treatments-xml/data/03/BD/87/03BD87BAFFD3E063FC41F9C6FC16FC05.xml

<document id="B5871CBD1FAA9CC07733A80D9CEEE908" ID-DOI="10.5281/zenodo.4504476" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504476" ID-ZooBank="2CB78941-EAAD-461E-BFFC-D4262A16AE3A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612466473556" checkinUser="carolina" docAuthor="Do, Van Tu, Nguyen, Tong Cuong &amp; Dang, Van Dong" docDate="2017" docId="03BD87BAFFD3E063FC41F9C6FC16FC05" docLanguage="en" docName="RafflesBZool.65.455-465.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Tiwaripotamon hamyen Do &amp; Nguyen &amp; Dang 2017, new species" docType="treatment" docUuid="4380D18D-4907-4288-BE57-2C57E343F917" docUuidSource="ZooBank" docVersion="5" lastPageNumber="464" masterDocId="FF84FFC2FFD5E06AFFF5FFAFFFFEFFD0" masterDocTitle="Two new species of freshwater crabs of the genus Tiwaripotamon Bott, 1970 (Crustacea, Decapoda, Brachyura, Potamidae) from northern Vietnam" masterLastPageNumber="465" masterPageNumber="455" pageNumber="461" updateTime="1698866241085" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2C5BCCDA478A37B05BE5CD49D6560CEE">Two new species of freshwater crabs of the genus Tiwaripotamon Bott, 1970 (Crustacea, Decapoda, Brachyura, Potamidae) from northern Vietnam</mods:title>
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<mods:namePart id="A70C8CD917E949AC9B8224792573369B">Do, Van Tu</mods:namePart>
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<mods:namePart id="E7F4680249EA27E268482E9B76065BC0">Nguyen, Tong Cuong</mods:namePart>
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<mods:namePart id="7DF667F85E6BBF15F53FFC20FC456155">Dang, Van Dong</mods:namePart>
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<paragraph id="8BAB36ACFFD3E06CFC41F9C6FAACF951" blockId="6.[948,1362,1641,1697]" box="[948,1362,1641,1665]" pageId="6" pageNumber="461">
<heading id="D0E381C0FFD3E06CFC41F9C6FAACF951" bold="true" box="[948,1362,1641,1665]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="461" reason="2">
<emphasis id="B960EABEFFD3E06CFC41F9C6FAACF951" bold="true" box="[948,1362,1641,1665]" pageId="6" pageNumber="461">
<taxonomicName id="4C144D2FFFD3E06CFC41F9C6FB43F951" ID-CoL="7CDNT" authority="Do &amp; Nguyen &amp; Dang, 2017" authorityName="Do &amp; Nguyen &amp; Dang" authorityYear="2017" box="[948,1213,1641,1665]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="461" phylum="Arthropoda" rank="species" species="hamyen" status="sp. nov.">
<emphasis id="B960EABEFFD3E06CFC41F9C6FB43F951" bold="true" box="[948,1213,1641,1665]" italics="true" pageId="6" pageNumber="461">Tiwaripotamon hamyen</emphasis>
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, 
<taxonomicNameLabel id="A25357C5FFD3E06CFB39F9C6FAACF951" box="[1228,1362,1641,1665]" pageId="6" pageNumber="461" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BAB36ACFFD3E06CFBB2F926FB40F971" blockId="6.[948,1362,1641,1697]" box="[1095,1214,1673,1697]" pageId="6" pageNumber="461">
(
<figureCitation id="132F2A29FFD3E06CFBBAF926FB48F971" box="[1103,1206,1673,1697]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[136,171,1279,1301]" captionStartId-1="7.[113,148,1958,1980]" captionStartId-2="8.[135,170,1096,1118]" captionTargetBox-0="[135,1473,189,1264]" captionTargetBox-1="[114,1452,294,1941]" captionTargetBox-2="[136,1474,189,1081]" captionTargetId-0="figure-318@6.[135,1473,189,1264]" captionTargetId-1="figure-14@7.[107,1465,287,1955]" captionTargetId-2="figure-465@8.[136,1474,189,1081]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." captionText-1="Fig. 7. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." captionText-2="Fig. 8. A–D, Tiwaripotamon hamyen, new species, with differences living colour in carapace and ambulatory legs." figureDoi-0="http://doi.org/10.5281/zenodo.4504488" figureDoi-1="http://doi.org/10.5281/zenodo.4504490" figureDoi-2="http://doi.org/10.5281/zenodo.4504492" httpUri-0="https://zenodo.org/record/4504488/files/figure.png" httpUri-1="https://zenodo.org/record/4504490/files/figure.png" httpUri-2="https://zenodo.org/record/4504492/files/figure.png" pageId="6" pageNumber="461">Figs. 6–8</figureCitation>
)
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<paragraph id="8BAB36ACFFD3E06CFCB6F966FB4CF891" blockId="6.[835,1476,1737,1857]" pageId="6" pageNumber="461">
<emphasis id="B960EABEFFD3E06CFCB6F966FBDAF931" bold="true" box="[835,1060,1737,1761]" pageId="6" pageNumber="461">Material examined.</emphasis>
<materialsCitation id="3B7C3CF1FFD3E06CFBDBF966FB4CF891" collectingDate="2015-08-27" collectionCode="FC" collectorName="Chu H. Q." country="Vietnam" elevation="232" latitude="22.055683" location="Binh Xa" longLatPrecision="1" longitude="105.16064" municipality="Ham Yen district" pageId="6" pageNumber="461" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="holotype">
<typeStatus id="54AF880EFFD3E06CFBDBF966FB6BF931" box="[1070,1173,1737,1761]" pageId="6" pageNumber="461" type="holotype">Holotype</typeStatus>
: male (41.7 × 32.3) (IEBR- 
<collectionCode id="ED05AE69FFD3E06CFCB6F946FC9AF8D1" box="[835,868,1769,1793]" pageId="6" pageNumber="461">FC</collectionCode>
THx01), 
<geoCoordinate id="EE20506BFFD3E06CFC3FF947FBACF8D1" box="[970,1106,1768,1793]" degrees="22" direction="north" minutes="03.341" orientation="latitude" pageId="6" pageNumber="461" precision="1" value="22.055683">22°03.341′N</geoCoordinate>
<geoCoordinate id="EE20506BFFD3E06CFBA2F947FB15F8D1" box="[1111,1259,1768,1793]" degrees="105" direction="east" minutes="09.638" orientation="longitude" pageId="6" pageNumber="461" precision="1" value="105.16064">105°09.638′E</geoCoordinate>
, 
<location id="8ECB6077FFD3E06CFB00F946FAB3F8D1" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFD3E06CFB00F946FAB3F8D1" box="[1269,1357,1769,1793]" country="Vietnam" latitude="22.055683" longLatPrecision="1" longitude="105.16064" municipality="Ham Yen district" name="Binh Xa" pageId="6" pageNumber="461" stateProvince="Tuyen Quang">Binh Xa</location>
commune, 
<collectingMunicipality id="6BCFACD6FFD3E06CFCB6F8A5FC01F8F1" box="[835,1023,1801,1825]" pageId="6" pageNumber="461">Ham Yen district</collectingMunicipality>
, 
<collectingRegion id="49D0F84EFFD3E06CFBFFF8A5FAFDF8F1" box="[1034,1283,1801,1826]" country="Vietnam" name="Tuyen Quang" pageId="6" pageNumber="461">Tuyen Quang province</collectingRegion>
, 
<collectingCountry id="F303763CFFD3E06CFAFBF8A6FA93F8F1" box="[1294,1389,1801,1825]" name="Vietnam" pageId="6" pageNumber="461">Vietnam</collectingCountry>
, 
<quantity id="4CEC9B49FFD3E06CFA8DF8A6FA40F8F1" box="[1400,1470,1801,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="6" pageNumber="461" unit="m" value="232.0">
<elevation id="0039D19FFFD3E06CFA8DF8A6FA40F8F1" box="[1400,1470,1801,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="6" pageNumber="461" unit="m" value="232.0">232 m</elevation>
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, coll. 
<collectorName id="26E1537AFFD3E06CFC8EF886FC10F891" box="[891,1006,1833,1857]" pageId="6" pageNumber="461">Chu H. Q.</collectorName>
, 
<date id="FFAA106CFFD3E06CFC09F886FB50F891" box="[1020,1198,1833,1857]" pageId="6" pageNumber="461" value="2015-08-27">
<collectingDate id="EFEEE984FFD3E06CFC09F886FB50F891" box="[1020,1198,1833,1857]" pageId="6" pageNumber="461" value="2015-08-27">27 August 2015</collectingDate>
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.
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<paragraph id="8BAB36ACFFD3E06CFCB6F8C6FBC4F831" blockId="6.[834,1476,1897,2017]" pageId="6" pageNumber="461">
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<typeStatus id="54AF880EFFD3E06CFCB6F8C6FC49F851" box="[835,951,1897,1921]" pageId="6" pageNumber="461" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D12FD25FFD3E06CFC31F8C6FBE9F851" box="[964,1047,1897,1921]" pageId="6" pageNumber="461" type="male">5 males</specimenCount>
(37.2 × 28.4, 36.4 × 28.3, 35.7 × 27.8, 35.0 × 27.3, 29.4 × 23.2) (
<collectionCode id="ED05AE69FFD3E06CFB94F826FB37F871" box="[1121,1225,1929,1953]" pageId="6" pageNumber="461">IEBR-FC</collectionCode>
THx02–06), 
<specimenCount id="9D12FD25FFD3E06CFAA9F826FA3DF871" box="[1372,1475,1929,1953]" pageId="6" pageNumber="461" type="female">3 females</specimenCount>
(43.8 × 33.0, 40.6 × 29.8, 40.0 × 30.8) (
<collectionCode id="ED05AE69FFD3E06CFB22F806FAC0F811" box="[1239,1342,1961,1985]" pageId="6" pageNumber="461">IEBR-FC</collectionCode>
THx07–09), same data as holotype.
</paragraph>
<caption id="DF6B6624FFD2E06DFF84F809FAC1F808" ID-DOI="http://doi.org/10.5281/zenodo.4504490" ID-Zenodo-Dep="4504490" httpUri="https://zenodo.org/record/4504490/files/figure.png" pageId="7" pageNumber="462" startId="7.[113,148,1958,1980]" targetBox="[114,1452,294,1941]" targetPageId="7">
<paragraph id="8BAB36ACFFD2E06DFF84F809FAC1F808" blockId="7.[113,1452,1958,2009]" pageId="7" pageNumber="462">
Fig. 7. 
<taxonomicName id="4C144D2FFFD2E06DFF42F808FE62F86B" authority="Do &amp; Nguyen &amp; Dang, 2017" authorityName="Do &amp; Nguyen &amp; Dang" authorityYear="2017" box="[183,412,1958,1980]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="462" phylum="Arthropoda" rank="species" species="hamyen" status="sp. nov.">
<emphasis id="B960EABEFFD2E06DFF42F808FE62F86B" box="[183,412,1958,1980]" italics="true" pageId="7" pageNumber="462">Tiwaripotamon hamyen</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25357C5FFD2E06DFE5DF809FDE1F86C" box="[424,543,1958,1980]" pageId="7" pageNumber="462" rank="species">new species</taxonomicNameLabel>
, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment.
</paragraph>
</caption>
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<paragraph id="8BAB36ACFFDDE062FF72FBE7FB0EFB8E" blockId="8.[135,1264,1096,1118]" box="[135,1264,1096,1118]" pageId="8" pageNumber="463">
Fig. 8. A–D, 
<taxonomicName id="4C144D2FFFDDE062FEF8FBE6FE0CFB8D" authority="Do &amp; Nguyen &amp; Dang, 2017" authorityName="Do &amp; Nguyen &amp; Dang" authorityYear="2017" box="[269,498,1096,1118]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="463" phylum="Arthropoda" rank="species" species="hamyen" status="sp. nov.">
<emphasis id="B960EABEFFDDE062FEF8FBE6FE0CFB8D" box="[269,498,1096,1118]" italics="true" pageId="8" pageNumber="463">Tiwaripotamon hamyen</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25357C5FFDDE062FDF5FBE6FD89FB8E" box="[512,631,1096,1118]" pageId="8" pageNumber="463" rank="species">new species</taxonomicNameLabel>
, with differences living colour in carapace and ambulatory legs.
</paragraph>
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<paragraph id="8BAB36ACFFDDE062FF7DFB3AFDDBF9BD" blockId="8.[135,776,1173,1645]" pageId="8" pageNumber="463">
<emphasis id="B960EABEFFDDE062FF7DFB3AFE66FB7D" bold="true" box="[136,408,1173,1197]" pageId="8" pageNumber="463">Comparative material.</emphasis>
<taxonomicName id="4C144D2FFFDDE062FE56FB39FEEAFB1D" authority="Ng &amp; Yeo, 2001: 1" authorityName="Ng &amp; Yeo" authorityPageNumber="1" authorityYear="2001" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="463" phylum="Arthropoda" rank="species" species="edostilus">
<emphasis id="B960EABEFFDDE062FE56FB39FD45FB7D" box="[419,699,1173,1197]" italics="true" pageId="8" pageNumber="463">Tiwaripotamon edostilus</emphasis>
<materialsCitation id="3B7C3CF1FFDDE062FD33FB39FEEAFB1D" collectingDate="2013-03-18" collectingDateMax="2013-03-19" collectingDateMin="2013-03-18" collectionCode="IEBR-FC, V, T" country="Vietnam" elevation="93" location="Cat Ba island" municipality="Do" pageId="8" pageNumber="463" specimenCode="TE01" specimenCount="1" specimenCount-male="1" stateProvince="Hai Phong">
<bibRefCitation id="EF854B5DFFDDE062FD33FB39FEEAFB1D" author="Ng PKL &amp; Yeo DCJ" pageId="8" pageNumber="463" pagination="275 - 287" refId="ref5474" refString="Ng PKL &amp; Yeo DCJ (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21: 275 - 287." type="journal article" year="2001">
Ng &amp; Yeo, 2001: 
<specimenCount id="9D12FD25FFDDE062FEF2FB1AFEEAFB1D" box="[263,276,1205,1229]" pageId="8" pageNumber="463" type="male">1</specimenCount>
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male (26.1 × 21.4), (
<collectionCode id="ED05AE69FFDDE062FE0FFB1AFD9CFB1D" box="[506,610,1205,1229]" pageId="8" pageNumber="463">IEBR-FC</collectionCode>
<specimenCode id="DBB29ED7FFDDE062FD9FFB1AFD57FB1D" box="[618,681,1205,1229]" collectionCode="TE" pageId="8" pageNumber="463">TE01</specimenCode>
), 
<location id="8ECB6077FFDDE062FD48FB1AFF35FB3D" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFDDE062FD48FB1AFF35FB3D" country="Vietnam" municipality="Do" name="Cat Ba island" pageId="8" pageNumber="463" stateProvince="Hai Phong">Cat Ba island</location>
, 
<collectingRegion id="49D0F84EFFDDE062FF2DFB7AFEB4FB3D" box="[216,330,1237,1261]" country="Vietnam" name="Hai Phong" pageId="8" pageNumber="463">Hai Phong</collectingRegion>
city, 
<collectingCountry id="F303763CFFDDE062FE7FFB7AFE17FB3D" box="[394,489,1237,1261]" name="Vietnam" pageId="8" pageNumber="463">Vietnam</collectingCountry>
, 
<quantity id="4CEC9B49FFDDE062FE00FB7AFD9AFB3D" box="[501,612,1237,1261]" metricMagnitude="1" metricUnit="m" metricValue="9.25" metricValueMax="17.0" metricValueMin="1.5" pageId="8" pageNumber="463" unit="m" value="92.5" valueMax="170.0" valueMin="15.0">
<elevation id="0039D19FFFDDE062FE00FB7AFD9AFB3D" box="[501,612,1237,1261]" metricMagnitude="1" metricUnit="m" metricValue="9.25" metricValueMax="17.0" metricValueMin="1.5" pageId="8" pageNumber="463" unit="m" value="92.5" valueMax="170.0" valueMin="15.0">15–170 m</elevation>
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, coll. 
<collectionCode id="ED05AE69FFDDE062FD52FB79FD40FB3D" box="[679,702,1238,1261]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="463" type="Museum">V</collectionCode>
. 
<collectionCode id="ED05AE69FFDDE062FD37FB79FD2BFB3D" box="[706,725,1238,1261]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="463">T</collectionCode>
. 
<collectingMunicipality id="6BCFACD6FFDDE062FD14FB79FCFAFB3D" box="[737,772,1238,1261]" pageId="8" pageNumber="463">Do</collectingMunicipality>
, 
<date id="FFAA106CFFDDE062FF7FFB5AFEA3FADD" box="[138,349,1269,1293]" pageId="8" pageNumber="463" value="2013-03-18" valueMax="2013-03-19" valueMin="2013-03-18">
<collectingDate id="EFEEE984FFDDE062FF7FFB5AFEA3FADD" box="[138,349,1269,1293]" pageId="8" pageNumber="463" value="2013-03-18" valueMax="2013-03-19" valueMin="2013-03-18">18–19 March 2013</collectingDate>
</date>
. 
<materialsCitation id="3B7C3CF1FFDDE062FE92FB59FF3EFA5D" collectingDate="2013-05-14" collectionCode="IEBR-FC, V, T" country="Vietnam" elevation="500" location="Cuc Phuong National Park" municipality="Do" pageId="8" pageNumber="463" specimenCount="2" specimenCount-male="2" stateProvince="Ninh Binh">
<taxonomicName id="4C144D2FFFDDE062FE92FB59FF05FAFD" authority="(Dang &amp; Ho, 2002)" baseAuthorityName="Dang &amp; Ho" baseAuthorityYear="2002" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="463" phylum="Arthropoda" rank="species" species="vietnamicum">
<emphasis id="B960EABEFFDDE062FE92FB59FD63FADD" box="[359,669,1270,1293]" italics="true" pageId="8" pageNumber="463">Tiwaripotamon vietnamicum</emphasis>
(
<bibRefCitation id="EF854B5DFFDDE062FD5BFB5AFF0AFAFD" author="Dang NT &amp; Ho TH" pageId="8" pageNumber="463" pagination="1 - 8" refId="ref5285" refString="Dang NT &amp; Ho TH (2002) Two new crab species of Potamidae from Vietnam. Journal of Biology, 24: 1 - 8. [In Vietnamese, with summary in English]." type="journal article" year="2002">Dang &amp; Ho, 2002</bibRefCitation>
)
</taxonomicName>
: 
<specimenCount id="9D12FD25FFDDE062FEFDFABAFEA3FAFD" box="[264,349,1301,1325]" pageId="8" pageNumber="463" type="male">2 males</specimenCount>
(44.5 × 32.8, 41.3 × 31.2), (
<collectionCode id="ED05AE69FFDDE062FD6AFABAFCF9FAFD" box="[671,775,1301,1325]" pageId="8" pageNumber="463">IEBR-FC</collectionCode>
TVn01, NCHUZOOL 13612), 
<location id="8ECB6077FFDDE062FE2EFA9AFCFDFA9D" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFDDE062FE2EFA9AFCFDFA9D" box="[475,771,1333,1357]" country="Vietnam" municipality="Do" name="Cuc Phuong National Park" pageId="8" pageNumber="463" stateProvince="Ninh Binh">Cuc Phuong National Park</location>
, 
<collectingRegion id="49D0F84EFFDDE062FF7DFAFAFEA3FABD" box="[136,349,1365,1389]" country="Vietnam" name="Ninh Binh" pageId="8" pageNumber="463">Ninh Binh province</collectingRegion>
, 
<collectingCountry id="F303763CFFDDE062FE93FAFAFE3BFABD" box="[358,453,1365,1389]" name="Vietnam" pageId="8" pageNumber="463">Vietnam</collectingCountry>
, 
<quantity id="4CEC9B49FFDDE062FE3AFAFAFDEDFABD" box="[463,531,1365,1389]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="463" unit="m" value="500.0">
<elevation id="0039D19FFFDDE062FE3AFAFAFDEDFABD" box="[463,531,1365,1389]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="463" unit="m" value="500.0">500 m</elevation>
</quantity>
, coll. 
<collectionCode id="ED05AE69FFDDE062FDA7FAF9FD97FABD" box="[594,617,1366,1389]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="463" type="Museum">V</collectionCode>
. 
<collectionCode id="ED05AE69FFDDE062FD99FAF9FD81FABD" box="[620,639,1366,1389]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="463">T</collectionCode>
. 
<collectingMunicipality id="6BCFACD6FFDDE062FD7DFAF9FD55FABD" box="[648,683,1366,1389]" pageId="8" pageNumber="463">Do</collectingMunicipality>
, 
<date id="FFAA106CFFDDE062FD40FAFAFF3EFA5D" pageId="8" pageNumber="463" value="2013-05-14">
<collectingDate id="EFEEE984FFDDE062FD40FAFAFF3EFA5D" pageId="8" pageNumber="463" value="2013-05-14">14 May 2013</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B7C3CF1FFDDE062FF3FFAD9FDADFA3D" collectingDate="2013-07-02" collectionCode="IEBR-FC" collectorName="N. L. Doan &amp; X. N. Nguyen" country="Vietnam" elevation="758" location="Tung Ba" municipality="Vi Xuyen district" pageId="8" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Ha Giang">
<taxonomicName id="4C144D2FFFDDE062FF3FFAD9FD32FA5D" authority="Shih &amp; Do, 2014: 1" authorityName="Shih &amp; Do" authorityPageNumber="1" authorityYear="2014" box="[202,716,1397,1421]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="463" phylum="Arthropoda" rank="species" species="vixuyenense">
<emphasis id="B960EABEFFDDE062FF3FFAD9FE08FA5D" box="[202,502,1398,1421]" italics="true" pageId="8" pageNumber="463">Tiwaripotamon vixuyenense</emphasis>
<bibRefCitation id="EF854B5DFFDDE062FE09FADAFD32FA5D" author="Shih HT &amp; Do VT" box="[508,716,1397,1421]" pageId="8" pageNumber="463" pagination="26 - 38" refId="ref5617" refString="Shih HT &amp; Do VT (2014) A new species of Tiwaripotamon Bott, 1970, from northern Vietnam, with notes on T. vietnamicum (Dang &amp; Ho, 2002) and T. edostilus Ng &amp; Yeo, 2001 (Crustacea, Brachyura, Potamidae). Zootaxa, 3764: 26 - 38." type="journal article" year="2014">
Shih &amp; Do, 2014: 
<specimenCount id="9D12FD25FFDDE062FD4AFADAFD32FA5D" box="[703,716,1397,1421]" pageId="8" pageNumber="463" type="male">1</specimenCount>
</bibRefCitation>
</taxonomicName>
male (26.4 × 20.5) (
<collectionCode id="ED05AE69FFDDE062FED8FA3AFE68FA7D" box="[301,406,1429,1453]" pageId="8" pageNumber="463">IEBR-FC</collectionCode>
TVx01), 
<location id="8ECB6077FFDDE062FDF3FA39FD99FA7D" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFDDE062FDF3FA39FD99FA7D" box="[518,615,1430,1454]" country="Vietnam" municipality="Vi Xuyen district" name="Tung Ba" pageId="8" pageNumber="463" stateProvince="Ha Giang">Tung Ba</location>
commune, 
<collectingMunicipality id="6BCFACD6FFDDE062FD19FA3AFED8FA1D" pageId="8" pageNumber="463">Vi Xuyen district</collectingMunicipality>
, 
<collectingRegion id="49D0F84EFFDDE062FEC6FA19FDF9FA1D" box="[307,519,1461,1485]" country="Vietnam" name="Ha Giang" pageId="8" pageNumber="463">Ha Giang province</collectingRegion>
, 
<collectingCountry id="F303763CFFDDE062FDE0FA1AFD8AFA1D" box="[533,628,1461,1485]" name="Vietnam" pageId="8" pageNumber="463">Vietnam</collectingCountry>
, 
<quantity id="4CEC9B49FFDDE062FD74FA1AFD34FA1D" box="[641,714,1461,1485]" metricMagnitude="2" metricUnit="m" metricValue="7.58" pageId="8" pageNumber="463" unit="m" value="758.0">
<elevation id="0039D19FFFDDE062FD74FA1AFD34FA1D" box="[641,714,1461,1485]" metricMagnitude="2" metricUnit="m" metricValue="7.58" pageId="8" pageNumber="463" unit="m" value="758.0">758 m</elevation>
</quantity>
, coll. 
<collectorName id="26E1537AFFDDE062FF7DFA79FF00FA3D" box="[136,254,1494,1517]" pageId="8" pageNumber="463">N.L. Doan</collectorName>
&amp; 
<collectorName id="26E1537AFFDDE062FED2FA79FE43FA3E" box="[295,445,1494,1518]" pageId="8" pageNumber="463">X.N. Nguyen</collectorName>
, 
<date id="FFAA106CFFDDE062FE39FA7AFDADFA3D" box="[460,595,1493,1517]" pageId="8" pageNumber="463" value="2013-07-02">
<collectingDate id="EFEEE984FFDDE062FE39FA7AFDADFA3D" box="[460,595,1493,1517]" pageId="8" pageNumber="463" value="2013-07-02">2 July 2013</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B7C3CF1FFDDE062FD94FA79FDDFF9BD" collectingDate="2014-06-07" collectionCode="IEBR-FC, T" country="Vietnam" county="Pham" elevation="572" location="Coong village" longitude="106.66417" municipality="Ha Lang district" pageId="8" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
<taxonomicName id="4C144D2FFFDDE062FD94FA79FEFEF9DD" authorityName="Do et al" authorityYear="2016" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="463" phylum="Arthropoda" rank="species" species="pluviosum">
<emphasis id="B960EABEFFDDE062FD94FA79FEFEF9DD" italics="true" pageId="8" pageNumber="463">Tiwaripotamon pluviosum</emphasis>
</taxonomicName>
: 
<specimenCount id="9D12FD25FFDDE062FEE3FA5AFE96F9DD" box="[278,360,1525,1549]" pageId="8" pageNumber="463" type="male">1 male</specimenCount>
(32.2 × 23.2) (
<collectionCode id="ED05AE69FFDDE062FDDEFA5AFD62F9DD" box="[555,668,1525,1549]" pageId="8" pageNumber="463">IEBR-FC</collectionCode>
TPx01), 22°43′466″N 
<geoCoordinate id="EE20506BFFDDE062FED1F9BBFE3DF9FD" box="[292,451,1556,1581]" degrees="106" direction="east" minutes="39" orientation="longitude" pageId="8" pageNumber="463" precision="15" seconds="051" value="106.66417">106°39′051″E</geoCoordinate>
, 
<location id="8ECB6077FFDDE062FE26F9BAFD88F9FD" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFDDE062FE26F9BAFD88F9FD" box="[467,630,1557,1581]" country="Vietnam" county="Pham" longitude="106.66417" municipality="Ha Lang district" name="Coong village" pageId="8" pageNumber="463" stateProvince="Cao Bang">Coong village</location>
, 
<location id="8ECB6077FFDDE062FD70F9B9FCF9F9FD" LSID="urn:lsid:plazi:treatment:03BD87BAFFD3E063FC41F9C6FC16FC05:8ECB6077FFDDE062FD70F9B9FCF9F9FD" box="[645,775,1557,1581]" country="Vietnam" county="Pham" longitude="106.66417" municipality="Ha Lang district" name="Duc Quang" pageId="8" pageNumber="463" stateProvince="Cao Bang">Duc Quang</location>
commune, 
<collectingMunicipality id="6BCFACD6FFDDE062FEF4F999FE48F99D" box="[257,438,1589,1614]" pageId="8" pageNumber="463">Ha Lang district</collectingMunicipality>
, 
<collectingRegion id="49D0F84EFFDDE062FE36F99AFD66F99D" box="[451,664,1589,1614]" country="Vietnam" name="Cao Bang" pageId="8" pageNumber="463">Cao Bang province</collectingRegion>
, 
<collectingCountry id="F303763CFFDDE062FD50F99AFCFAF99D" box="[677,772,1589,1613]" name="Vietnam" pageId="8" pageNumber="463">Vietnam</collectingCountry>
, 
<quantity id="4CEC9B49FFDDE062FF7DF9FAFF2EF9BD" box="[136,208,1621,1645]" metricMagnitude="2" metricUnit="m" metricValue="5.72" pageId="8" pageNumber="463" unit="m" value="572.0">
<elevation id="0039D19FFFDDE062FF7DF9FAFF2EF9BD" box="[136,208,1621,1645]" metricMagnitude="2" metricUnit="m" metricValue="5.72" pageId="8" pageNumber="463" unit="m" value="572.0">572 m</elevation>
</quantity>
, coll. 
<collectionCode id="ED05AE69FFDDE062FEE1F9FAFED9F9BD" box="[276,295,1621,1645]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="463">T</collectionCode>
.C. 
<collectingCounty id="62CA4E20FFDDE062FEB9F9FAFE72F9BD" box="[332,396,1621,1645]" pageId="8" pageNumber="463">Pham</collectingCounty>
, 
<date id="FFAA106CFFDDE062FE6DF9F9FDDFF9BD" box="[408,545,1621,1645]" pageId="8" pageNumber="463" value="2014-06-07">
<collectingDate id="EFEEE984FFDDE062FE6DF9F9FDDFF9BD" box="[408,545,1621,1645]" pageId="8" pageNumber="463" value="2014-06-07">7 June 2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6527FFDDE062FF7DF93AFBBCFB1D" pageId="8" pageNumber="463" type="diagnosis">
<paragraph id="8BAB36ACFFDDE062FF7DF93AFBBCFB1D" blockId="8.[135,777,1685,2094]" lastBlockId="8.[835,1474,1174,1230]" pageId="8" pageNumber="463">
<emphasis id="B960EABEFFDDE062FF7DF93AFEFDF97D" bold="true" box="[136,259,1685,1709]" pageId="8" pageNumber="463">Diagnosis.</emphasis>
Medium-sized (CW: 
<quantity id="4CEC9B49FFDDE062FDF3F93AFD79F97D" box="[518,647,1685,1709]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.4" metricValueMin="2.8" pageId="8" pageNumber="463" unit="mm" value="36.0" valueMax="44.0" valueMin="28.0">28–44 mm</quantity>
). Carapace about 1.3 times broader than long, transverse, low; dorsal surface flat, glabrous; regions poorly defined, cervical groves indiscernible, H-shaped depression very shallow. Anterolateral margins of carapace very weakly serrated (in males) and nearly smooth (in females); postorbital cristae indistinct, smooth, not obviously confluent with epibranchial tooth. Epibranchial tooth very small, low, not prominent. Posterior margin of epistome with low median blunt tooth. Exopod of third maxilliped with very short flagellum, about 0.1 times width of merus. Ambulatory legs long and slender; 4 
<superScript id="7C619BE4FFDDE062FF63F85CFF5CF7D1" attach="left" box="[150,162,2035,2049]" fontSize="6" pageId="8" pageNumber="463">th</superScript>
pair with length of merus about 4.1 times width. G1 terminal segment curved outwards, hook-shaped, without dorsal flap in posterior part. Telson triangular with lateral margins gently convex.
</paragraph>
</subSubSection>
<subSubSection id="C30E6527FFDDE063FCB6FB5AFE60F8C5" lastPageId="9" lastPageNumber="464" pageId="8" pageNumber="463" type="description">
<paragraph id="8BAB36ACFFDDE062FCB6FB5AFB31F7FD" blockId="8.[834,1476,1269,2094]" pageId="8" pageNumber="463">
<emphasis id="B960EABEFFDDE062FCB6FB5AFC28FADD" bold="true" box="[835,982,1269,1293]" pageId="8" pageNumber="463">Description.</emphasis>
Carapace transverse, low, about 1.3 time broader than long (n=9); dorsal surface flat, glabrous; regions poorly defined, cervical grooves indiscernible, H-shaped depression very shallow, indiscernible (
<figureCitation id="132F2A29FFDDE062FAAAFAF9FA48FABD" box="[1375,1462,1365,1390]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="8" pageNumber="463">Fig. 5A</figureCitation>
). Epigastric cristae very low, weak and smooth, separated by very short, shallow groove that opens up into inverted V-shape posteriorly, no groove between epigastric cristae and postorbital cristae; postorbital cristae weakens, indistinct, smooth, not obviously confluent with epibranchial tooth; regions behind epigastric and postorbital cristae smooth (
<figureCitation id="132F2A29FFDDE062FA6DF9B9FC98F99D" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="8" pageNumber="463">Fig. 5A</figureCitation>
). Frontal margin slightly emarginate medially; frontal region turned downwards, smooth; supra- and infraorbital margins cristate, supraorbital margin sinuous, infraorbital margin almost straight; orbital region smooth, relatively narrow; subhepatic and subbranchial regions smooth (
<figureCitation id="132F2A29FFDDE062FA79F919FC96F93D" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="8" pageNumber="463">Figs. 5A</figureCitation>
, 
<figureCitation id="132F2A29FFDDE062FC8DF97AFC65F93D" box="[888,923,1749,1773]" captionStart="Fig" captionStartId="6.[136,171,1279,1301]" captionTargetBox="[135,1473,189,1264]" captionTargetId="figure-318@6.[135,1473,189,1264]" captionTargetPageId="6" captionText="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." figureDoi="http://doi.org/10.5281/zenodo.4504488" httpUri="https://zenodo.org/record/4504488/files/figure.png" pageId="8" pageNumber="463">6B</figureCitation>
). External orbital angle triangular, outer margin convex, weakly cristate to smooth; epibranchial tooth very small and low, separated from external orbital angle by narrow and shallow triangular cleft; anterolateral margin slightly convex, very weakly serrated (in males) and nearly smooth (in females); posterolateral margin entire, almost straight, not strongly convergent posteriorly; branchial and metabranchial regions smooth (
<figureCitation id="132F2A29FFDDE062FB59F819FAEFF81D" box="[1196,1297,1973,1998]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="8" pageNumber="463">Figs. 5A</figureCitation>
, 
<figureCitation id="132F2A29FFDDE062FAD4F81AFABAF81D" box="[1313,1348,1973,1997]" captionStart="Fig" captionStartId="6.[136,171,1279,1301]" captionTargetBox="[135,1473,189,1264]" captionTargetId="figure-318@6.[135,1473,189,1264]" captionTargetPageId="6" captionText="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." figureDoi="http://doi.org/10.5281/zenodo.4504488" httpUri="https://zenodo.org/record/4504488/files/figure.png" pageId="8" pageNumber="463">6B</figureCitation>
). Epistome anterior margin with median triangle; posterior margin with low median blunt tooth, slightly crenulated laterally, with outer part deeply concave (
<figureCitation id="132F2A29FFDDE062FB98F7B9FB3DF7FD" box="[1133,1219,2069,2094]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="8" pageNumber="463">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FF12FDEFFEA5" blockId="9.[113,753,189,373]" pageId="9" pageNumber="464">
Ischium of third maxilliped subrectangular, about 1.4 times longer than broad, with shallow longitudinal median sulcus; merus squarish, longer than half (0.6 times) of ischium length; exopod relatively short, equal or slightly exceeding upper edge of ischium, with very short flagellum, about 0.1 times width of merus (n=9) (
<figureCitation id="132F2A29FFDCE063FE5BFEF2FDFBFEA5" box="[430,517,349,373]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="9" pageNumber="464">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FE32FD8DFD85" blockId="9.[112,753,413,597]" pageId="9" pageNumber="464">
Chelipeds (
<figureCitation id="132F2A29FFDCE063FF00FE32FEAFFE65" box="[245,337,413,437]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="9" pageNumber="464">Fig. 5D</figureCitation>
) subequal; outer surface smooth to slightly rugose, subequal in length to palm, tips overlapping. Right chelipeds with length of palm+pollex about 3.1 times palm height, carpus with smooth outer surface, with strong, obliquely directed, subdistal spine on inner margin; merus with serrated edges, without subterminal spine.
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FDD2FD40FC65" blockId="9.[112,753,637,949]" pageId="9" pageNumber="464">
Ambulatory legs (
<figureCitation id="132F2A29FFDCE063FED8FDD2FE80FD45" box="[301,382,637,661]" captionStart="Fig" captionStartId="5.[113,148,1360,1382]" captionTargetBox="[113,1451,189,1344]" captionTargetId="figure-331@5.[113,1451,189,1344]" captionTargetPageId="5" captionText="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi="http://doi.org/10.5281/zenodo.4504486" httpUri="https://zenodo.org/record/4504486/files/figure.png" pageId="9" pageNumber="464">Fig. 5A</figureCitation>
) glabrous, long and slender; second leg with dactylus about 8.4 times longer than proximal width, propodus about 4.7 times longer than broad and equal or slightly longer than dactylus, carpus about 0.6 times length of dactylus, merus about 1.4 times longer than dactylus (n=9); fourth leg with dactylus about 8.3 times longer than proximal width, propodus slightly longer than dactylus, carpus about 0.7 times length of dactylus (n=8), merus without serrated upper margins, about 4.1 times longer than proximal width (n=8) and about 1.4 times longer than dactylus (n=8).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FC72FEBFFB45" blockId="9.[112,752,989,1173]" pageId="9" pageNumber="464">
Suture between anterior thoracic sternites 2 and 3 complete, distinct, slightly convex in the middle; groove between sternites 3 and 4 complete, distinct (
<figureCitation id="132F2A29FFDCE063FE17FBB2FDCBFBE5" box="[482,565,1053,1077]" captionStart="Fig" captionStartId="7.[113,148,1958,1980]" captionTargetBox="[114,1452,294,1941]" captionTargetId="figure-14@7.[107,1465,287,1955]" captionTargetPageId="7" captionText="Fig. 7. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." figureDoi="http://doi.org/10.5281/zenodo.4504490" httpUri="https://zenodo.org/record/4504490/files/figure.png" pageId="9" pageNumber="464">Fig. 7A</figureCitation>
). Male abdominal cavity reaching imaginary line joining posterior points of cheliped bases (below the suture between thoracic sternites 3 and 4) (
<figureCitation id="132F2A29FFDCE063FF2AFBD2FECBFB45" box="[223,309,1149,1173]" captionStart="Fig" captionStartId="7.[113,148,1958,1980]" captionTargetBox="[114,1452,294,1941]" captionTargetId="figure-14@7.[107,1465,287,1955]" captionTargetPageId="7" captionText="Fig. 7. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." figureDoi="http://doi.org/10.5281/zenodo.4504490" httpUri="https://zenodo.org/record/4504490/files/figure.png" pageId="9" pageNumber="464">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FB12FD3EFAA5" blockId="9.[113,754,1213,1398]" pageId="9" pageNumber="464">
Male abdomen broadly triangular; telson triangular, broader than long (about 1.4 times), lateral margins gently convex, tip rounded, subequal in length to sixth segment; segment 6 with proximal width about 2.2 times length, lateral margins almost straight; lateral margins of segments 4 to 5 straight; lateral margins of segment 3 gently convex (
<figureCitation id="132F2A29FFDCE063FDA9FAF2FD4AFAA5" box="[604,692,1373,1397]" captionStart="Fig" captionStartId="6.[136,171,1279,1301]" captionTargetBox="[135,1473,189,1264]" captionTargetId="figure-318@6.[135,1473,189,1264]" captionTargetPageId="6" captionText="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." figureDoi="http://doi.org/10.5281/zenodo.4504488" httpUri="https://zenodo.org/record/4504488/files/figure.png" pageId="9" pageNumber="464">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84FA32FE60F8C5" blockId="9.[112,754,1437,1813]" pageId="9" pageNumber="464">
Gl gently sinuous, reaching just to over half of the fourth sternum; terminal segment slender, about 0.4 times length of subterminal segment, about 2.9 times longer than proximal width, without dorsal flap, curved outwards, hook-shaped, opening on tip visible from dorsal and ventral views, distal part of groove for G2 visible on ventral side, tip truncated; subterminal segment relatively slender, sinuous, without neck-like distal part (
<figureCitation id="132F2A29FFDCE063FE9CF9D2FE30F945" box="[361,462,1661,1685]" captionStart="Fig" captionStartId="6.[136,171,1279,1301]" captionTargetBox="[135,1473,189,1264]" captionTargetId="figure-318@6.[135,1473,189,1264]" captionTargetPageId="6" captionText="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." figureDoi="http://doi.org/10.5281/zenodo.4504488" httpUri="https://zenodo.org/record/4504488/files/figure.png" pageId="9" pageNumber="464">Figs. 6B</figureCitation>
, 
<figureCitation id="132F2A29FFDCE063FE2BF9D2FE15F945" box="[478,491,1661,1685]" captionStart="Fig" captionStartId="7.[113,148,1958,1980]" captionTargetBox="[114,1452,294,1941]" captionTargetId="figure-14@7.[107,1465,287,1955]" captionTargetPageId="7" captionText="Fig. 7. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." figureDoi="http://doi.org/10.5281/zenodo.4504490" httpUri="https://zenodo.org/record/4504490/files/figure.png" pageId="9" pageNumber="464">7</figureCitation>
A–D). G2 shorter than G1 (0.9 times length of G1) with distal segment about 0.3 times length of basal segment (
<figureCitation id="132F2A29FFDCE063FE4DF912FDDBF905" box="[440,549,1725,1749]" captionStart="Fig" captionStartId="7.[113,148,1958,1980]" captionTargetBox="[114,1452,294,1941]" captionTargetId="figure-14@7.[107,1465,287,1955]" captionTargetPageId="7" captionText="Fig. 7. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." figureDoi="http://doi.org/10.5281/zenodo.4504490" httpUri="https://zenodo.org/record/4504490/files/figure.png" pageId="9" pageNumber="464">Fig. 7E, F</figureCitation>
). Female gonopore in thoracic sternite 6, round, without operculum, opened posterio-mesially (
<figureCitation id="132F2A29FFDCE063FECEF952FE6FF8C5" box="[315,401,1789,1813]" captionStart="Fig" captionStartId="6.[136,171,1279,1301]" captionTargetBox="[135,1473,189,1264]" captionTargetId="figure-318@6.[135,1473,189,1264]" captionTargetPageId="6" captionText="Fig. 6. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 43.8 mm), IEBR-FC THx07." figureDoi="http://doi.org/10.5281/zenodo.4504488" httpUri="https://zenodo.org/record/4504488/files/figure.png" pageId="9" pageNumber="464">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E6527FFDCE063FF84F892FBF4FE65" pageId="9" pageNumber="464" type="etymology">
<paragraph id="8BAB36ACFFDCE063FF84F892FD48F8A5" blockId="9.[112,752,1853,1909]" pageId="9" pageNumber="464">
<emphasis id="B960EABEFFDCE063FF84F892FF0DF885" bold="true" box="[113,243,1853,1877]" pageId="9" pageNumber="464">Etymology.</emphasis>
The new species is named after the 
<typeStatus id="54AF880EFFDCE063FD92F891FD6BF886" box="[615,661,1854,1878]" pageId="9" pageNumber="464">type</typeStatus>
locality, Ham Yen. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8BAB36ACFFDCE063FF84F832FE2BF825" blockId="9.[113,752,1949,2038]" pageId="9" pageNumber="464">
<emphasis id="B960EABEFFDCE063FF84F832FEC8F865" bold="true" box="[113,310,1949,1973]" pageId="9" pageNumber="464">Live colouration.</emphasis>
Carapace and chela milky to light gray, ambulatory legs vary from milky to light gray or purple mixed light orange (
<figureCitation id="132F2A29FFDCE063FEBAF872FE71F825" box="[335,399,2013,2037]" captionStart="Fig" captionStartId="8.[135,170,1096,1118]" captionTargetBox="[136,1474,189,1081]" captionTargetId="figure-465@8.[136,1474,189,1081]" captionTargetPageId="8" captionText="Fig. 8. A–D, Tiwaripotamon hamyen, new species, with differences living colour in carapace and ambulatory legs." figureDoi="http://doi.org/10.5281/zenodo.4504492" httpUri="https://zenodo.org/record/4504492/files/figure.png" pageId="9" pageNumber="464">Fig. 8</figureCitation>
A–D).
</paragraph>
<paragraph id="8BAB36ACFFDCE063FCD8FF12FBF4FE65" blockId="9.[811,1453,189,437]" pageId="9" pageNumber="464">
<emphasis id="B960EABEFFDCE063FCD8FF12FC16FF05" bold="true" box="[813,1000,189,213]" pageId="9" pageNumber="464">Ecological notes.</emphasis>
This species inhabits limestone mountains. Our observation of this species and other 
<taxonomicName id="4C144D2FFFDCE063FAF0FF71FA55FF25" authorityName="Bott" authorityYear="1970" box="[1285,1451,222,245]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B960EABEFFDCE063FAF0FF71FA55FF25" box="[1285,1451,222,245]" italics="true" pageId="9" pageNumber="464">Tiwaripotamon</emphasis>
</taxonomicName>
species from 
<collectingCountry id="F303763CFFDCE063FC3DFF52FBD9FEC5" box="[968,1063,253,277]" name="Vietnam" pageId="9" pageNumber="464">Vietnam</collectingCountry>
confirm that they have terrestrial habits. They were often found far from water sources like streams and ponds. We observed a female 
<taxonomicName id="4C144D2FFFDCE063FAD3FE91FA55FE85" authorityName="Ng &amp; Yeo" authorityYear="2001" box="[1318,1451,317,341]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="edostilus">
<emphasis id="B960EABEFFDCE063FAD3FE91FA55FE85" box="[1318,1451,317,341]" italics="true" pageId="9" pageNumber="464">T. edostilus</emphasis>
</taxonomicName>
carrying juveniles walking on the forest floor not close to any water body. It means these juveniles did not develop in streams or ponds.
</paragraph>
</subSubSection>
<subSubSection id="C30E6527FFDCE063FCD8FE72FC16FC05" pageId="9" pageNumber="464" type="discussion">
<paragraph id="8BAB36ACFFDCE063FCD8FE72FC16FC05" blockId="9.[811,1454,477,981]" pageId="9" pageNumber="464">
<emphasis id="B960EABEFFDCE063FCD8FE72FC65FE25" bold="true" box="[813,923,477,501]" pageId="9" pageNumber="464">Remarks.</emphasis>
<taxonomicName id="4C144D2FFFDCE063FC56FE71FB5DFE25" authorityName="Do &amp; Nguyen &amp; Dang" authorityYear="2017" box="[931,1187,477,501]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="hamyen">
<emphasis id="B960EABEFFDCE063FC56FE71FB5DFE25" box="[931,1187,477,501]" italics="true" pageId="9" pageNumber="464">Tiwaripotamon hamyen</emphasis>
</taxonomicName>
is easily separated from all other 
<taxonomicName id="4C144D2FFFDCE063FC60FE51FBC3FDC5" authorityName="Bott" authorityYear="1970" box="[917,1085,510,533]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B960EABEFFDCE063FC60FE51FBC3FDC5" box="[917,1085,510,533]" italics="true" pageId="9" pageNumber="464">Tiwaripotamon</emphasis>
</taxonomicName>
species by: (1) median tooth of epistome posterior margin is low and blunt (vs. median tooth of epistome posterior margin is triangular); (2) ambulatory legs are relatively less longer and slender compared to the congeners (the merus of 4 
<superScript id="7C619BE4FFDCE063FBB2FDD4FBADFD59" attach="left" box="[1095,1107,635,649]" fontSize="6" pageId="9" pageNumber="464">th</superScript>
pair leg of this species is about 4.1 longer than broad compared to 4.3 for 
<taxonomicName id="4C144D2FFFDCE063FAF7FD31FA5BFD65" authorityName="Dai &amp; Naiyanetr" authorityYear="1994" box="[1282,1445,670,693]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="pingguoense">
<emphasis id="B960EABEFFDCE063FAF7FD31FA5BFD65" box="[1282,1445,670,693]" italics="true" pageId="9" pageNumber="464">T. pingguoense</emphasis>
</taxonomicName>
, 4.5 for 
<taxonomicName id="4C144D2FFFDCE063FC72FD11FBD9FD05" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[903,1063,702,725]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B960EABEFFDCE063FC72FD11FBD9FD05" box="[903,1063,702,725]" italics="true" pageId="9" pageNumber="464">T. annamense</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EF854B5DFFDCE063FB9EFD12FADCFD05" author="Do VT &amp; Shih HT &amp; Huang C" box="[1131,1314,701,725]" pageId="9" pageNumber="464" pagination="213 - 219" refId="ref5384" refString="Do VT, Shih HT &amp; Huang C (2016) A new species of freshwater crab Tiwaripotamon Bott, 1970 (Crustacea, Brachyura, Potamidae) from northern Vietnam and southern China. Raffles Bulletin of Zoology, 64: 213 - 219." type="journal article" year="2016">Do et al., 2016</bibRefCitation>
), 4.8 for 
<taxonomicName id="4C144D2FFFDCE063FA60FD11FC65FD25" authorityName="Do et al" authorityYear="2016" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="pluviosum">
<emphasis id="B960EABEFFDCE063FA60FD11FC65FD25" italics="true" pageId="9" pageNumber="464">T. pluviosum</emphasis>
</taxonomicName>
, 5.1 times for 
<taxonomicName id="4C144D2FFFDCE063FBB1FD71FB13FD25" baseAuthorityName="Dang &amp; Ho" baseAuthorityYear="2002" box="[1092,1261,734,757]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="vietnamicum">
<emphasis id="B960EABEFFDCE063FBB1FD71FB13FD25" box="[1092,1261,734,757]" italics="true" pageId="9" pageNumber="464">T. vietnamicum</emphasis>
</taxonomicName>
, 5.3 times for 
<taxonomicName id="4C144D2FFFDCE063FA63FD71FC61FCC5" authorityName="Dai &amp; Naiyanetr" authorityYear="1994" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="xiurenense">
<emphasis id="B960EABEFFDCE063FA63FD71FC61FCC5" italics="true" pageId="9" pageNumber="464">T. xiurenense</emphasis>
</taxonomicName>
(unpublished data), 5.6 times for large 
<taxonomicName id="4C144D2FFFDCE063FAC7FD51FA54FCC5" authorityName="Ng &amp; Yeo" authorityYear="2001" box="[1330,1450,765,789]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="edostilus">
<emphasis id="B960EABEFFDCE063FAC7FD51FA54FCC5" box="[1330,1450,765,789]" italics="true" pageId="9" pageNumber="464">T. edostilus</emphasis>
</taxonomicName>
specimens, and 6.5 for 
<taxonomicName id="4C144D2FFFDCE063FBD8FCB1FA56FCE5" authority="(Shih &amp; Do, 2014)" baseAuthorityName="Shih &amp; Do" baseAuthorityYear="2014" box="[1069,1448,797,821]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="vixuyenense">
<emphasis id="B960EABEFFDCE063FBD8FCB1FB32FCE5" box="[1069,1228,798,821]" italics="true" pageId="9" pageNumber="464">T. vixuyenense</emphasis>
(
<bibRefCitation id="EF854B5DFFDCE063FB28FCB2FA5EFCE5" author="Shih HT &amp; Do VT" box="[1245,1440,797,821]" pageId="9" pageNumber="464" pagination="26 - 38" refId="ref5617" refString="Shih HT &amp; Do VT (2014) A new species of Tiwaripotamon Bott, 1970, from northern Vietnam, with notes on T. vietnamicum (Dang &amp; Ho, 2002) and T. edostilus Ng &amp; Yeo, 2001 (Crustacea, Brachyura, Potamidae). Zootaxa, 3764: 26 - 38." type="journal article" year="2014">Shih &amp; Do, 2014</bibRefCitation>
)
</taxonomicName>
; (3) G1 terminal segment is curved outwards, hook shape (vs. curved upwards, not hook shape) (
<bibRefCitation id="EF854B5DFFDCE063FB12FCF2FC98FC45" author="Dai AY &amp; Naiyanetr P" pageId="9" pageNumber="464" pagination="47 - 72" refId="ref5246" refString="Dai AY &amp; Naiyanetr P (1994) A revision of the genus Tiwaripotamon Bott, 1970, the freshwater crabs from China (Decapoda: Brachyura: Potamidae). Sinozoologia, 11: 47 - 72." type="journal article" year="1994">Dai &amp; Naiyanetr, 1994</bibRefCitation>
: 
<figureCitation id="132F2A29FFDCE063FC86FCD2FC25FC45" box="[883,987,893,917]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[113,148,1262,1284]" captionStartId-1="2.[136,171,1240,1262]" captionTargetBox-0="[113,1450,190,1249]" captionTargetBox-1="[136,1474,190,1225]" captionTargetId-0="figure-14@1.[100,1455,175,1250]" captionTargetId-1="figure-381@2.[136,1474,190,1225]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Collection sites of Tiwaripotamon xuanson, new species and T. hamyen, new species." captionText-1="Fig. 2. Tiwaripotamon xuanson, new species, holotype, male (CW 33.1 mm), IEBR-FC TXx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi-0="http://doi.org/10.5281/zenodo.4504478" figureDoi-1="http://doi.org/10.5281/zenodo.4504480" httpUri-0="https://zenodo.org/record/4504478/files/figure.png" httpUri-1="https://zenodo.org/record/4504480/files/figure.png" pageId="9" pageNumber="464">Figs. 1–2</figureCitation>
; 
<bibRefCitation id="EF854B5DFFDCE063FC1FFCD2FB59FC45" author="Ng PKL &amp; Yeo DCJ" box="[1002,1191,893,917]" pageId="9" pageNumber="464" pagination="275 - 287" refId="ref5474" refString="Ng PKL &amp; Yeo DCJ (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21: 275 - 287." type="journal article" year="2001">Ng &amp; Yeo, 2001</bibRefCitation>
: 
<figureCitation id="132F2A29FFDCE063FB40FCD2FAE0FC45" box="[1205,1310,893,917]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="1.[113,148,1262,1284]" captionStartId-1="2.[136,171,1240,1262]" captionStartId-2="3.[112,147,1310,1332]" captionStartId-3="4.[135,170,2024,2046]" captionStartId-4="5.[113,148,1360,1382]" captionTargetBox-0="[113,1450,190,1249]" captionTargetBox-1="[136,1474,190,1225]" captionTargetBox-2="[113,1451,189,1295]" captionTargetBox-3="[136,1472,228,2009]" captionTargetBox-4="[113,1451,189,1344]" captionTargetId-0="figure-14@1.[100,1455,175,1250]" captionTargetId-1="figure-381@2.[136,1474,190,1225]" captionTargetId-2="figure-310@3.[113,1451,189,1295]" captionTargetId-3="figure-8@4.[121,1474,218,2017]" captionTargetId-4="figure-331@5.[113,1451,189,1344]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="Fig. 1. Collection sites of Tiwaripotamon xuanson, new species and T. hamyen, new species." captionText-1="Fig. 2. Tiwaripotamon xuanson, new species, holotype, male (CW 33.1 mm), IEBR-FC TXx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." captionText-2="Fig. 3. Tiwaripotamon xuanson, new species, holotype, male (CW 33.1 mm), IEBR-FC TXx01. A, ventral view; B, sternoabdominal cavity showing G1s and G2s; C, female gonopore (CW 32.6 mm), IEBR-FC TXx06; D, A habitat in Ten mountain, Xuan Son National Park, Thanh Son District, Phu Tho Province, Vietnam." captionText-3="Fig. 4. Tiwaripotamon xuanson, new species, holotype, male (CW 33.1 mm), IEBR-FC TXx01. A–F, right G1: A, ventral view; B, ventral view of terminal segment; C, dorsal view; D, dorsal view of terminal segment; E, right G2; F, right G2 of terminal segment." captionText-4="Fig. 5. Tiwaripotamon hamyen, new species, holotype, male (CW 41.7 mm), IEBR-FC THx01. A, carapace, dorsal view; B, frontal view; C, left third maxilliped; D, chela outer view." figureDoi-0="http://doi.org/10.5281/zenodo.4504478" figureDoi-1="http://doi.org/10.5281/zenodo.4504480" figureDoi-2="http://doi.org/10.5281/zenodo.4504482" figureDoi-3="http://doi.org/10.5281/zenodo.4504484" figureDoi-4="http://doi.org/10.5281/zenodo.4504486" httpUri-0="https://zenodo.org/record/4504478/files/figure.png" httpUri-1="https://zenodo.org/record/4504480/files/figure.png" httpUri-2="https://zenodo.org/record/4504482/files/figure.png" httpUri-3="https://zenodo.org/record/4504484/files/figure.png" httpUri-4="https://zenodo.org/record/4504486/files/figure.png" pageId="9" pageNumber="464">Figs. 1–5</figureCitation>
; 
<bibRefCitation id="EF854B5DFFDCE063FAD9FCD2FC96FC65" author="Shih HT &amp; Do VT" pageId="9" pageNumber="464" pagination="26 - 38" refId="ref5617" refString="Shih HT &amp; Do VT (2014) A new species of Tiwaripotamon Bott, 1970, from northern Vietnam, with notes on T. vietnamicum (Dang &amp; Ho, 2002) and T. edostilus Ng &amp; Yeo, 2001 (Crustacea, Brachyura, Potamidae). Zootaxa, 3764: 26 - 38." type="journal article" year="2014">Shih &amp; Do, 2014</bibRefCitation>
: Figs: 2–7; 
<bibRefCitation id="EF854B5DFFDCE063FC00FC32FB34FC65" author="Dang NT &amp; Ho TH" box="[1013,1226,925,949]" pageId="9" pageNumber="464" refId="ref5322" refString="Dang NT &amp; Ho TH (2012) Tom, Cua nuoc ngot Viet Nam (Palaemonidae, Atyidae, Parathelphusidae, Potamidae). (Freshwater Crabs and Shrimps from Vietnam [Palaemonidae, Atyidae, Parathelphusidae, Potamidae]). Publishing House for Science and Technology, Hanoi, Vietnam, 264 pp. [In Vietnamese]." type="book" year="2012">Dang &amp; Ho, 2012</bibRefCitation>
: Fig. 78; 
<bibRefCitation id="EF854B5DFFDCE063FACBFC32FC98FC05" author="Do VT &amp; Shih HT &amp; Huang C" pageId="9" pageNumber="464" pagination="213 - 219" refId="ref5384" refString="Do VT, Shih HT &amp; Huang C (2016) A new species of freshwater crab Tiwaripotamon Bott, 1970 (Crustacea, Brachyura, Potamidae) from northern Vietnam and southern China. Raffles Bulletin of Zoology, 64: 213 - 219." type="journal article" year="2016">Do et al., 2016</bibRefCitation>
: Figs: 1–5).
</paragraph>
</subSubSection>
</treatment>
</document>