 1908), Maladera punctulata (Frey, 1972), M. rotunda (Arrow, 1946), M. tibialis (Brenske, 1898), M. trifidiforceps sp. n., M. wandingana sp. n." figureDoi="http://doi.org/10.5281/zenodo.4496670" httpUri="https://zenodo.org/record/4496670/files/figure.png" pageId="331" pageNumber="332">Fig. 130</figureCitation>
|
||
) and
|
||
<tableCitation id="C696034B6A2AFE8EAD6AFAE5FD163FE5" box="[628,712,655,681]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant & Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="331" pageNumber="332">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |