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<document id="BFAE950AD476FF50F46C2B938DF20875" ID-DOI="10.11646/zootaxa.5418.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10779797" ID-ZooBank="20F7368E-110A-4D71-9E28-ABBBA2EEC9C5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709587612430" checkinUser="plazi" docAuthor="Nobre, Welington B. &amp; Ruiz, Gustavo R. S." docDate="2024" docId="03BD87C12138DE66FF35A784F768FE85" docLanguage="en" docName="zootaxa.5418.5.3.pdf" docOrigin="Zootaxa 5418 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyphelia variegata Simon 1902" docType="treatment" docVersion="2" lastPageNumber="475" masterDocId="FF84FFB9213BDE62FFA2A126F340FFE0" masterDocTitle="On the boundary of the jumping spider genus Rhyphelia Simon, 1902 (Araneae: Salticidae: Euophryini), with description of ten new species" masterLastPageNumber="500" masterPageNumber="471" pageNumber="474" updateTime="1709588318203" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2CD0DA7ED1B196D7550359953996FC91">On the boundary of the jumping spider genus Rhyphelia Simon, 1902 (Araneae: Salticidae: Euophryini), with description of ten new species</mods:title>
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<mods:namePart id="C044207C6B8E7D12C6B65088EB8AD8EE">Nobre, Welington B.</mods:namePart>
<mods:affiliation id="A9F4AFC64F8338FD9B68280CE46EF4DF">Programa de Pós-Graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi, Brazil</mods:affiliation>
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<mods:namePart id="DFD6FFAC1A598CE25D00DAF3CBCECD4A">Ruiz, Gustavo R. S.</mods:namePart>
<mods:affiliation id="FA38993569629CDE5B8DB7D90EBCF55A">Instituto de Ciências Biológicas, Universidade Federal do Pará, Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil</mods:affiliation>
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<paragraph id="8BAB36D72138DE61FF35A784F16CF95C" blockId="3.[151,556,1698,1760]" box="[151,556,1698,1725]" pageId="3" pageNumber="474">
<heading id="D0E381BB2138DE61FF35A784F16CF95C" bold="true" box="[151,556,1698,1725]" fontSize="11" level="1" pageId="3" pageNumber="474" reason="1">
<taxonomicName id="4C144D542138DE61FF35A784F16CF95C" ID-CoL="6WVJ7" authority="Simon, 1902" authorityName="Simon" authorityYear="1902" box="[151,556,1698,1725]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B960EAC52138DE61FF35A784F16CF95C" bold="true" box="[151,556,1698,1725]" pageId="3" pageNumber="474">
<emphasis id="B960EAC52138DE61FF35A784F2CAF95C" bold="true" box="[151,394,1698,1725]" italics="true" pageId="3" pageNumber="474">Rhyphelia variegata</emphasis>
<bibRefCitation id="EF854B262138DE61FE30A784F16CF95C" author="Simon, E." box="[402,556,1698,1724]" pageId="3" pageNumber="474" pagination="24 - 56" refId="ref11632" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406." type="journal article" year="1902">Simon, 1902</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30E655C2138DE61FF35A7E0F261F900" box="[151,289,1734,1760]" pageId="3" pageNumber="474" type="description">
<paragraph id="8BAB36D72138DE61FF35A7E0F261F900" blockId="3.[151,556,1698,1760]" box="[151,289,1734,1760]" pageId="3" pageNumber="474">
<figureCitation id="132F2A522138DE61FF35A7E0F261F900" box="[151,289,1734,1760]" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="3" pageNumber="474">Figs 1A1D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E655C2138DE61FF35A62DF2A4F880" pageId="3" pageNumber="474" type="reference_group">
<paragraph id="8BAB36D72138DE61FF35A62DF2A4F880" blockId="3.[151,1436,1803,1888]" pageId="3" pageNumber="474">
<treatmentCitationGroup id="AB0411F92138DE61FF35A62DF2A4F880" pageId="3" pageNumber="474">
<treatmentCitation id="0AB510C62138DE61FF35A62DF15FF8C3" author="Simon, E." box="[151,543,1803,1827]" page="363" pageId="3" pageNumber="474" year="1902">
<taxonomicName id="4C144D542138DE61FF35A62DF15FF8C3" ID-CoL="6WVJ7" authority="Simon, 1902: 363" authorityName="Simon" authorityPageNumber="363" authorityYear="1902" box="[151,543,1803,1827]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B960EAC52138DE61FF35A62DF225F8C2" box="[151,357,1803,1827]" italics="true" pageId="3" pageNumber="474">Rhyphelia variegata</emphasis>
<bibRefCitation id="EF854B262138DE61FEC9A62DF15FF8C3" author="Simon, E." box="[363,543,1803,1827]" pageId="3" pageNumber="474" pagination="24 - 56" refId="ref11632" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406." type="journal article" year="1902">Simon, 1902: 363</bibRefCitation>
</taxonomicName>
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(Male
<typeStatus id="54AF88752138DE61FDC5A62DF180F8C3" box="[615,704,1803,1827]" pageId="3" pageNumber="474" type="holotype">holotype</typeStatus>
from Teresópolis,
<collectingRegion id="49D0F8352138DE61FCDCA62DF752F8C3" box="[894,1042,1803,1827]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="474">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F30376472138DE61FBB9A62DF71BF8C3" box="[1051,1115,1803,1827]" name="Brazil" pageId="3" pageNumber="474">Brazil</collectingCountry>
, deposited in Muséum National dHistoire Naturelle, Paris, examined);
<treatmentCitation id="0AB510C62138DE61FDFCA60FF05FF8A1" author="Simon, E." box="[606,799,1833,1857]" page="1053" pageId="3" pageNumber="474" year="1903">
<bibRefCitation id="EF854B262138DE61FDFCA60FF05FF8A1" author="Simon, E." box="[606,799,1833,1857]" pageId="3" pageNumber="474" pagination="669 - 1080" refId="ref11672" refString="Simon, E. (1903) n. k. In: Histoire naturelle des araignees. Tome Second. Deuxieme Edition. Roret, Paris, pp. 669 - 1080." type="book chapter" year="1903">Simon 1903: 1053</bibRefCitation>
</treatmentCitation>
, figs 11201121;
<treatmentCitation id="0AB510C62138DE61FC7AA60FF7DBF8A1" author="Galiano, M. E." box="[984,1179,1833,1857]" page="429" pageId="3" pageNumber="474" year="1963">
<bibRefCitation id="EF854B262138DE61FC7AA60FF7DBF8A1" author="Galiano, M. E." box="[984,1179,1833,1857]" pageId="3" pageNumber="474" pagination="273 - 470" refId="ref11018" refString="Galiano, M. E. (1963) Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales, Series C, 23, 273 - 470." type="journal article" year="1963">Galiano 1963: 429</bibRefCitation>
</treatmentCitation>
, pl. XXXIII, figs 1012;
<bibRefCitation id="EF854B262138DE61FF65A66EF2A0F880" author="World Spider Catalog" box="[199,480,1864,1888]" pageId="3" pageNumber="474" refId="ref11738" refString="World Spider Catalog (2023) World Spider Catalog. Version 24.5. Natural History Museum Bern, Bern. Avaliable from: http: // wsc. nmbe. ch (acessed 12 December 2023)" type="url" year="2023">World Spider Catalog 2023</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E655C2138DE66FF35A6A8F749FF35" lastPageId="4" lastPageNumber="475" pageId="3" pageNumber="474" type="diagnosis">
<paragraph id="8BAB36D72138DE66FF35A6A8F749FF35" blockId="3.[151,1437,1934,2033]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="475" pageId="3" pageNumber="474">
<emphasis id="B960EAC52138DE61FF35A6A8F252F848" bold="true" box="[151,274,1934,1960]" pageId="3" pageNumber="474">Diagnosis.</emphasis>
The male of
<taxonomicName id="4C144D542138DE61FE0AA6A9F172F848" authorityName="Simon" authorityYear="1902" box="[424,562,1935,1960]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B960EAC52138DE61FE0AA6A9F172F848" box="[424,562,1935,1960]" italics="true" pageId="3" pageNumber="474">R. variegata</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C144D542138DE61FCB5A6A9F0EEF848" authority="Nobre &amp; Ruiz, 2024" authorityName="Nobre &amp; Ruiz" authorityYear="2024" box="[791,942,1934,1960]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="brevistylus" status="sp. nov.">
<emphasis id="B960EAC52138DE61FCB5A6A9F0EEF848" box="[791,942,1934,1960]" italics="true" pageId="3" pageNumber="474">R. brevistylus</emphasis>
</taxonomicName>
<emphasis id="B960EAC52138DE61FC14A6A8F750F848" bold="true" box="[950,1040,1934,1960]" pageId="3" pageNumber="474">
<taxonomicNameLabel id="A25357BE2138DE61FC14A6A8F750F848" box="[950,1040,1934,1960]" pageId="3" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the longer carapace (about 1.5x longer than wide,
<figureCitation id="132F2A522138DE61FEC9A694F28AF82C" box="[363,458,1970,1996]" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="3" pageNumber="474">Figs 1A</figureCitation>
,
<figureCitation id="132F2A522138DE61FE7BA694F2BEF82C" box="[473,510,1970,1996]" captionStart="FIGURE 2" captionStartId="5.[151,250,1888,1912]" captionTargetBox="[155,1435,181,1861]" captionTargetId="figure-17@5.[151,1435,181,1863]" captionTargetPageId="5" captionText="FIGURE 2. Rhyphelia brevistylus sp. nov. AB male holotype (A dorsal view; B ventral view); CD female (C dorsal view; D ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10779803" httpUri="https://zenodo.org/record/10779803/files/figure.png" pageId="3" pageNumber="474">2A</figureCitation>
; shorter, 1.31.4x longer than wide in the remaining species), by the developed tegular lobe (
<figureCitation id="132F2A522138DE61FE89A6F0F2C6F810" box="[299,390,2006,2032]" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="3" pageNumber="474">Figs 1C</figureCitation>
,
<figureCitation id="132F2A522138DE61FE31A6F0F2F7F810" box="[403,439,2006,2032]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[196,1391,220,1918]" captionTargetId="figure-26@6.[196,1391,181,1918]" captionTargetPageId="6" captionText="FIGURE 3. Rhyphelia brevistylus sp. nov. AB male holotype, left palp (A ventral view; B retrolateral view); C epigyne, ventral view. D Rhyphelia bilineata sp. nov. epigyne, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10779805" httpUri="https://zenodo.org/record/10779805/files/figure.png" pageId="3" pageNumber="474">3A</figureCitation>
) and by the spurs on the border of the endites (
<figureCitation id="132F2A522138DE61FC65A6F0F75EF810" box="[967,1054,2006,2032]" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="3" pageNumber="474">Fig. 1B</figureCitation>
). The male of
<taxonomicName id="4C144D542138DE61FB60A6F1F60CF810" authorityName="Simon" authorityYear="1902" box="[1218,1356,2007,2032]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B960EAC52138DE61FB60A6F1F60CF810" box="[1218,1356,2007,2032]" italics="true" pageId="3" pageNumber="474">R. variegata</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C144D54213FDE66FE83A1B1F2F7FF51" authorityName="Nobre &amp; Ruiz" authorityYear="2024" box="[289,439,151,177]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="475" phylum="Arthropoda" rank="species" species="brevistylus">
<emphasis id="B960EAC5213FDE66FE83A1B1F2F7FF51" box="[289,439,151,177]" italics="true" pageId="4" pageNumber="475">R. brevistylus</emphasis>
</taxonomicName>
by the long embolus, which reaches almost the length of the tegulum (
<figureCitation id="132F2A52213FDE66FB61A1B1F657FF51" box="[1219,1303,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="4" pageNumber="475">Fig. 1C</figureCitation>
) (about one third the length of tegulum in
<taxonomicName id="4C144D54213FDE66FE45A19DF13EFF35" authorityName="Nobre &amp; Ruiz" authorityYear="2024" box="[487,638,187,213]" class="Arachnida" family="Salticidae" genus="Rhyphelia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="475" phylum="Arthropoda" rank="species" species="brevistylus">
<emphasis id="B960EAC5213FDE66FE45A19DF13EFF35" box="[487,638,187,213]" italics="true" pageId="4" pageNumber="475">R. brevistylus</emphasis>
</taxonomicName>
;
<figureCitation id="132F2A52213FDE66FD2FA19DF1A6FF35" box="[653,742,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[196,1391,220,1918]" captionTargetId="figure-26@6.[196,1391,181,1918]" captionTargetPageId="6" captionText="FIGURE 3. Rhyphelia brevistylus sp. nov. AB male holotype, left palp (A ventral view; B retrolateral view); C epigyne, ventral view. D Rhyphelia bilineata sp. nov. epigyne, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10779805" httpUri="https://zenodo.org/record/10779805/files/figure.png" pageId="4" pageNumber="475">Fig. 3A</figureCitation>
). The female is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E655C213FDE66FF65A1F8F3ACFEFD" pageId="4" pageNumber="475" type="description">
<paragraph id="8BAB36D7213FDE66FF65A1F8F3ACFEFD" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="475">
<emphasis id="B960EAC5213FDE66FF65A1F8F235FF18" bold="true" box="[199,373,222,248]" pageId="4" pageNumber="475">Redescription.</emphasis>
See
<bibRefCitation id="EF854B26213FDE66FE13A1F9F122FF19" author="Galiano, M. E." box="[433,610,223,249]" pageId="4" pageNumber="475" pagination="273 - 470" refId="ref11018" refString="Galiano, M. E. (1963) Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales, Series C, 23, 273 - 470." type="journal article" year="1963">Galiano (1963)</bibRefCitation>
. Herein we give new illustrations of the
<typeStatus id="54AF8875213FDE66FB9DA1F9F7E1FF19" box="[1087,1185,223,249]" pageId="4" pageNumber="475" type="holotype">holotype</typeStatus>
for comparison (
<figureCitation id="132F2A52213FDE66FAC9A1F9F3A0FEFD" captionStart="FIGURE 1" captionStartId="4.[151,250,1507,1531]" captionTargetBox="[151,1435,381,1484]" captionTargetId="figure-109@4.[151,1435,381,1484]" captionTargetPageId="4" captionText="FIGURE 1. Rhyphelia variegata Simon, 1902, male holotype. A dorsal view; B chelicera and endites, ventral view; CD right male palp, digitally inverted (C ventral view; D retrolateral view)." figureDoi="http://doi.org/10.5281/zenodo.10779799" httpUri="https://zenodo.org/record/10779799/files/figure.png" pageId="4" pageNumber="475">Figs 1AD</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E655C213FDE66FF65A000F768FE85" pageId="4" pageNumber="475" type="distribution">
<paragraph id="8BAB36D7213FDE66FF65A000F768FE85" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="475">
<emphasis id="B960EAC5213FDE66FF65A000F221FEA0" bold="true" box="[199,353,294,320]" pageId="4" pageNumber="475">Distribution.</emphasis>
The original description mentions Caracas (
<collectingCountry id="F3037647213FDE66FCEBA001F080FEA1" box="[841,960,295,321]" name="Venezuela" pageId="4" pageNumber="475">Venezuela</collectingCountry>
) as the
<typeStatus id="54AF8875213FDE66FBB1A001F703FEA1" box="[1043,1091,295,321]" pageId="4" pageNumber="475">type</typeStatus>
locality, but the label states the actual
<typeStatus id="54AF8875213FDE66FF43A06DF251FE85" box="[225,273,331,357]" pageId="4" pageNumber="475">type</typeStatus>
locality in the state of
<collectingRegion id="49D0F835213FDE66FDB0A06DF1F7FE85" box="[530,695,331,357]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="475">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F3037647213FDE66FD60A06DF046FE85" box="[706,774,331,357]" name="Brazil" pageId="4" pageNumber="475">Brazil</collectingCountry>
(
<bibRefCitation id="EF854B26213FDE66FCB7A06DF0EFFE85" author="Galiano, M. E." box="[789,943,331,357]" pageId="4" pageNumber="475" pagination="273 - 470" refId="ref11018" refString="Galiano, M. E. (1963) Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales, Series C, 23, 273 - 470." type="journal article" year="1963">Galiano 1963</bibRefCitation>
) (
<figureCitation id="132F2A52213FDE66FC65A06DF75BFE85" box="[967,1051,331,357]" captionStart="FIGURE 22" captionStartId="27.[151,250,1040,1064]" captionTargetBox="[243,1344,181,1016]" captionTargetId="figure-17@27.[243,1344,181,1016]" captionTargetPageId="27" captionText="FIGURE 22. Map with individual symbols for the species of Rhyphelia described and recorded prior to this work (new species described herein are mapped as red triangles and separate in the next map). Brazilian states are in italics." figureDoi="http://doi.org/10.5281/zenodo.10779865" httpUri="https://zenodo.org/record/10779865/files/figure.png" pageId="4" pageNumber="475">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>