treatments-xml/data/03/BD/87/03BD87B6FFBFF431CFDB736AFF0AFE73.xml

<document ID-DOI="10.11646/zootaxa.5277.1.4" ID-GBIF-Dataset="ad5ba4d1-c2ec-42e3-a76d-a5c9b395a6da" ID-ISSN="1175-5326" ID-Zenodo-Dep="7892052" ID-ZooBank="272D07DF-8471-4ADE-9689-DB8F588ACF35" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682930765663" checkinUser="plazi" docAuthor="Short, Andrew E. Z., Torres-Gavosto, Juan Martin &amp; Hettinger, Ty" docDate="2023" docId="03BD87B6FFBFF431CFDB736AFF0AFE73" docLanguage="en" docName="zootaxa.5277.1.4.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oocyclus radiatus Short &amp; Torres-Gavosto &amp; Hettinger 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="102" masterDocId="FF84FFCEFFB5F43ACF4C7204FFE1FFDE" masterDocTitle="A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="112" masterPageNumber="91" pageNumber="101" updateTime="1683122599468" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew E. Z.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7467-7116</mods:nameIdentifier>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. &amp; aezshort @ gmail. com; https: // orcid. org / 0000 - 0002 - 7467 - 7116</mods:affiliation>
<mods:nameIdentifier type="email">aezshort@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torres-Gavosto, Juan Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">0009-0007-9274-2923</mods:nameIdentifier>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. &amp; https: // orcid. org / 0009 - 0007 - 9274 - 2923</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hettinger, Ty</mods:namePart>
<mods:affiliation>Division of Entomology, Biodiversity Institute &amp; Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-01</mods:number>
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<mods:detail type="volume">
<mods:number>5277</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>91</mods:start>
<mods:end>112</mods:end>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7892052</mods:identifier>
<mods:identifier type="ZooBank">272D07DF-8471-4ADE-9689-DB8F588ACF35</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7892084" ID-Zenodo-Dep="7892084" LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73" httpUri="http://treatment.plazi.org/id/03BD87B6FFBFF431CFDB736AFF0AFE73" lastPageId="11" lastPageNumber="102" pageId="10" pageNumber="101">
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<heading bold="true" box="[151,442,366,393]" fontSize="11" level="1" pageId="10" pageNumber="101" reason="1">
<emphasis bold="true" box="[151,442,366,393]" pageId="10" pageNumber="101">
<taxonomicName authority="Short &amp; Torres-Gavosto &amp; Hettinger, 2023" authorityName="Short &amp; Torres-Gavosto &amp; Hettinger" authorityYear="2023" box="[151,368,366,393]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="radiatus" status="sp. n.">
<emphasis bold="true" box="[151,368,366,393]" italics="true" pageId="10" pageNumber="101">Oocyclus radiatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[376,442,367,393]" pageId="10" pageNumber="101" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,341,403,429]" pageId="10" pageNumber="101" type="description">
<paragraph blockId="10.[151,442,366,429]" box="[151,341,403,429]" pageId="10" pageNumber="101">
(
<figureCitation box="[159,259,403,429]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="10" pageNumber="101">Figs. 3A</figureCitation>
, 
<figureCitation box="[271,306,403,429]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="10" pageNumber="101">6B</figureCitation>
, 
<figureCitation box="[318,333,403,429]" captionStart="FIGURE 7" captionStartId="19.[151,250,1886,1910]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@19.[151,1436,181,1868]" captionTargetPageId="19" captionText="FIGURE 7. Distribution of Oocyclus spp. in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.7892078" httpUri="https://zenodo.org/record/7892078/files/figure.png" pageId="10" pageNumber="101">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="101" type="materials_examined">
<paragraph blockId="10.[151,1437,474,2013]" pageId="10" pageNumber="101">
<emphasis bold="true" box="[151,328,474,501]" pageId="10" pageNumber="101">Type Material.</emphasis>
<materialsCitation collectingDate="1975-07-10" collectionCode="ECUADOR-PEACE, CORPS, USNM" pageId="10" pageNumber="101" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[335,537,474,501]" pageId="10" pageNumber="101">
<typeStatus box="[335,444,475,501]" pageId="10" pageNumber="101">Holotype</typeStatus>
(male):
</emphasis>
“ECUADOR, Pich.,/ Sto. Domingo de/ los Colorados, / 44KmE., 
<date box="[1283,1427,474,501]" pageId="10" pageNumber="101" value="1975-07-10">
<collectingDate box="[1283,1427,474,501]" pageId="10" pageNumber="101" value="1975-07-10">10 July 1975</collectingDate>
</date>
/ Jeffrey Cohen”, “ 
<collectionCode box="[343,676,511,537]" pageId="10" pageNumber="101">ECUADOR-PEACE CORPS</collectionCode>
,/ SMITHSONIAN INSTITUTION/ AQUATIC INSECT SURVEY” (
<collectionCode box="[159,246,547,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<emphasis bold="true" box="[269,511,546,573]" pageId="10" pageNumber="101">
<typeStatus box="[269,389,547,573]" pageId="10" pageNumber="101">Paratypes</typeStatus>
(
<specimenCount box="[408,497,546,572]" pageId="10" pageNumber="101" type="generic">77 exs.</specimenCount>
):
</emphasis>
<materialsCitation collectingDate="1975-05-08" collectionCode="USNM, SEMC" collectorName="P. Spangler; A. Langley &amp; J. Cohen" country="Ecuador" location="Colorados" pageId="10" pageNumber="101" specimenCount="20" stateProvince="Manabi" typeStatus="paratype">
<emphasis bold="true" box="[522,903,546,573]" pageId="10" pageNumber="101">
<collectingCountry box="[522,668,546,572]" name="Ecuador" pageId="10" pageNumber="101">ECUADOR</collectingCountry>
: 
<collectingRegion box="[684,898,546,573]" country="Ecuador" name="Manabi" pageId="10" pageNumber="101">Manabí Province</collectingRegion>
:
</emphasis>
<quantity box="[914,989,547,573]" metricMagnitude="4" metricUnit="m" metricValue="7.9" pageId="10" pageNumber="101" unit="km" value="79.0">79 km</quantity>
<collectorName box="[999,1222,547,573]" pageId="10" pageNumber="101">W. Santo Domingo</collectorName>
de los 
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73:8ECB607BFFBFF430CA6E7027FA79FDE3" box="[1314,1432,547,573]" country="Ecuador" name="Colorados" pageId="10" pageNumber="101" stateProvince="Manabi">Colorados</location>
, 
<date box="[151,250,582,609]" pageId="10" pageNumber="101" value="1975-05-08">
<collectingDate box="[151,250,582,609]" pageId="10" pageNumber="101" value="1975-05-08">8.v.1975</collectingDate>
</date>
, leg. 
<collectorName box="[307,433,583,609]" pageId="10" pageNumber="101">P. Spangler</collectorName>
, 
<collectorName box="[442,568,583,609]" pageId="10" pageNumber="101">A. Langley</collectorName>
, &amp; 
<collectorName box="[605,704,583,609]" pageId="10" pageNumber="101">J. Cohen</collectorName>
, seepage over rock outcrops in roadcuts (
<specimenCount box="[1157,1239,583,609]" pageId="10" pageNumber="101" type="generic">20 exs.</specimenCount>
, 
<collectionCode box="[1249,1333,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
, 
<collectionCode box="[1343,1424,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="101" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation box="[151,1187,618,645]" collectingDate="1978-01-08" collectionCode="USNM" collectorName="P. Spangler" country="Ecuador" elevation="380" location="Flavio Alfaro" pageId="10" pageNumber="101" specimenCount="3" stateProvince="Manabi" typeStatus="paratype">
<quantity box="[151,245,618,645]" metricMagnitude="4" metricUnit="m" metricValue="2.65" pageId="10" pageNumber="101" unit="km" value="26.5">26.5 km</quantity>
E. 
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73:8ECB607BFFBFF430CE52706FFE59FD5B" box="[286,440,619,645]" country="Ecuador" name="Flavio Alfaro" pageId="10" pageNumber="101" stateProvince="Manabi">Flavio Alfaro</location>
, 
<quantity box="[453,527,619,645]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="101" unit="m" value="380.0">
<elevation box="[453,527,619,645]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="101" unit="m" value="380.0">380 m</elevation>
</quantity>
, 
<date box="[541,634,619,645]" pageId="10" pageNumber="101" value="1978-01-08">
<collectingDate box="[541,634,619,645]" pageId="10" pageNumber="101" value="1978-01-08">8.i.1978</collectingDate>
</date>
, leg. 
<collectorName box="[698,826,619,645]" pageId="10" pageNumber="101">P. Spangler</collectorName>
, roadside seep (
<specimenCount box="[1010,1080,619,645]" pageId="10" pageNumber="101" type="generic">3 exs.</specimenCount>
, 
<collectionCode box="[1093,1179,619,645]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1975-05-11" collectionCode="USNM, SEMC" collectorName="P. Spangler; A. Langley &amp; J. Cohen" country="Ecuador" location="Portoviejo" pageId="10" pageNumber="101" specimenCount="35" stateProvince="Manabi" typeStatus="paratype">
<quantity box="[1199,1272,619,645]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="101" unit="km" value="38.0">38 km</quantity>
E. 
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73:8ECB607BFFBFF430CA6D706FFA78FD5B" box="[1313,1433,619,645]" country="Ecuador" name="Portoviejo" pageId="10" pageNumber="101" stateProvince="Manabi">Portoviejo</location>
, 
<date box="[151,263,654,681]" pageId="10" pageNumber="101" value="1975-05-11">
<collectingDate box="[151,263,654,681]" pageId="10" pageNumber="101" value="1975-05-11">11.v.1975</collectingDate>
</date>
, leg. 
<collectorName box="[323,451,655,681]" pageId="10" pageNumber="101">P. Spangler</collectorName>
, 
<collectorName box="[461,588,655,681]" pageId="10" pageNumber="101">A. Langley</collectorName>
, &amp; 
<collectorName box="[629,730,655,681]" pageId="10" pageNumber="101">J. Cohen</collectorName>
, splash zone at waterfall, (
<specimenCount box="[1028,1111,654,681]" pageId="10" pageNumber="101" type="generic">35 exs.</specimenCount>
, 
<collectionCode box="[1123,1207,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
, 
<collectionCode box="[1218,1299,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="101" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation collectingDate="1975-07-05" collectionCode="USNM" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" location="Colorados" pageId="10" pageNumber="101" specimenCount="11" stateProvince="Pichincha" typeStatus="paratype">
<emphasis bold="true" pageId="10" pageNumber="101">
<collectingRegion country="Ecuador" name="Pichincha" pageId="10" pageNumber="101">Pichincha Province</collectingRegion>
:
</emphasis>
<quantity box="[274,347,691,717]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="101" unit="km" value="14.0">14 km</quantity>
<collectorName box="[355,566,691,717]" pageId="10" pageNumber="101">E. Santo Domingo</collectorName>
de los 
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73:8ECB607BFFBFF430CDC770B7FCE0FD13" box="[651,769,691,717]" country="Ecuador" name="Colorados" pageId="10" pageNumber="101" stateProvince="Pichincha">Colorados</location>
, 
<date box="[782,897,690,717]" pageId="10" pageNumber="101" value="1975-07-05">
<collectingDate box="[782,897,690,717]" pageId="10" pageNumber="101" value="1975-07-05">5.vii.1975</collectingDate>
</date>
, leg. 
<collectorName box="[959,1087,691,717]" pageId="10" pageNumber="101">A. Langley</collectorName>
&amp; 
<collectorName box="[1125,1225,691,717]" pageId="10" pageNumber="101">J. Cohen</collectorName>
(
<specimenCount box="[1242,1325,691,717]" pageId="10" pageNumber="101" type="generic">11 exs.</specimenCount>
, 
<collectionCode box="[1338,1424,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation box="[151,1281,726,753]" collectingDate="1975-07-05" collectionCode="USNM" collectorName="A. Langley &amp; J. Cohen" country="Ecuador" location="Colorados" pageId="10" pageNumber="101" specimenCount="8" stateProvince="Pichincha" typeStatus="paratype">
<quantity box="[151,223,726,753]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="101" unit="km" value="25.0">25 km</quantity>
<collectorName box="[230,440,727,753]" pageId="10" pageNumber="101">E. Santo Domingo</collectorName>
de los 
<location LSID="urn:lsid:plazi:treatment:03BD87B6FFBFF431CFDB736AFF0AFE73:8ECB607BFFBFF430CD4670D3FD61FD2F" box="[522,640,727,753]" country="Ecuador" name="Colorados" pageId="10" pageNumber="101" stateProvince="Pichincha">Colorados</location>
, 
<date box="[651,767,726,753]" pageId="10" pageNumber="101" value="1975-07-05">
<collectingDate box="[651,767,726,753]" pageId="10" pageNumber="101" value="1975-07-05">5.vii.1975</collectingDate>
</date>
, leg. 
<collectorName box="[827,954,727,753]" pageId="10" pageNumber="101">A. Langley</collectorName>
&amp; 
<collectorName box="[991,1090,727,753]" pageId="10" pageNumber="101">J. Cohen</collectorName>
(
<specimenCount box="[1107,1175,727,753]" pageId="10" pageNumber="101" type="generic">8 exs.</specimenCount>
, 
<collectionCode box="[1187,1273,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1975-07-10" collectionCode="USNM, SEMC" country="Ecuador" pageId="10" pageNumber="101" specimenCount="9" stateProvince="Pichincha" typeStatus="paratype">
same data as holotype (
<specimenCount box="[267,337,763,789]" pageId="10" pageNumber="101" type="generic">9 exs.</specimenCount>
, 
<collectionCode box="[350,434,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
, 
<collectionCode box="[447,528,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="101" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation collectingDate="1975-05-18" collectionCode="USNM" collectorName="P. Spangler;A. Langley &amp; J. Cohen" country="Ecuador" location="Sucumbios Province" pageId="10" pageNumber="101" specimenCount="1" stateProvince="Sucumbios" typeStatus="paratype">
<emphasis bold="true" box="[548,803,762,789]" pageId="10" pageNumber="101">
<collectingRegion box="[548,798,762,789]" country="Ecuador" name="Sucumbios" pageId="10" pageNumber="101">Sucumbíos Province</collectingRegion>
:
</emphasis>
<quantity box="[811,898,763,789]" metricMagnitude="5" metricUnit="m" metricValue="1.46" pageId="10" pageNumber="101" unit="km" value="146.0">146 km</quantity>
<collectorName box="[906,1079,763,789]" pageId="10" pageNumber="101">W. Lago Agrio</collectorName>
, 
<date box="[1092,1205,762,789]" pageId="10" pageNumber="101" value="1975-05-18">
<collectingDate box="[1092,1205,762,789]" pageId="10" pageNumber="101" value="1975-05-18">18.v.1975</collectingDate>
</date>
, leg. 
<collectorName box="[1268,1397,763,789]" pageId="10" pageNumber="101">P. Spangler</collectorName>
, 
<collectorName pageId="10" pageNumber="101">A. Langley</collectorName>
, &amp; 
<collectorName box="[284,385,799,825]" pageId="10" pageNumber="101">J. Cohen</collectorName>
, roadside ditch (
<specimenCount box="[572,627,799,825]" pageId="10" pageNumber="101" type="generic">1 ex.</specimenCount>
, 
<collectionCode box="[638,724,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="101" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="101" type="diagnosis">
<paragraph blockId="10.[151,1437,474,2013]" pageId="10" pageNumber="101">
<emphasis bold="true" box="[199,322,834,860]" pageId="10" pageNumber="101">Diagnosis.</emphasis>
Larger species. Posterolateral corners of pronotum angulate, but not spinose. Prosternal carinae set with two spines anteriorly. Elytra with first row of systematic punctures forming a dense, broken row bearing erect setae. Elytral margins slightly explanate, without a fringe of long setae. Abdominal ventrites bicolored, with dark anterior margins and pale posterior margins.
</paragraph>
<paragraph blockId="10.[151,1437,474,2013]" pageId="10" pageNumber="101">
Of larger-bodied species in 
<collectingCountry box="[531,624,979,1005]" name="Ecuador" pageId="10" pageNumber="101">Ecuador</collectingCountry>
, 
<taxonomicName authorityName="Short &amp; Torres-Gavosto &amp; Hettinger" authorityYear="2023" box="[644,777,979,1005]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="radiatus">
<emphasis box="[644,777,979,1005]" italics="true" pageId="10" pageNumber="101">O. radiatus</emphasis>
</taxonomicName>
is most similar to 
<emphasis box="[1013,1140,979,1005]" italics="true" pageId="10" pageNumber="101">
<taxonomicName authorityName="Short &amp; Torres-Gavosto &amp; Hettinger" authorityYear="2023" box="[1013,1136,979,1005]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="pakcha">O. pakcha</taxonomicName>
,
</emphasis>
which also has angulate posterolateral pronotal corners, however it can easily be distinguished by the broken irregular first row of erect setae on the elytra while 
<taxonomicName authorityName="Short &amp; Torres-Gavosto &amp; Hettinger" authorityYear="2023" box="[426,541,1051,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="pakcha">
<emphasis box="[426,541,1051,1077]" italics="true" pageId="10" pageNumber="101">O. pakcha</emphasis>
</taxonomicName>
has a dense continuous row. It is also similar to 
<taxonomicName authorityName="Short &amp; Torres-Gavosto &amp; Hettinger" authorityYear="2023" box="[1074,1208,1051,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="ustulatus">
<emphasis box="[1074,1094,1051,1077]" italics="true" pageId="10" pageNumber="101">O</emphasis>
. 
<emphasis box="[1107,1208,1051,1077]" italics="true" pageId="10" pageNumber="101">ustulatus</emphasis>
</taxonomicName>
, but that species has spinose posterolateral angles of the pronotum and a pale yellow abdominal venter.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="102" pageId="10" pageNumber="101" type="description">
<paragraph blockId="10.[151,1437,474,2013]" pageId="10" pageNumber="101">
<emphasis bold="true" box="[199,524,1122,1149]" pageId="10" pageNumber="101">Description. Size and Form.</emphasis>
Length = 
<quantity box="[630,765,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.4" metricValueMin="4.3" pageId="10" pageNumber="101" unit="mm" value="4.85" valueMax="5.4" valueMin="4.3">4.3–5.4 mm</quantity>
.Very broadly oval, slightly convex.(
<figureCitation box="[1165,1252,1123,1149]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="10" pageNumber="101">Fig. 3A</figureCitation>
). 
<emphasis bold="true" box="[1269,1342,1122,1148]" pageId="10" pageNumber="101">Color.</emphasis>
Dorsum of head, pronotum, and elytra black, with faint iridescent green reflections. Anterolateral margins of pronotum with large, broad pale spot, extending nearly half the length of the margin. Elytra with very faint iridescent green maculae which often appear faint and with defuse borders. Maxillary and labial palps yellow. Mentum and stipes dark brown, similar in color to the venter of the head. Thoracic ventrites and legs are reddish to dark brown. Abdominal ventrites with a distinct banding pattern, with the anterior half of each ventrite reddish to dark brown and the posterior margin distinctly paler, being light brown to orange, with the thickness of the posterior orange band widening laterally.
</paragraph>
<paragraph blockId="10.[151,1437,474,2013]" lastBlockId="11.[151,1437,151,429]" lastPageId="11" lastPageNumber="102" pageId="10" pageNumber="101">
<emphasis bold="true" box="[151,222,1375,1401]" pageId="10" pageNumber="101">Head.</emphasis>
Ground punctation on labrum, clypeus and frons fine, distance between punctures 2.0–3.0× the width of one puncture. Systematic punctures on labrum consisting of several indistinct punctures, sometimes bearing a short seta. Frons with an irregular row of systematic punctures bearing setae mesad of each eye. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation. Maxillary palps short, about as long as width of mentum; segment 2 slightly bulbous, apical segment approximately the same length as penultimate. Labial palps less than half as long as width of mentum; with clump of long setae on second segment. Mentum quadrate with fine scattered punctures, anterior margin slightly convex and depressed. 
<emphasis bold="true" box="[1341,1436,1591,1617]" pageId="10" pageNumber="101">Thorax.</emphasis>
Ground punctation on pronotum and elytra composed of very fine, evenly distributed punctures. Pronotal systematic punctures with short fine setae, at least 2× the size of surrounding ground punctures, anterior and posterior series each forming an irregular field. Lateral margins of pronotum set with a few sparsely distributed setiferous punctures; with punctures almost appearing absent. Posterolateral corners of pronotum angulate, but not spinose. Elytral margins slightly explante posteriorly. Sutural punctation on elytra absent or unmodified from ground punctation; sutural interval not raised in posterior half. Elytra with five rows of systematic punctures: the first row is represented as a grove with a moderately dense, nearly continuous row of erect setae. Rows 2 and 3 represented by a nearly regular series of coarse punctures bearing slightly shorter and more sparsely set setae than row 1. Punctures in rows 4 and 5 similar to 2 and 3 but irregular in distribution. Prosternum with a clearly defined median carina; slightly elevated anteromedially, the elevation set with two spines. Elevated process of the mesoventrite narrow and elongate, more than three times as long as wide; surface nearly flat; set with 6–7 thickened, coarse spines. Metaventrite with oval glabrous area posteromedially, slightly less than twice as long as wide, length of glabrous area slightly more than half the length of metaventrite. Procoxae covered with dense pubescence that is interspersed with short, thickened spines. Protibiae with 12 spines on dorsal face. 
<emphasis bold="true" box="[690,814,222,248]" pageId="11" pageNumber="102">Abdomen.</emphasis>
Ventrites covered with fine setae, the longest setae not longer than the longest setae surrounding the glabrous area of the metaventrite. Aedeagus as in 
<figureCitation box="[1218,1333,259,285]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="11" pageNumber="102">Figure 6B</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="102" type="distribution">
<paragraph blockId="11.[151,1437,151,429]" pageId="11" pageNumber="102">
<emphasis bold="true" box="[199,353,294,320]" pageId="11" pageNumber="102">Distribution.</emphasis>
Known from a series of localities in the north of 
<collectingCountry box="[891,986,295,321]" name="Ecuador" pageId="11" pageNumber="102">Ecuador</collectingCountry>
, most on the western slopes of the Andes but also one locality on the eastern slope (
<figureCitation box="[621,690,331,357]" captionStart="FIGURE 7" captionStartId="19.[151,250,1886,1910]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@19.[151,1436,181,1868]" captionTargetPageId="19" captionText="FIGURE 7. Distribution of Oocyclus spp. in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.7892078" httpUri="https://zenodo.org/record/7892078/files/figure.png" pageId="11" pageNumber="102">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="102" type="etymology">
<paragraph blockId="11.[151,1437,151,429]" pageId="11" pageNumber="102">
<emphasis bold="true" box="[199,334,367,393]" pageId="11" pageNumber="102">Etymology.</emphasis>
Named after the “sunrise” like coloration of the abdominal ventrites, which are half dark and half orange.
</paragraph>
</subSubSection>
</treatment>
</document>