treatments-xml/data/03/BD/87/03BD8788FFABE65438E8FD71FEDBF88D.xml

<document id="EA7EAADAC76AF1CE881F122BBDB67D67" ID-CLB-Dataset="7330" ID-DOI="10.11646/zootaxa.4952.2.5" ID-GBIF-Dataset="210b95ea-4596-487c-914d-03df0694d33e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4674078" ID-ZooBank="B8276A74-838A-4B54-87C4-8589F81A58F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617948646286" checkinUser="plazi" docAuthor="Schmidt, Ray C., Knobloch, Elise C. &amp; Barrientos, Christian" docDate="2021" docId="03BD8788FFABE65438E8FD71FEDBF88D" docLanguage="en" docName="zootaxa.4952.2.5.pdf" docOrigin="Zootaxa 4952 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Distichodus mbiniensis Schmidt &amp; Knobloch &amp; Barrientos 2021, sp. nov." docType="treatment" docUuid="8B5B9875-DB26-4417-B719-0869D8CFA19E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="308" masterDocId="FF84FFF0FFA5E645387FFFF2FFDFFFBA" masterDocTitle="Cast netting new species: Integrative taxonomy of Distichodus notospilus (Characiformes: Distichodontidae) discovers new species and overlooked areas of endemism in Central Africa" masterLastPageNumber="313" masterPageNumber="291" pageNumber="305" updateTime="1698936259970" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6B6CD4DDACF7A370ABB8ABBB3621A011">Cast netting new species: Integrative taxonomy of Distichodus notospilus (Characiformes: Distichodontidae) discovers new species and overlooked areas of endemism in Central Africa</mods:title>
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<mods:namePart id="236F99106AB7E24A08ACB7D08E34EF96">Schmidt, Ray C.</mods:namePart>
<mods:affiliation id="EE96DFE389FC8EACEB787FF613DE913D">Biology Department, Randolph-Macon College, Ashland, VA 23005, USA; &amp; Smithsonian Research Associate, National Museum of Natural History, Division of Fishes, Washington, DC 20560, USA</mods:affiliation>
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<mods:namePart id="848C48A3B244C7F652A793D64D748969">Knobloch, Elise C.</mods:namePart>
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<mods:namePart id="55DC09CDE81577C2F7816126D160E395">Barrientos, Christian</mods:namePart>
<mods:affiliation id="07D68141C2DF6BABE9B2CDFD652701AE">Wildlife Conservation Society, Equatorial Guinea. Edificio Candy Vista Mar Of. 208 Bata Litoral, Equatorial Guinea</mods:affiliation>
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<taxonomicName id="4C144D1DFFABE64B38E8FD71FE6FFD24" authority="Schmidt &amp; Knobloch &amp; Barrientos, 2021" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[151,432,643,670]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis" status="sp. nov.">
<emphasis id="B960EA8CFFABE64B38E8FD71FE6FFD24" bold="true" box="[151,432,643,670]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
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<paragraph id="8BAB369EFFABE64B38E8FD3EFE32FD5C" blockId="14.[151,961,643,743]" box="[151,493,716,743]" pageId="14" pageNumber="305">
<figureCitation id="132F2A1BFFABE64B38E8FD3EFEDDFD5C" box="[151,258,716,742]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[151,1436,211,1649]" captionTargetId="figure-19@7.[151,1436,211,1649]" captionTargetPageId="7" captionText="FIGURE 2. Phylogeny of Distichodus spp. inferred from 996 bp of cytochrome b (A). Phylogeny inferred from concatenated dataset of 996 bp CYTB, 729 bp MYH6, and 627 bp of CO1(B). Posterior probabilities from Bayesian inference above major branches; above 0.95 not shown and branches with support values &lt;70 were collapsed. Metadata and accession numbers for specimens included in the analyses shown in table 1 and average uncorrected p-distances among these different taxa shown in table 3." figureDoi="http://doi.org/10.5281/zenodo.4674082" httpUri="https://zenodo.org/record/4674082/files/figure.png" pageId="14" pageNumber="305">Figures 2</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B3971FD3EFEC2FD5C" box="[270,285,716,742]" captionStart="FIGURE 3" captionStartId="8.[151,250,1883,1907]" captionTargetBox="[395,1191,181,1859]" captionTargetId="figure-28@8.[395,1191,181,1859]" captionTargetPageId="8" captionText="FIGURE 3. Phylogeny of Distichodus spp. inferred from 627 bp of CO1. Posterior probabilities from Bayesian inference above major branches; above 0.95 not shown and branches with support values &lt;70 were collapsed. Metadata and accession numbers for specimens included in the analyses shown in table 1 and average uncorrected p-distances among these different taxa shown in table 3." figureDoi="http://doi.org/10.5281/zenodo.4674084" httpUri="https://zenodo.org/record/4674084/files/figure.png" pageId="14" pageNumber="305">3</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B3955FD3EFEE5FD5C" box="[298,314,716,742]" captionStart="FIGURE 4" captionStartId="10.[151,250,844,868]" captionTargetBox="[151,1436,237,821]" captionTargetId="figure-52@10.[151,1436,237,821]" captionTargetPageId="10" captionText="FIGURE 4. Plot of PC2 and PC3 from principal component analysis of 24 log-transformed measurements from 51 specimens (A). Reduced-major axis regression of eye diameter as a percentage of snout length on log-transformed standard length (B). Trendlines are shown for each species; slopes for each species are not different (p-value = 0.7613), and the elevation among species are significantly different (p-value = 0.0002) with Distichodus microps being significantly different (p-value &lt;0.01) from D. mbiniensis and topotypic D. notospilus. The holotype of Distichodus microps and D. mbiniensis are denoted by a filled triangle and star respectively. Component loadings and results of Mann-Whitney U tests shown in table 3." figureDoi="http://doi.org/10.5281/zenodo.4674086" httpUri="https://zenodo.org/record/4674086/files/figure.png" pageId="14" pageNumber="305">4</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B3939FD3FFE89FD5D" box="[326,342,717,743]" captionStart="FIGURE 7" captionStartId="16.[151,250,970,994]" captionTargetBox="[171,1414,441,919]" captionTargetId="figure-155@16.[151,1437,417,947]" captionTargetPageId="16" captionText="FIGURE 7. Distichodus mbiniensis, a new species, holotype, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674092" httpUri="https://zenodo.org/record/4674092/files/figure.png" pageId="14" pageNumber="305">7</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B391DFD3EFEAEFD5C" box="[354,369,716,742]" captionStart="FIGURE 8" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[166,1436,1106,1569]" captionTargetId="figure-195@16.[151,1436,1091,1572]" captionTargetPageId="16" captionText="FIGURE 8. Lateral views of head region in Distichodus microps sp. nov. (A), Distichodus mbiniensis sp. nov. (B), and topotypic Distichodus notospilus (C). Same specimens and locality information as previous figures. Photographs by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674094" httpUri="https://zenodo.org/record/4674094/files/figure.png" pageId="14" pageNumber="305">8</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B3901FD3EFE51FD5C" box="[382,398,716,742]" captionStart="FIGURE 9" captionStartId="17.[151,250,850,874]" captionTargetBox="[151,1437,273,827]" captionTargetId="figure-62@17.[151,1437,273,827]" captionTargetPageId="17" captionText="FIGURE 9. Live coloration of Distichodus microps, USNM 451457, ALC 81.6 mm SL, voucher EqGui2017_0274, Equatorial Guinea, Litoral, Rio Mbia at Menzong, 32 m elev, 2.07529° N, 9.92378° E photograph by R.C. Schmidt (A); Distichodus mbiniensis, USNM 451319, ALC 61.3 mm SL, voucher EqGui2017_0130, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m, 1.29091° N, 10.9174° E, photograph by R. C. Schmidt (B); Topotypic Distichodus notospilus, OS 19544, ALC, 104.1 mm SL, Gabon, Province de l’Ogooué-Lolo, Right bank of Ogooué River at Doumé village, 0.84245° S, 12.96249° E, Photograph by B. Sidlauskas, included with permission (C); Holotype of Distichodus mbiniensis, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by R. C. Schmidt (D)." figureDoi="http://doi.org/10.5281/zenodo.4674096" httpUri="https://zenodo.org/record/4674096/files/figure.png" pageId="14" pageNumber="305">9</figureCitation>
; 
<tableCitation id="C6960325FFABE64B39E5FD3EFE32FD5C" box="[410,493,716,742]" captionStart="TABLE 6" captionStartId="15.[151,239,151,175]" captionText="TABLE 6. Morphometric measurements and meristics of Distichodus mbiniensis sp. nov. (n=13; holotype and 12 paratypes). Standard length expressed in mm. All other measurements expressed in percent SL unless noted. Meristic data from holotype are identified by an “*”." pageId="14" pageNumber="305">Table 6</tableCitation>
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<typeStatus id="54AF883CFFABE64B38E8FCE6FED9FC94" box="[151,262,788,814]" pageId="14" pageNumber="305" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBB29EE5FFABE64B396EFCE6FE1FFC95" box="[273,448,788,815]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451317</specimenCode>
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SL, voucher 
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2017_0127, 
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at 
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elev, 
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, 2017 
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expedition team, 
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<typeStatus id="54AF883CFFABE64B38B8FC72FE9EFC20" box="[199,321,896,922]" pageId="14" pageNumber="305" type="paratype">Paratypes</typeStatus>
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<specimenCode id="DBB29EE5FFABE64B3932FC72FDCBFC21" box="[333,532,896,923]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="305" type="Museum">MRAC 2020,005</specimenCode>
,P,0001, 1 ALC, 
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SL, voucher 
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2017_0128, collection information the same as the 
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.— 
<materialsCitation id="3B7C3CC3FFABE64B39ADFC56FD99FC58" ID-GBIF-Occurrence="3073877307" collectionCode="MRAC" collectorName="EqGui" pageId="14" pageNumber="305" specimenCode="MRAC 2020,005" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB29EE5FFABE64B39ADFC56FD43FC05" box="[466,668,932,959]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="305" type="Museum">MRAC 2020,005</specimenCode>
,P,0002, 1 ALC, 
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SL, voucher 
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2017_0135, collection information the same as the 
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</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B3A19FC3AFE67FBBC" ID-GBIF-Occurrence="3073877305" collectionCode="MRAC" pageId="14" pageNumber="305" specimenCode="MRAC 2020,005" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB29EE5FFABE64B3A19FC3AFCEEFC59" box="[614,817,968,995]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="305" type="Museum">MRAC 2020,005</specimenCode>
,P,0003, 1 ALC, 
<quantity id="4CEC9B7BFFABE64B3B8FFC3AFBBBFC59" box="[1008,1124,968,995]" metricMagnitude="-1" metricUnit="m" metricValue="1.139" pageId="14" pageNumber="305" unit="mm" value="113.9">113.9 mm</quantity>
SL, collection information the same as the 
<typeStatus id="54AF883CFFABE64B392AFC1EFE67FBBC" box="[341,440,1004,1030]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B39A6FC1EFE3DFB90" ID-GBIF-Occurrence="3073877309" collectionCode="USNM" collectorName="EqGui" pageId="14" pageNumber="305" specimenCode="USNM 451319" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB29EE5FFABE64B39A6FC1EFD53FBBD" box="[473,652,1004,1031]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451319</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B3A9DFC1EFC96FBBD" box="[738,841,1004,1031]" metricMagnitude="-2" metricUnit="m" metricValue="6.13" pageId="14" pageNumber="305" unit="mm" value="61.3">61.3 mm</quantity>
SL, voucher 
<collectorName id="26E15348FFABE64B3B96FC1EFBEBFBBC" box="[1001,1076,1004,1030]" pageId="14" pageNumber="305">EqGui</collectorName>
2017_0130, collection information the same as the 
<typeStatus id="54AF883CFFABE64B3900FBE2FE3DFB90" box="[383,482,1040,1066]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B3A7CFBE2FDD5FBF4" ID-GBIF-Occurrence="3073877303" collectionCode="USNM" collectorName="EqGui" pageId="14" pageNumber="305" specimenCode="USNM 451318" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB29EE5FFABE64B3A7CFBE2FD6BFB91" box="[515,692,1040,1067]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451318</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B3B7AFBE2FCA7FB91" box="[773,888,1040,1067]" metricMagnitude="-1" metricUnit="m" metricValue="1.2209999999999999" pageId="14" pageNumber="305" unit="mm" value="122.1">122.1 mm</quantity>
SL, voucher 
<collectorName id="26E15348FFABE64B3C6EFBE2FB83FB90" box="[1041,1116,1040,1066]" pageId="14" pageNumber="305">EqGui</collectorName>
2017_0129, collection information the same as the 
<typeStatus id="54AF883CFFABE64B39D8FBC6FDD5FBF4" box="[423,522,1076,1102]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B3A54FBC6FE9BFBC8" ID-GBIF-Occurrence="3073877313" collectionCode="USNM" pageId="14" pageNumber="305" specimenCode="USNM 451320, 3" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB29EE5FFABE64B3A54FBC6FD02FBF5" box="[555,733,1076,1103]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451320</specimenCode>
, 
<specimenCode id="DBB29EE5FFABE64B3A95FBC6FD27FBF4" box="[746,760,1076,1102]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">3</specimenCode>
ALC, 78.4–119.0 mm SL, collection information the same as the 
<typeStatus id="54AF883CFFABE64B389EFBAAFE9BFBC8" box="[225,324,1112,1138]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B391AFBAAFF10FB00" ID-GBIF-Occurrence="3073877311" collectingDate="2017-06-27" collectionCode="USNM" country="Equatorial Guinea" county="EqGui" elevation="660" latitude="1.64875" location="Alam" longLatPrecision="1" longitude="11.08593" municipality="Rio Nuik" pageId="14" pageNumber="305" specimenCode="USNM 451324" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B391AFBAAFDC9FBC9" box="[357,534,1112,1139]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451324</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B3A16FBAAFD11FBC9" box="[617,718,1112,1139]" metricMagnitude="-2" metricUnit="m" metricValue="7.37" pageId="14" pageNumber="305" unit="mm" value="73.7">73.7 mm</quantity>
SL, voucher 
<collectingCounty id="62CA4E12FFABE64B3B17FBAAFC6CFBC8" box="[872,947,1112,1138]" pageId="14" pageNumber="305">EqGui</collectingCounty>
2017_0191, 
<collectingCountry id="F303760EFFABE64B3C41FBAAFAD0FBC8" box="[1086,1295,1112,1138]" name="Equatorial Guinea" pageId="14" pageNumber="305">Equatorial Guinea</collectingCountry>
, 
<collectingRegion id="49D0F87CFFABE64B3D64FBAAFA47FBC8" box="[1307,1432,1112,1138]" country="Equatorial Guinea" name="Wele-Nzas" pageId="14" pageNumber="305">Wele-Nzas</collectingRegion>
, 
<collectingMunicipality id="6BCFACE4FFABE64B38E8FB8EFEDDFB2C" box="[151,258,1148,1174]" pageId="14" pageNumber="305">Rio Nuik</collectingMunicipality>
near 
<location id="8ECB6045FFABE64B393AFB8EFE59FB2C" LSID="urn:lsid:plazi:treatment:03BD8788FFABE65438E8FD71FEDBF88D:8ECB6045FFABE64B393AFB8EFE59FB2C" box="[325,390,1148,1174]" country="Equatorial Guinea" county="EqGui" latitude="1.64875" longLatPrecision="1" longitude="11.08593" municipality="Rio Nuik" name="Alam" pageId="14" pageNumber="305" stateProvince="Wele-Nzas">Alam</location>
, 
<quantity id="4CEC9B7BFFABE64B39EAFB8EFE00FB2D" box="[405,479,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="14" pageNumber="305" unit="m" value="660.0">
<elevation id="0039D1ADFFABE64B39EAFB8EFE00FB2D" box="[405,479,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="14" pageNumber="305" unit="m" value="660.0">660 m</elevation>
</quantity>
elev, 
<geoCoordinate id="EE205059FFABE64B3A57FB8EFD70FB2D" box="[552,687,1148,1175]" degrees="1.64875" direction="north" orientation="latitude" pageId="14" pageNumber="305" precision="1" value="1.64875">1.64875° N</geoCoordinate>
, 
<geoCoordinate id="EE205059FFABE64B3AC1FB8EFC91FB2D" box="[702,846,1148,1175]" degrees="11.08593" direction="east" orientation="longitude" pageId="14" pageNumber="305" precision="1" value="11.08593">11.08593° E</geoCoordinate>
, 2017 
<collectingCountry id="F303760EFFABE64B3BDFFB8EFBAFFB2C" box="[928,1136,1148,1174]" name="Equatorial Guinea" pageId="14" pageNumber="305">Equatorial Guinea</collectingCountry>
expedition team, 
<date id="FFAA105EFFABE64B3D3CFB8EFF10FB00" pageId="14" pageNumber="305" value="2017-06-27">
<collectingDate id="EFEEE9B6FFABE64B3D3CFB8EFF10FB00" pageId="14" pageNumber="305" value="2017-06-27">27 June 2017</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B388FFB52FED4FAB8" ID-GBIF-Occurrence="3073877302" collectingDate="2017-06-23" collectionCode="USNM" country="Equatorial Guinea" county="EqGui" elevation="664" latitude="1.31069" location="Evinayong - Aconibe Rd" longLatPrecision="1" longitude="10.86025" municipality="Rio Abia" pageId="14" pageNumber="305" specimenCode="USNM 451321" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B388FFB52FE7DFB01" box="[240,418,1184,1211]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451321</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B398DFB52FDBAFB01" box="[498,613,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="1.0859999999999999" pageId="14" pageNumber="305" unit="mm" value="108.6">108.6 mm</quantity>
SL, voucher 
<collectingCounty id="62CA4E12FFABE64B3A81FB52FC96FB00" box="[766,841,1184,1210]" pageId="14" pageNumber="305">EqGui</collectingCounty>
2017_0147, 
<collectingCountry id="F303760EFFABE64B3BACFB52FB7BFB00" box="[979,1188,1184,1210]" name="Equatorial Guinea" pageId="14" pageNumber="305">Equatorial Guinea</collectingCountry>
, 
<collectingRegion id="49D0F87CFFABE64B3CD0FB52FAF3FB00" box="[1199,1324,1184,1210]" country="Equatorial Guinea" name="Wele-Nzas" pageId="14" pageNumber="305">Wele-Nzas</collectingRegion>
, 
<collectingMunicipality id="6BCFACE4FFABE64B3D48FB52FA43FB00" box="[1335,1436,1184,1210]" pageId="14" pageNumber="305">Rio Abia</collectingMunicipality>
at 
<location id="8ECB6045FFABE64B38CBFB36FE0AFB64" LSID="urn:lsid:plazi:treatment:03BD8788FFABE65438E8FD71FEDBF88D:8ECB6045FFABE64B38CBFB36FE0AFB64" box="[180,469,1220,1246]" country="Equatorial Guinea" county="EqGui" latitude="1.31069" longLatPrecision="1" longitude="10.86025" municipality="Rio Abia" name="Evinayong - Aconibe Rd" pageId="14" pageNumber="305" stateProvince="Wele-Nzas">Evinayong—Aconibe Rd</location>
, 
<quantity id="4CEC9B7BFFABE64B399DFB36FDF4FB65" box="[482,555,1220,1247]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="14" pageNumber="305" unit="m" value="664.0">
<elevation id="0039D1ADFFABE64B399DFB36FDF4FB65" box="[482,555,1220,1247]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="14" pageNumber="305" unit="m" value="664.0">664 m</elevation>
</quantity>
elev, 
<geoCoordinate id="EE205059FFABE64B3A0EFB36FD28FB65" box="[625,759,1220,1247]" degrees="1.31069" direction="north" orientation="latitude" pageId="14" pageNumber="305" precision="1" value="1.31069">1.31069° N</geoCoordinate>
, 
<geoCoordinate id="EE205059FFABE64B3B7BFB36FC4BFB65" box="[772,916,1220,1247]" degrees="10.86025" direction="east" orientation="longitude" pageId="14" pageNumber="305" precision="1" value="10.86025">10.86025° E</geoCoordinate>
, 2017 
<collectingCountry id="F303760EFFABE64B3B9CFB36FB6DFB64" box="[995,1202,1220,1246]" name="Equatorial Guinea" pageId="14" pageNumber="305">Equatorial Guinea</collectingCountry>
expedition team, 
<date id="FFAA105EFFABE64B3DFFFB36FED4FAB8" pageId="14" pageNumber="305" value="2017-06-23">
<collectingDate id="EFEEE9B6FFABE64B3DFFFB36FED4FAB8" pageId="14" pageNumber="305" value="2017-06-23">23 June 2017</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B3953FB1AFA43FAB9" ID-GBIF-Occurrence="3073877304" box="[300,1436,1256,1283]" collectionCode="USNM" country="Equatorial Guinea" pageId="14" pageNumber="305" specimenCode="USNM 451322" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B3953FB1AFE01FAB9" box="[300,478,1256,1283]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451322</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B3A4EFB1AFD7CFAB9" box="[561,675,1256,1283]" metricMagnitude="-1" metricUnit="m" metricValue="1.138" pageId="14" pageNumber="305" unit="mm" value="113.8">113.8 mm</quantity>
SL, voucher EqGui2017_0149, collection information the same as
</materialsCitation>
<materialsCitation id="3B7C3CC3FFABE64B38E8FAFEFE9CFA9D" ID-GBIF-Occurrence="3073877308" box="[151,323,1292,1319]" collectionCode="USNM" country="Equatorial Guinea" pageId="14" pageNumber="305" specimenCode="USNM 451321" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B38E8FAFEFE9CFA9D" box="[151,323,1292,1319]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451321</specimenCode>
</materialsCitation>
.— 
<materialsCitation id="3B7C3CC3FFABE64B391BFAFEFBA0FA9D" ID-GBIF-Occurrence="3073877310" box="[356,1151,1292,1319]" collectionCode="USNM" country="Equatorial Guinea" pageId="14" pageNumber="305" specimenCode="USNM 451323" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B391BFAFEFDCAFA9D" box="[356,533,1292,1319]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451323</specimenCode>
, ALC, 
<quantity id="4CEC9B7BFFABE64B3A1BFAFEFD17FA9D" box="[612,712,1292,1319]" metricMagnitude="-2" metricUnit="m" metricValue="9.03" pageId="14" pageNumber="305" unit="mm" value="90.3">90.3 mm</quantity>
SL, collection information the same as
</materialsCitation>
<materialsCitation id="3B7C3CC3FFABE64B3CF9FAFEFAE9FA9D" ID-GBIF-Occurrence="3073877316" box="[1158,1334,1292,1319]" collectionCode="USNM" country="Equatorial Guinea" pageId="14" pageNumber="305" specimenCode="USNM 451321" specimenCount="1" stateProvince="Wele-Nzas" typeStatus="paratype">
<specimenCode id="DBB29EE5FFABE64B3CF9FAFEFAE9FA9D" box="[1158,1334,1292,1319]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="305" type="Museum">USNM 451321</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6515FFABE64B38B8FAC2FBA8F84C" pageId="14" pageNumber="305" type="diagnosis">
<paragraph id="8BAB369EFFABE64B38B8FAC2FBA8F84C" blockId="14.[151,1437,788,2039]" pageId="14" pageNumber="305">
<emphasis id="B960EA8CFFABE64B38B8FAC2FE9DFAF0" bold="true" box="[199,322,1328,1354]" pageId="14" pageNumber="305">Diagnosis.</emphasis>
<taxonomicName id="4C144D1DFFABE64B3938FAC2FD94FAF0" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[327,587,1328,1354]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B3938FAC2FD94FAF0" box="[327,587,1328,1354]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
is readily distinguished from the larger-bodied, higher-scale count 
<taxonomicName id="4C144D1DFFABE64B3D4BFAC2FF61FAD4" authorityName="J.P.Muller &amp; Troschel" authorityYear="1844" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="genus">
<emphasis id="B960EA8CFFABE64B3D4BFAC2FF61FAD4" italics="true" pageId="14" pageNumber="305">Distichodus</emphasis>
</taxonomicName>
spp. (group B in 
<bibRefCitation id="EF854B6FFFABE64B3900FAA6FDB5FAD4" author="Arroyave, J. &amp; Denton, J. S. &amp; Stiassny, M. L." box="[383,618,1364,1390]" pageId="14" pageNumber="305" pagination="1 - 28" refId="ref12268" refString="Arroyave, J., Denton, J. S. &amp; Stiassny, M. L. (2020) Pattern and timing of diversification in the African freshwater fish genus Distichodus (Characiformes: Distichodontidae). BMC Evolutionary Biology, 20, 1 - 28. https: // doi. org / 10.1186 / s 12862 - 020 - 01615 - 6" type="journal article" year="2020">
Arroyave 
<emphasis id="B960EA8CFFABE64B398FFAA7FDF7FAD4" box="[496,552,1364,1390]" italics="true" pageId="14" pageNumber="305">et al.</emphasis>
2020
</bibRefCitation>
; 
<taxonomicName id="4C144D1DFFABE64B3A0AFAA7FD3AFAD4" authorityName="Schilthuis" authorityYear="1891" box="[629,741,1365,1390]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="antonii">
<emphasis id="B960EA8CFFABE64B3A0AFAA7FD3AFAD4" box="[629,741,1365,1390]" italics="true" pageId="14" pageNumber="305">D. antonii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3A8BFAA7FC7AFAD4" authorityName="Boulenger" authorityYear="1898" box="[756,933,1364,1390]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="atroventralis">
<emphasis id="B960EA8CFFABE64B3A8BFAA7FC7AFAD4" box="[756,933,1364,1390]" italics="true" pageId="14" pageNumber="305">D. atroventralis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3BCCFAA7FB91FAD4" authorityName="Boulenger" authorityYear="1898" box="[947,1102,1364,1390]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="B960EA8CFFABE64B3BCCFAA7FB91FAD4" box="[947,1102,1364,1390]" italics="true" pageId="14" pageNumber="305">D. fasciolatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3C23FAA7FB0FFAD4" authorityName="Schilthuis" authorityYear="1891" box="[1116,1232,1364,1390]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="lusosso">
<emphasis id="B960EA8CFFABE64B3C23FAA7FB0FFAD4" box="[1116,1232,1364,1390]" italics="true" pageId="14" pageNumber="305">D. lusosso</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3CA1FAA7FA4AFAD4" authorityName="Peters" authorityYear="1852" box="[1246,1429,1364,1390]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mossambicus">
<emphasis id="B960EA8CFFABE64B3CA1FAA7FA4AFAD4" box="[1246,1429,1364,1390]" italics="true" pageId="14" pageNumber="305">D. mossambicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B38E8FA8BFE9EFA28" authorityName="Boulenger" authorityYear="1897" box="[151,321,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="sexfasciatus">
<emphasis id="B960EA8CFFABE64B38E8FA8BFE9EFA28" box="[151,321,1400,1426]" italics="true" pageId="14" pageNumber="305">D. sexfasciatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B392FFA8BFE71FA28" authorityName="Nichols &amp; Griscom" authorityYear="1917" box="[336,430,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="langi">
<emphasis id="B960EA8CFFABE64B392FFA8BFE71FA28" box="[336,430,1400,1426]" italics="true" pageId="14" pageNumber="305">D. langi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B39C3FA8BFD9CFA28" box="[444,579,1401,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="rostratus">
<emphasis id="B960EA8CFFABE64B39C3FA8BFD9CFA28" box="[444,579,1401,1426]" italics="true" pageId="14" pageNumber="305">D. rostratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3A2DFA8BFCD2FA28" box="[594,781,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="engycephalus">
<emphasis id="B960EA8CFFABE64B3A2DFA8BFCD2FA28" box="[594,781,1400,1426]" italics="true" pageId="14" pageNumber="305">D. engycephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3B63FA8BFC6DFA28" box="[796,946,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kasaiensis">
<emphasis id="B960EA8CFFABE64B3B63FA8BFC6DFA28" box="[796,946,1400,1426]" italics="true" pageId="14" pageNumber="305">D. kasaiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3BBEFA8BFBFDFA28" box="[961,1058,1401,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="ingae">
<emphasis id="B960EA8CFFABE64B3BBEFA8BFBFDFA28" box="[961,1058,1401,1426]" italics="true" pageId="14" pageNumber="305">D. ingae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3C4EFA8BFB57FA28" box="[1073,1160,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="polli">
<emphasis id="B960EA8CFFABE64B3C4EFA8BFB57FA28" box="[1073,1160,1400,1426]" italics="true" pageId="14" pageNumber="305">D. polli</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3CE8FA8BFAD1FA28" authorityName="Pfeffer" authorityYear="1896" box="[1175,1294,1401,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="petersii">
<emphasis id="B960EA8CFFABE64B3CE8FA8BFAD1FA28" box="[1175,1294,1401,1426]" italics="true" pageId="14" pageNumber="305">D. petersii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3D62FA8BFA49FA28" box="[1309,1430,1400,1426]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="nefasch">
<emphasis id="B960EA8CFFABE64B3D62FA8BFA49FA28" box="[1309,1430,1400,1426]" italics="true" pageId="14" pageNumber="305">D. nefasch</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B38E8FA6FFEE7FA0C" authorityName="Gunther" authorityYear="1864" box="[151,312,1436,1462]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="brevipinnis">
<emphasis id="B960EA8CFFABE64B38E8FA6FFEE7FA0C" box="[151,312,1436,1462]" italics="true" pageId="14" pageNumber="305">D. brevipinnis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C144D1DFFABE64B3908FA6FFE28FA0C" authorityName="Peters" authorityYear="1852" box="[375,503,1436,1462]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="schenga">
<emphasis id="B960EA8CFFABE64B3908FA6FFE28FA0C" box="[375,503,1436,1462]" italics="true" pageId="14" pageNumber="305">D. schenga</emphasis>
</taxonomicName>
) in having fewer lateral line scales (40–42 versus&gt;60) and in achieving a smaller maximum standard length (&lt;
<quantity id="4CEC9B7BFFABE64B39A5FA32FDC1FA61" box="[474,542,1472,1499]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="305" unit="cm" value="15.0">15 cm</quantity>
versus&gt; 
<quantity id="4CEC9B7BFFABE64B3AFDFA32FD14FA61" box="[642,715,1472,1499]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="305" unit="cm" value="30.0">30 cm</quantity>
). This new species is also distinguished from the smaller-bodied, lower-scale count 
<taxonomicName id="4C144D1DFFABE64B391DFA16FE37FA44" authorityName="J.P.Muller &amp; Troschel" authorityYear="1844" box="[354,488,1508,1534]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="genus">
<emphasis id="B960EA8CFFABE64B391DFA16FE37FA44" box="[354,488,1508,1534]" italics="true" pageId="14" pageNumber="305">Distichodus</emphasis>
</taxonomicName>
spp. (group A in 
<bibRefCitation id="EF854B6FFFABE64B3ADAFA16FC52FA44" author="Arroyave, J. &amp; Denton, J. S. &amp; Stiassny, M. L." box="[677,909,1508,1534]" pageId="14" pageNumber="305" pagination="1 - 28" refId="ref12268" refString="Arroyave, J., Denton, J. S. &amp; Stiassny, M. L. (2020) Pattern and timing of diversification in the African freshwater fish genus Distichodus (Characiformes: Distichodontidae). BMC Evolutionary Biology, 20, 1 - 28. https: // doi. org / 10.1186 / s 12862 - 020 - 01615 - 6" type="journal article" year="2020">
Arroyave 
<emphasis id="B960EA8CFFABE64B3B6AFA17FC92FA44" box="[789,845,1508,1534]" italics="true" pageId="14" pageNumber="305">et al.</emphasis>
2020
</bibRefCitation>
) in the following ways. 
<taxonomicName id="4C144D1DFFABE64B3CE7FA16FA43FA44" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[1176,1436,1508,1534]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B3CE7FA16FA43FA44" box="[1176,1436,1508,1534]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C144D1DFFABE64B39F4F9FBFDB6F998" authorityName="Pellegrin" authorityYear="1926" box="[395,617,1544,1570]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="decemmaculatus">
<emphasis id="B960EA8CFFABE64B39F4F9FBFDB6F998" box="[395,617,1544,1570]" italics="true" pageId="14" pageNumber="305">D. decemmaculatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C144D1DFFABE64B3AE0F9FBFCC7F998" authorityName="Mamonekene &amp; Vreven" authorityYear="2008" box="[671,792,1544,1570]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="teugelsi">
<emphasis id="B960EA8CFFABE64B3AE0F9FBFCC7F998" box="[671,792,1544,1570]" italics="true" pageId="14" pageNumber="305">D. teugelsi</emphasis>
</taxonomicName>
in having two rows of teeth on the lower jaw versus one. 
<taxonomicName id="4C144D1DFFABE64B38E8F9DEFE43F9FC" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[151,412,1580,1606]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B38E8F9DEFE43F9FC" box="[151,412,1580,1606]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
can be readily distinguished from 
<emphasis id="B960EA8CFFABE64B3B5CF9DFFB87F9FC" box="[803,1112,1580,1606]" italics="true" pageId="14" pageNumber="305">
<taxonomicName id="4C144D1DFFABE64B3B5CF9DFFC50F9FC" authorityName="Boulenger" authorityYear="1899" box="[803,911,1580,1606]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="noboli">D. noboli</taxonomicName>
, 
<taxonomicName id="4C144D1DFFABE64B3BE4F9DFFB87F9FC" authorityName="Pellegrin" authorityYear="1900" box="[923,1112,1580,1606]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="hypostomatus">D. hypostomatus</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C144D1DFFABE64B3CE9F9DFFAF4F9FC" authorityName="Boulenger" authorityYear="1898" box="[1174,1323,1580,1606]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B960EA8CFFABE64B3CE9F9DFFAF4F9FC" box="[1174,1323,1580,1606]" italics="true" pageId="14" pageNumber="305">D. maculatus</emphasis>
</taxonomicName>
in having fewer scales along the lateral line (40–42 versus 45, 53–60, and 75 respectively), and distinguished from 
<taxonomicName id="4C144D1DFFABE64B3D38F9A3FF3AF934" authorityName="Norman" authorityYear="1922" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="rufigiensis">
<emphasis id="B960EA8CFFABE64B3D38F9A3FF3AF934" italics="true" pageId="14" pageNumber="305">D. rufigiensis</emphasis>
</taxonomicName>
in not having prominent vertical bars along the sides. 
<taxonomicName id="4C144D1DFFABE64B3B46F986FBE2F934" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[825,1085,1652,1678]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B3B46F986FBE2F934" box="[825,1085,1652,1678]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C144D1DFFABE64B3D4DF987FA43F934" authorityName="Holly" authorityYear="1926" box="[1330,1436,1652,1678]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kolleri">
<emphasis id="B960EA8CFFABE64B3D4DF987FA43F934" box="[1330,1436,1652,1678]" italics="true" pageId="14" pageNumber="305">D. kolleri</emphasis>
</taxonomicName>
in having fewer dorsal-fin rays (16–18 total rays versus 21–26), and distinguished from 
<taxonomicName id="4C144D1DFFABE64B3CFDF96BFB04F908" authorityName="Boulenger" authorityYear="1899" box="[1154,1243,1688,1714]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="altus">
<emphasis id="B960EA8CFFABE64B3CFDF96BFB04F908" box="[1154,1243,1688,1714]" italics="true" pageId="14" pageNumber="305">D. altus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C144D1DFFABE64B3D6AF96BFAA1F908" box="[1301,1406,1688,1714]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="affinis">
<emphasis id="B960EA8CFFABE64B3D6AF96BFAA1F908" box="[1301,1406,1688,1714]" italics="true" pageId="14" pageNumber="305">D. affinis</emphasis>
</taxonomicName>
in having fewer total anal-fin rays (13–14 versus 21–22). 
<taxonomicName id="4C144D1DFFABE64B3B7BF94EFBD5F96C" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[772,1034,1724,1750]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B3B7BF94EFBD5F96C" box="[772,1034,1724,1750]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C144D1DFFABE64B3D79F94FFA43F96C" box="[1286,1436,1724,1750]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="notospilus">
<emphasis id="B960EA8CFFABE64B3D79F94FFA43F96C" box="[1286,1436,1724,1750]" italics="true" pageId="14" pageNumber="305">D. notospilus</emphasis>
</taxonomicName>
in having more scales along the lateral line (41–42, rarely 40 versus 37–39, rarely 40; 
<tableCitation id="C6960325FFABE64B3C13F912FB14F941" box="[1132,1227,1760,1787]" captionStart="TABLE 5" captionStartId="12.[151,239,151,175]" captionText="TABLE 5. Morphometric measurements and meristics of topotypic Distichodus notospilus (n=26). Standard length expressed in mm. All other measurements expressed in percent SL unless noted." pageId="14" pageNumber="305">Tables 5</tableCitation>
&amp; 
<tableCitation id="C6960325FFABE64B3C8EF912FADEF941" box="[1265,1281,1760,1787]" captionStart="TABLE 6" captionStartId="15.[151,239,151,175]" captionText="TABLE 6. Morphometric measurements and meristics of Distichodus mbiniensis sp. nov. (n=13; holotype and 12 paratypes). Standard length expressed in mm. All other measurements expressed in percent SL unless noted. Meristic data from holotype are identified by an “*”." pageId="14" pageNumber="305">6</tableCitation>
), a much less prominent elongated dark spot at the base of the caudal fin, a shorter dorsal fin (21.4–27.2 versus 22.7–34.2% SL), and shallower caudal peduncle (11.8–13.0 versus 12.3–14.1% SL). 
<taxonomicName id="4C144D1DFFABE64B3BFAF8DAFB56F8F8" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[901,1161,1832,1858]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFABE64B3BFAF8DAFB56F8F8" box="[901,1161,1832,1858]" italics="true" pageId="14" pageNumber="305">Distichodus mbiniensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C144D1DFFABE64B3DFEF8DBFF2FF8DC" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="microps">
<emphasis id="B960EA8CFFABE64B3DFEF8DBFF2FF8DC" italics="true" pageId="14" pageNumber="305">D. microps</emphasis>
</taxonomicName>
in having a deeper body (usually seven scales from LL to the pelvic fin versus six), a subterminal mouth versus the nearly inferior mouth of 
<taxonomicName id="4C144D1DFFABE64B3A56F883FD7AF830" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[553,677,1905,1930]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="microps">
<emphasis id="B960EA8CFFABE64B3A56F883FD7AF830" box="[553,677,1905,1930]" italics="true" pageId="14" pageNumber="305">D. microps</emphasis>
</taxonomicName>
, a shallower (11.8–13.0 versus 12.4–14.2% SL) and shorter caudal peduncle (9.7–12.6 versus 11.9–14.8% SL), a larger eye (58.6–88.0 versus 56.7–80.4% of snout length, 
<figureCitation id="132F2A1BFFABE64B3D63F866FAA3F814" box="[1308,1404,1940,1966]" captionStart="FIGURE 4" captionStartId="10.[151,250,844,868]" captionTargetBox="[151,1436,237,821]" captionTargetId="figure-52@10.[151,1436,237,821]" captionTargetPageId="10" captionText="FIGURE 4. Plot of PC2 and PC3 from principal component analysis of 24 log-transformed measurements from 51 specimens (A). Reduced-major axis regression of eye diameter as a percentage of snout length on log-transformed standard length (B). Trendlines are shown for each species; slopes for each species are not different (p-value = 0.7613), and the elevation among species are significantly different (p-value = 0.0002) with Distichodus microps being significantly different (p-value &lt;0.01) from D. mbiniensis and topotypic D. notospilus. The holotype of Distichodus microps and D. mbiniensis are denoted by a filled triangle and star respectively. Component loadings and results of Mann-Whitney U tests shown in table 3." figureDoi="http://doi.org/10.5281/zenodo.4674086" httpUri="https://zenodo.org/record/4674086/files/figure.png" pageId="14" pageNumber="305">Figs. 4B</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B3DF8F866FA48F815" box="[1415,1431,1940,1967]" captionStart="FIGURE 5" captionStartId="13.[151,250,943,967]" captionTargetBox="[175,1421,400,902]" captionTargetId="figure-120@13.[151,1436,381,920]" captionTargetPageId="13" captionText="FIGURE 5. Distichodus microps, a new species, holotype, USNM 451325, ALC, 98.0 mm SL, Equatorial Guinea, Litoral, Rio Mbia at Menzong, 32 m elev, 2.07529° N, 9.92378° E; Photograph by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674088" httpUri="https://zenodo.org/record/4674088/files/figure.png" pageId="14" pageNumber="305">5</figureCitation>
, 
<figureCitation id="132F2A1BFFABE64B38E8F84BFF78F869" box="[151,167,1977,2003]" captionStart="FIGURE 7" captionStartId="16.[151,250,970,994]" captionTargetBox="[171,1414,441,919]" captionTargetId="figure-155@16.[151,1437,417,947]" captionTargetPageId="16" captionText="FIGURE 7. Distichodus mbiniensis, a new species, holotype, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674092" httpUri="https://zenodo.org/record/4674092/files/figure.png" pageId="14" pageNumber="305">7</figureCitation>
, &amp; 8), a wider subopercle (
<figureCitation id="132F2A1BFFABE64B39AAF84AFDC7F868" box="[469,536,1976,2002]" captionStart="FIGURE 8" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[166,1436,1106,1569]" captionTargetId="figure-195@16.[151,1436,1091,1572]" captionTargetPageId="16" captionText="FIGURE 8. Lateral views of head region in Distichodus microps sp. nov. (A), Distichodus mbiniensis sp. nov. (B), and topotypic Distichodus notospilus (C). Same specimens and locality information as previous figures. Photographs by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674094" httpUri="https://zenodo.org/record/4674094/files/figure.png" pageId="14" pageNumber="305">Fig. 8</figureCitation>
), and a shallower infraorbital 1 with a more curved anterolateral margin versus a deeper infraorbital 1 with a more rectangular anterolateral margin in 
<taxonomicName id="4C144D1DFFABE64B3BE5F82FFBCAF84C" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[922,1045,2013,2038]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="microps">
<emphasis id="B960EA8CFFABE64B3BE5F82FFBCAF84C" box="[922,1045,2013,2038]" italics="true" pageId="14" pageNumber="305">D. microps</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A1BFFABE64B3C5AF82EFBB6F84C" box="[1061,1129,2012,2038]" captionStart="FIGURE 8" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[166,1436,1106,1569]" captionTargetId="figure-195@16.[151,1436,1091,1572]" captionTargetPageId="16" captionText="FIGURE 8. Lateral views of head region in Distichodus microps sp. nov. (A), Distichodus mbiniensis sp. nov. (B), and topotypic Distichodus notospilus (C). Same specimens and locality information as previous figures. Photographs by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674094" httpUri="https://zenodo.org/record/4674094/files/figure.png" pageId="14" pageNumber="305">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6B6616FFAAE64A38E8FF65FDE9FF42" pageId="15" pageNumber="306" startId="15.[151,239,151,175]">
<paragraph id="8BAB369EFFAAE64A38E8FF65FDE9FF42" blockId="15.[151,1436,151,248]" pageId="15" pageNumber="306">
<emphasis id="B960EA8CFFAAE64A38E8FF65FED5FF15" bold="true" box="[151,266,151,175]" pageId="15" pageNumber="306">TABLE 6.</emphasis>
Morphometric measurements and meristics of 
<taxonomicName id="4C144D1DFFAAE64A3B79FF65FC21FF15" authority="Schmidt &amp; Knobloch &amp; Barrientos, 2021" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[774,1022,151,175]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="15" pageNumber="306" phylum="Chordata" rank="species" species="mbiniensis" status="sp. nov.">
<emphasis id="B960EA8CFFAAE64A3B79FF65FC21FF15" box="[774,1022,151,175]" italics="true" pageId="15" pageNumber="306">Distichodus mbiniensis</emphasis>
</taxonomicName>
<emphasis id="B960EA8CFFAAE64A3C79FF6AFB83FF0A" bold="true" box="[1030,1116,152,176]" pageId="15" pageNumber="306">
<taxonomicNameLabel id="A25357F7FFAAE64A3C79FF6AFB83FF0A" box="[1030,1116,152,176]" pageId="15" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(n=13; holotype and 12 paratypes). Standard length expressed in mm. All other measurements expressed in percent SL unless noted. Meristic data from holotype are identified by an “*”.
</paragraph>
</caption>
<paragraph id="8BAB369EFFAAE64A38D9FEFAFB6FF973" pageId="15" pageNumber="306">
<table id="F914C43EFFAA19BA38D9FEFAFAB7F973" box="[166,1384,264,1737]" colsContinueFrom="11.[166,1342,625,1776]" gridcols="4" gridrows="35" pageId="15" pageNumber="306">
<tr id="352434DCFFAA19BA38D9FEFAFAB7FE9A" box="[166,1384,264,288]" gridrow="0" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FEFAFDDEFE9A" box="[166,513,264,288]" gridcol="0" gridrow="0" pageId="15" pageNumber="306">MORPHOMETRICS</th>
<th id="76F55DA0FFAA19BA3B54FEFAFC56FE9A" box="[811,905,264,288]" gridcol="1" gridrow="0" pageId="15" pageNumber="306">Holotype</th>
<th id="76F55DA0FFAA19BA3BABFEFAFB6DFE9A" box="[980,1202,264,288]" gridcol="2" gridrow="0" pageId="15" pageNumber="306">Range</th>
<th id="76F55DA0FFAA19BA3C94FEFAFAB7FE9A" box="[1259,1384,264,288]" gridcol="3" gridrow="0" pageId="15" pageNumber="306">Mean±%SD</th>
</tr>
<tr id="352434DCFFAA19BA38D9FEC2FAB7FEF2" box="[166,1384,304,328]" gridrow="1" pageId="15" pageNumber="306" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9FEC2FDDEFEF2" box="[166,513,304,328]" gridcol="0" gridrow="1" pageId="15" pageNumber="306">Standard Length (mm)</th>
<td id="76F55DA0FFAA19BA3B54FEC2FC56FEF2" box="[811,905,304,328]" gridcol="1" gridrow="1" pageId="15" pageNumber="306">99.9</td>
<td id="76F55DA0FFAA19BA3BABFEC2FB6DFEF2" box="[980,1202,304,328]" gridcol="2" gridrow="1" pageId="15" pageNumber="306">61.3–122.1</td>
</tr>
<tr id="352434DCFFAA19BA38D9FEAAFAB7FECA" box="[166,1384,344,368]" gridrow="2" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FEAAFDDEFECA" box="[166,513,344,368]" gridcol="0" gridrow="2" pageId="15" pageNumber="306">Predorsal length</th>
<td id="76F55DA0FFAA19BA3B54FEAAFC56FECA" box="[811,905,344,368]" gridcol="1" gridrow="2" pageId="15" pageNumber="306">52.1</td>
<td id="76F55DA0FFAA19BA3BABFEAAFB6DFECA" box="[980,1202,344,368]" gridcol="2" gridrow="2" pageId="15" pageNumber="306">49.5–54.4</td>
<td id="76F55DA0FFAA19BA3C94FEAAFAB7FECA" box="[1259,1384,344,368]" gridcol="3" gridrow="2" pageId="15" pageNumber="306">52.6±1.5</td>
</tr>
<tr id="352434DCFFAA19BA38D9FE72FAB7FE23" box="[166,1384,384,409]" gridrow="3" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FE72FDDEFE23" box="[166,513,384,409]" gridcol="0" gridrow="3" pageId="15" pageNumber="306">Head length</th>
<td id="76F55DA0FFAA19BA3B54FE72FC56FE23" box="[811,905,384,409]" gridcol="1" gridrow="3" pageId="15" pageNumber="306">23.9</td>
<td id="76F55DA0FFAA19BA3BABFE72FB6DFE23" box="[980,1202,384,409]" gridcol="2" gridrow="3" pageId="15" pageNumber="306">21.8–28.9</td>
<td id="76F55DA0FFAA19BA3C94FE72FAB7FE23" box="[1259,1384,384,409]" gridcol="3" gridrow="3" pageId="15" pageNumber="306">25±2.1</td>
</tr>
<tr id="352434DCFFAA19BA38D9FE5BFAB7FE7B" box="[166,1384,425,449]" gridrow="4" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FE5BFDDEFE7B" box="[166,513,425,449]" gridcol="0" gridrow="4" pageId="15" pageNumber="306">Eye diameter</th>
<td id="76F55DA0FFAA19BA3B54FE5BFC56FE7B" box="[811,905,425,449]" gridcol="1" gridrow="4" pageId="15" pageNumber="306">6.3</td>
<td id="76F55DA0FFAA19BA3BABFE5BFB6DFE7B" box="[980,1202,425,449]" gridcol="2" gridrow="4" pageId="15" pageNumber="306">5.7–8.5</td>
<td id="76F55DA0FFAA19BA3C94FE5BFAB7FE7B" box="[1259,1384,425,449]" gridcol="3" gridrow="4" pageId="15" pageNumber="306">6.7±0.9</td>
</tr>
<tr id="352434DCFFAA19BA38D9FE23FAB7FE53" box="[166,1384,465,489]" gridrow="5" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FE23FDDEFE53" box="[166,513,465,489]" gridcol="0" gridrow="5" pageId="15" pageNumber="306">Snout length</th>
<td id="76F55DA0FFAA19BA3B54FE23FC56FE53" box="[811,905,465,489]" gridcol="1" gridrow="5" pageId="15" pageNumber="306">8.5</td>
<td id="76F55DA0FFAA19BA3BABFE23FB6DFE53" box="[980,1202,465,489]" gridcol="2" gridrow="5" pageId="15" pageNumber="306">7.1–10.7</td>
<td id="76F55DA0FFAA19BA3C94FE23FAB7FE53" box="[1259,1384,465,489]" gridcol="3" gridrow="5" pageId="15" pageNumber="306">9.0±1.2</td>
</tr>
<tr id="352434DCFFAA19BA38D9FE0BFAB7FDAB" box="[166,1384,505,529]" gridrow="6" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FE0BFDDEFDAB" box="[166,513,505,529]" gridcol="0" gridrow="6" pageId="15" pageNumber="306">Interorbital distance</th>
<td id="76F55DA0FFAA19BA3B54FE0BFC56FDAB" box="[811,905,505,529]" gridcol="1" gridrow="6" pageId="15" pageNumber="306">9.6</td>
<td id="76F55DA0FFAA19BA3BABFE0BFB6DFDAB" box="[980,1202,505,529]" gridcol="2" gridrow="6" pageId="15" pageNumber="306">8.7–9.6</td>
<td id="76F55DA0FFAA19BA3C94FE0BFAB7FDAB" box="[1259,1384,505,529]" gridcol="3" gridrow="6" pageId="15" pageNumber="306">9.1±0.3</td>
</tr>
<tr id="352434DCFFAA19BA38D9FDD3FAB7FD83" box="[166,1384,545,569]" gridrow="7" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FDD3FDDEFD83" box="[166,513,545,569]" gridcol="0" gridrow="7" pageId="15" pageNumber="306">Internasal distance</th>
<td id="76F55DA0FFAA19BA3B54FDD3FC56FD83" box="[811,905,545,569]" gridcol="1" gridrow="7" pageId="15" pageNumber="306">5.4</td>
<td id="76F55DA0FFAA19BA3BABFDD3FB6DFD83" box="[980,1202,545,569]" gridcol="2" gridrow="7" pageId="15" pageNumber="306">4.8–6.2</td>
<td id="76F55DA0FFAA19BA3C94FDD3FAB7FD83" box="[1259,1384,545,569]" gridcol="3" gridrow="7" pageId="15" pageNumber="306">5.6±0.6</td>
</tr>
<tr id="352434DCFFAA19BA38D9FDB8FAB7FDD8" box="[166,1384,586,610]" gridrow="8" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FDB8FDDEFDD8" box="[166,513,586,610]" gridcol="0" gridrow="8" pageId="15" pageNumber="306">Prepectoral length</th>
<td id="76F55DA0FFAA19BA3B54FDB8FC56FDD8" box="[811,905,586,610]" gridcol="1" gridrow="8" pageId="15" pageNumber="306">26.1</td>
<td id="76F55DA0FFAA19BA3BABFDB8FB6DFDD8" box="[980,1202,586,610]" gridcol="2" gridrow="8" pageId="15" pageNumber="306">22.7–29.0</td>
<td id="76F55DA0FFAA19BA3C94FDB8FAB7FDD8" box="[1259,1384,586,610]" gridcol="3" gridrow="8" pageId="15" pageNumber="306">26.0±2.1</td>
</tr>
<tr id="352434DCFFAA19BA38D9FD80FAB7FD30" box="[166,1384,626,650]" gridrow="9" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FD80FDDEFD30" box="[166,513,626,650]" gridcol="0" gridrow="9" pageId="15" pageNumber="306">Prepelvic length</th>
<td id="76F55DA0FFAA19BA3B54FD80FC56FD30" box="[811,905,626,650]" gridcol="1" gridrow="9" pageId="15" pageNumber="306">53.0</td>
<td id="76F55DA0FFAA19BA3BABFD80FB6DFD30" box="[980,1202,626,650]" gridcol="2" gridrow="9" pageId="15" pageNumber="306">50.9–56.8</td>
<td id="76F55DA0FFAA19BA3C94FD80FAB7FD30" box="[1259,1384,626,650]" gridcol="3" gridrow="9" pageId="15" pageNumber="306">54.0±1.9</td>
</tr>
<tr id="352434DCFFAA19BA38D9FD68FAB7FD08" box="[166,1384,666,690]" gridrow="10" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FD68FDDEFD08" box="[166,513,666,690]" gridcol="0" gridrow="10" pageId="15" pageNumber="306">Preanal length</th>
<td id="76F55DA0FFAA19BA3B54FD68FC56FD08" box="[811,905,666,690]" gridcol="1" gridrow="10" pageId="15" pageNumber="306">78.7</td>
<td id="76F55DA0FFAA19BA3BABFD68FB6DFD08" box="[980,1202,666,690]" gridcol="2" gridrow="10" pageId="15" pageNumber="306">76.6–80.1</td>
<td id="76F55DA0FFAA19BA3C94FD68FAB7FD08" box="[1259,1384,666,690]" gridcol="3" gridrow="10" pageId="15" pageNumber="306">78.4±1.2</td>
</tr>
<tr id="352434DCFFAA19BA38D9FD30FAB7FD61" box="[166,1384,706,731]" gridrow="11" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FD30FDDEFD61" box="[166,513,706,731]" gridcol="0" gridrow="11" pageId="15" pageNumber="306">Dorsal-fin terminus to adipose fin</th>
<td id="76F55DA0FFAA19BA3B54FD30FC56FD61" box="[811,905,706,731]" gridcol="1" gridrow="11" pageId="15" pageNumber="306">21.9</td>
<td id="76F55DA0FFAA19BA3BABFD30FB6DFD61" box="[980,1202,706,731]" gridcol="2" gridrow="11" pageId="15" pageNumber="306">18.6–23.2</td>
<td id="76F55DA0FFAA19BA3C94FD30FAB7FD61" box="[1259,1384,706,731]" gridcol="3" gridrow="11" pageId="15" pageNumber="306">21.4±1.3</td>
</tr>
<tr id="352434DCFFAA19BA38D9FD19FAB7FCB9" box="[166,1384,747,771]" gridrow="12" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FD19FDDEFCB9" box="[166,513,747,771]" gridcol="0" gridrow="12" pageId="15" pageNumber="306">Body depth</th>
<td id="76F55DA0FFAA19BA3B54FD19FC56FCB9" box="[811,905,747,771]" gridcol="1" gridrow="12" pageId="15" pageNumber="306">38.7</td>
<td id="76F55DA0FFAA19BA3BABFD19FB6DFCB9" box="[980,1202,747,771]" gridcol="2" gridrow="12" pageId="15" pageNumber="306">33.6–42.4</td>
<td id="76F55DA0FFAA19BA3C94FD19FAB7FCB9" box="[1259,1384,747,771]" gridcol="3" gridrow="12" pageId="15" pageNumber="306">38.2±2.5</td>
</tr>
<tr id="352434DCFFAA19BA38D9FCE1FAB7FC91" box="[166,1384,787,811]" gridrow="13" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FCE1FDDEFC91" box="[166,513,787,811]" gridcol="0" gridrow="13" pageId="15" pageNumber="306">Pectoral-fin length</th>
<td id="76F55DA0FFAA19BA3B54FCE1FC56FC91" box="[811,905,787,811]" gridcol="1" gridrow="13" pageId="15" pageNumber="306">18.6</td>
<td id="76F55DA0FFAA19BA3BABFCE1FB6DFC91" box="[980,1202,787,811]" gridcol="2" gridrow="13" pageId="15" pageNumber="306">18.0–21.9</td>
<td id="76F55DA0FFAA19BA3C94FCE1FAB7FC91" box="[1259,1384,787,811]" gridcol="3" gridrow="13" pageId="15" pageNumber="306">19.9±1.4</td>
</tr>
<tr id="352434DCFFAA19BA38D9FCC9FAB7FCE9" box="[166,1384,827,851]" gridrow="14" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FCC9FDDEFCE9" box="[166,513,827,851]" gridcol="0" gridrow="14" pageId="15" pageNumber="306">Pectoral-fin base length</th>
<td id="76F55DA0FFAA19BA3B54FCC9FC56FCE9" box="[811,905,827,851]" gridcol="1" gridrow="14" pageId="15" pageNumber="306">4.5</td>
<td id="76F55DA0FFAA19BA3BABFCC9FB6DFCE9" box="[980,1202,827,851]" gridcol="2" gridrow="14" pageId="15" pageNumber="306">4.3–5.7</td>
<td id="76F55DA0FFAA19BA3C94FCC9FAB7FCE9" box="[1259,1384,827,851]" gridcol="3" gridrow="14" pageId="15" pageNumber="306">5.0±0.4</td>
</tr>
<tr id="352434DCFFAA19BA38D9FC91FAB7FCC6" box="[166,1384,867,892]" gridrow="15" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FC91FDDEFCC6" box="[166,513,867,892]" gridcol="0" gridrow="15" pageId="15" pageNumber="306">Pelvic-fin length</th>
<td id="76F55DA0FFAA19BA3B54FC91FC56FCC6" box="[811,905,867,892]" gridcol="1" gridrow="15" pageId="15" pageNumber="306">17.8</td>
<td id="76F55DA0FFAA19BA3BABFC91FB6DFCC6" box="[980,1202,867,892]" gridcol="2" gridrow="15" pageId="15" pageNumber="306">16.3–20.6</td>
<td id="76F55DA0FFAA19BA3C94FC91FAB7FCC6" box="[1259,1384,867,892]" gridcol="3" gridrow="15" pageId="15" pageNumber="306">18.8±1.5</td>
</tr>
<tr id="352434DCFFAA19BA38D9FC79FAB7FC1E" box="[166,1384,907,932]" gridrow="16" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FC79FDDEFC1E" box="[166,513,907,932]" gridcol="0" gridrow="16" pageId="15" pageNumber="306">Pelvic-fin base length</th>
<td id="76F55DA0FFAA19BA3B54FC79FC56FC1E" box="[811,905,907,932]" gridcol="1" gridrow="16" pageId="15" pageNumber="306">4.2</td>
<td id="76F55DA0FFAA19BA3BABFC79FB6DFC1E" box="[980,1202,907,932]" gridcol="2" gridrow="16" pageId="15" pageNumber="306">3.9–5.5</td>
<td id="76F55DA0FFAA19BA3C94FC79FAB7FC1E" box="[1259,1384,907,932]" gridcol="3" gridrow="16" pageId="15" pageNumber="306">4.6±0.5</td>
</tr>
<tr id="352434DCFFAA19BA38D9FC46FAB7FC76" box="[166,1384,948,972]" gridrow="17" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FC46FDDEFC76" box="[166,513,948,972]" gridcol="0" gridrow="17" pageId="15" pageNumber="306">Dorsal-fin height</th>
<td id="76F55DA0FFAA19BA3B54FC46FC56FC76" box="[811,905,948,972]" gridcol="1" gridrow="17" pageId="15" pageNumber="306">21.6</td>
<td id="76F55DA0FFAA19BA3BABFC46FB6DFC76" box="[980,1202,948,972]" gridcol="2" gridrow="17" pageId="15" pageNumber="306">21.4–27.2</td>
<td id="76F55DA0FFAA19BA3C94FC46FAB7FC76" box="[1259,1384,948,972]" gridcol="3" gridrow="17" pageId="15" pageNumber="306">23.6±1.9</td>
</tr>
<tr id="352434DCFFAA19BA38D9FC2EFAB7FC4E" box="[166,1384,988,1012]" gridrow="18" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FC2EFDDEFC4E" box="[166,513,988,1012]" gridcol="0" gridrow="18" pageId="15" pageNumber="306">Dorsal-fin base length</th>
<td id="76F55DA0FFAA19BA3B54FC2EFC56FC4E" box="[811,905,988,1012]" gridcol="1" gridrow="18" pageId="15" pageNumber="306">18.3</td>
<td id="76F55DA0FFAA19BA3BABFC2EFB6DFC4E" box="[980,1202,988,1012]" gridcol="2" gridrow="18" pageId="15" pageNumber="306">16.5–19.6</td>
<td id="76F55DA0FFAA19BA3C94FC2EFAB7FC4E" box="[1259,1384,988,1012]" gridcol="3" gridrow="18" pageId="15" pageNumber="306">18.5±0.9</td>
</tr>
<tr id="352434DCFFAA19BA38D9FBF6FAB7FBA6" box="[166,1384,1028,1052]" gridrow="19" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FBF6FDDEFBA6" box="[166,513,1028,1052]" gridcol="0" gridrow="19" pageId="15" pageNumber="306">Adipose-fin height</th>
<td id="76F55DA0FFAA19BA3B54FBF6FC56FBA6" box="[811,905,1028,1052]" gridcol="1" gridrow="19" pageId="15" pageNumber="306">7.7</td>
<td id="76F55DA0FFAA19BA3BABFBF6FB6DFBA6" box="[980,1202,1028,1052]" gridcol="2" gridrow="19" pageId="15" pageNumber="306">6.7–9.5</td>
<td id="76F55DA0FFAA19BA3C94FBF6FAB7FBA6" box="[1259,1384,1028,1052]" gridcol="3" gridrow="19" pageId="15" pageNumber="306">7.8±0.8</td>
</tr>
<tr id="352434DCFFAA19BA38D9FBDEFAB7FBFF" box="[166,1384,1068,1093]" gridrow="20" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FBDEFDDEFBFF" box="[166,513,1068,1093]" gridcol="0" gridrow="20" pageId="15" pageNumber="306">Adipose-fin base length</th>
<td id="76F55DA0FFAA19BA3B54FBDEFC56FBFF" box="[811,905,1068,1093]" gridcol="1" gridrow="20" pageId="15" pageNumber="306">2.8</td>
<td id="76F55DA0FFAA19BA3BABFBDEFB6DFBFF" box="[980,1202,1068,1093]" gridcol="2" gridrow="20" pageId="15" pageNumber="306">2.8–4.2</td>
<td id="76F55DA0FFAA19BA3C94FBDEFAB7FBFF" box="[1259,1384,1068,1093]" gridcol="3" gridrow="20" pageId="15" pageNumber="306">3.2±0.5</td>
</tr>
<tr id="352434DCFFAA19BA38D9FBA7FAB7FBD7" box="[166,1384,1109,1133]" gridrow="21" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FBA7FDDEFBD7" box="[166,513,1109,1133]" gridcol="0" gridrow="21" pageId="15" pageNumber="306">Anal-fin length</th>
<td id="76F55DA0FFAA19BA3B54FBA7FC56FBD7" box="[811,905,1109,1133]" gridcol="1" gridrow="21" pageId="15" pageNumber="306">13.4</td>
<td id="76F55DA0FFAA19BA3BABFBA7FB6DFBD7" box="[980,1202,1109,1133]" gridcol="2" gridrow="21" pageId="15" pageNumber="306">13.4–17.3</td>
<td id="76F55DA0FFAA19BA3C94FBA7FAB7FBD7" box="[1259,1384,1109,1133]" gridcol="3" gridrow="21" pageId="15" pageNumber="306">14.8±1.2</td>
</tr>
<tr id="352434DCFFAA19BA38D9FB8FFAB7FB2F" box="[166,1384,1149,1173]" gridrow="22" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FB8FFDDEFB2F" box="[166,513,1149,1173]" gridcol="0" gridrow="22" pageId="15" pageNumber="306">Anal-fin base length</th>
<td id="76F55DA0FFAA19BA3B54FB8FFC56FB2F" box="[811,905,1149,1173]" gridcol="1" gridrow="22" pageId="15" pageNumber="306">14.2</td>
<td id="76F55DA0FFAA19BA3BABFB8FFB6DFB2F" box="[980,1202,1149,1173]" gridcol="2" gridrow="22" pageId="15" pageNumber="306">11.7–14.8</td>
<td id="76F55DA0FFAA19BA3C94FB8FFAB7FB2F" box="[1259,1384,1149,1173]" gridcol="3" gridrow="22" pageId="15" pageNumber="306">13.3±1.0</td>
</tr>
<tr id="352434DCFFAA19BA38D9FB57FAB7FB07" box="[166,1384,1189,1213]" gridrow="23" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FB57FDDEFB07" box="[166,513,1189,1213]" gridcol="0" gridrow="23" pageId="15" pageNumber="306">Caudal peduncle depth</th>
<td id="76F55DA0FFAA19BA3B54FB57FC56FB07" box="[811,905,1189,1213]" gridcol="1" gridrow="23" pageId="15" pageNumber="306">12.2</td>
<td id="76F55DA0FFAA19BA3BABFB57FB6DFB07" box="[980,1202,1189,1213]" gridcol="2" gridrow="23" pageId="15" pageNumber="306">11.8–13.0</td>
<td id="76F55DA0FFAA19BA3C94FB57FAB7FB07" box="[1259,1384,1189,1213]" gridcol="3" gridrow="23" pageId="15" pageNumber="306">12.4±0.4</td>
</tr>
<tr id="352434DCFFAA19BA38D9FB3FFAB7FB5C" box="[166,1384,1229,1254]" gridrow="24" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FB3FFDDEFB5C" box="[166,513,1229,1254]" gridcol="0" gridrow="24" pageId="15" pageNumber="306">Caudal peduncle length</th>
<td id="76F55DA0FFAA19BA3B54FB3FFC56FB5C" box="[811,905,1229,1254]" gridcol="1" gridrow="24" pageId="15" pageNumber="306">11.5</td>
<td id="76F55DA0FFAA19BA3BABFB3FFB6DFB5C" box="[980,1202,1229,1254]" gridcol="2" gridrow="24" pageId="15" pageNumber="306">9.7–12.6</td>
<td id="76F55DA0FFAA19BA3C94FB3FFAB7FB5C" box="[1259,1384,1229,1254]" gridcol="3" gridrow="24" pageId="15" pageNumber="306">11.6±0.8</td>
</tr>
<tr id="352434DCFFAA19BA38D9FAECFAB7FA8D" box="[166,1384,1310,1335]" gridrow="25" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FAECFDDEFA8D" box="[166,513,1310,1335]" gridcol="0" gridrow="25" pageId="15" pageNumber="306">Eye diameter as % of snout length</th>
<td id="76F55DA0FFAA19BA3B54FAECFC56FA8D" box="[811,905,1310,1335]" gridcol="1" gridrow="25" pageId="15" pageNumber="306">74.1</td>
<td id="76F55DA0FFAA19BA3BABFAECFB6DFA8D" box="[980,1202,1310,1335]" gridcol="2" gridrow="25" pageId="15" pageNumber="306">58.6–88.0</td>
<td id="76F55DA0FFAA19BA3C94FAECFAB7FA8D" box="[1259,1384,1310,1335]" gridcol="3" gridrow="25" pageId="15" pageNumber="306">74.7±9.4</td>
</tr>
<tr id="352434DCFFAA19BA38D9FAB4FAB7FAE4" box="[166,1384,1350,1374]" gridrow="26" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FAB4FDDEFAE4" box="[166,513,1350,1374]" gridcol="0" gridrow="26" pageId="15" pageNumber="306">Eye diameter as % of head length</th>
<td id="76F55DA0FFAA19BA3B54FAB4FC56FAE4" box="[811,905,1350,1374]" gridcol="1" gridrow="26" pageId="15" pageNumber="306">26.4</td>
<td id="76F55DA0FFAA19BA3BABFAB4FB6DFAE4" box="[980,1202,1350,1374]" gridcol="2" gridrow="26" pageId="15" pageNumber="306">24.1–29.6</td>
<td id="76F55DA0FFAA19BA3C94FAB4FAB7FAE4" box="[1259,1384,1350,1374]" gridcol="3" gridrow="26" pageId="15" pageNumber="306">26.6±1.8</td>
</tr>
<tr id="352434DCFFAA19BA38D9FA9CFAB7FA3D" box="[166,1384,1390,1415]" gridrow="27" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FA9CFDDEFA3D" box="[166,513,1390,1415]" gridcol="0" gridrow="27" pageId="15" pageNumber="306">Snout length as % of head length</th>
<td id="76F55DA0FFAA19BA3B54FA9CFC56FA3D" box="[811,905,1390,1415]" gridcol="1" gridrow="27" pageId="15" pageNumber="306">35.6</td>
<td id="76F55DA0FFAA19BA3BABFA9CFB6DFA3D" box="[980,1202,1390,1415]" gridcol="2" gridrow="27" pageId="15" pageNumber="306">32–41.1</td>
<td id="76F55DA0FFAA19BA3C94FA9CFAB7FA3D" box="[1259,1384,1390,1415]" gridcol="3" gridrow="27" pageId="15" pageNumber="306">36.0±2.9</td>
</tr>
<tr id="352434DCFFAA19BA38D9FA4DFAB7FA6D" box="[166,1384,1471,1495]" gridrow="28" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9FA4DFAB7FA6D" box="[166,1384,1471,1495]" colspan="4" colspanRight="3" gridcol="0" gridrow="28" pageId="15" pageNumber="306">MERISTICS</th>
</tr>
<tr id="352434DCFFAA19BA38D9FA15FAB7FA45" box="[166,1384,1511,1535]" gridrow="29" pageId="15" pageNumber="306" rowspan-1="1" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9FA15FDDEFA45" box="[166,513,1511,1535]" gridcol="0" gridrow="29" pageId="15" pageNumber="306">Scales in lateral line</th>
<td id="76F55DA0FFAA19BA3BABFA15FB6DFA45" box="[980,1202,1511,1535]" gridcol="2" gridrow="29" pageId="15" pageNumber="306">40 (2), 41 (6), 42* (5)</td>
</tr>
<tr id="352434DCFFAA19BA38D9F9FDFAB7F992" box="[166,1384,1551,1576]" gridrow="30" pageId="15" pageNumber="306" rowspan-1="1" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9F9FDFDDEF992" box="[166,513,1551,1576]" gridcol="0" gridrow="30" pageId="15" pageNumber="306">Scales LL to dorsal</th>
<td id="76F55DA0FFAA19BA3BABF9FDFB6DF992" box="[980,1202,1551,1576]" gridcol="2" gridrow="30" pageId="15" pageNumber="306">7.5* (13)</td>
</tr>
<tr id="352434DCFFAA19BA38D9F9CAFAB7F9EA" box="[166,1384,1592,1616]" gridrow="31" pageId="15" pageNumber="306" rowspan-1="1" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9F9CAFDDEF9EA" box="[166,513,1592,1616]" gridcol="0" gridrow="31" pageId="15" pageNumber="306">Scales LL to pelvic</th>
<td id="76F55DA0FFAA19BA3BABF9CAFB6DF9EA" box="[980,1202,1592,1616]" gridcol="2" gridrow="31" pageId="15" pageNumber="306">6 (1), 7* (12)</td>
</tr>
<tr id="352434DCFFAA19BA38D9F992FAB7F9C2" box="[166,1384,1632,1656]" gridrow="32" pageId="15" pageNumber="306" rowspan-1="1" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9F992FDDEF9C2" box="[166,513,1632,1656]" gridcol="0" gridrow="32" pageId="15" pageNumber="306">Scales around caudal peduncle</th>
<td id="76F55DA0FFAA19BA3BABF992FB6DF9C2" box="[980,1202,1632,1656]" gridcol="2" gridrow="32" pageId="15" pageNumber="306">16* (13)</td>
</tr>
<tr id="352434DCFFAA19BA38D9F97AFAB7F91A" box="[166,1384,1672,1696]" gridrow="33" pageId="15" pageNumber="306">
<th id="76F55DA0FFAA19BA38D9F97AFDDEF91A" box="[166,513,1672,1696]" gridcol="0" gridrow="33" pageId="15" pageNumber="306">Dorsal-fin count</th>
<td id="76F55DA0FFAA19BA3B54F97AFAB7F91A" box="[811,1384,1672,1696]" colspan="3" colspanRight="2" gridcol="1" gridrow="33" pageId="15" pageNumber="306">ii-14 (1), iii-13 (1), iii-14* (10), iii-15 (1)</td>
</tr>
<tr id="352434DCFFAA19BA38D9F942FAB7F973" box="[166,1384,1712,1737]" gridrow="34" pageId="15" pageNumber="306" rowspan-1="1" rowspan-3="1">
<th id="76F55DA0FFAA19BA38D9F942FDDEF973" box="[166,513,1712,1737]" gridcol="0" gridrow="34" pageId="15" pageNumber="306">Anal-fin count</th>
<td id="76F55DA0FFAA19BA3BABF942FB6DF973" box="[980,1202,1712,1737]" gridcol="2" gridrow="34" pageId="15" pageNumber="306">iii-10* (10), iii-11 (3)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30E6515FFAAE65438B8F902FDD7FF43" lastPageId="17" lastPageNumber="308" pageId="15" pageNumber="306" type="description">
<paragraph id="8BAB369EFFAAE64A38B8F902FC20F850" blockId="15.[151,1436,1776,2027]" pageId="15" pageNumber="306">
<emphasis id="B960EA8CFFAAE64A38B8F902FE86F8B0" bold="true" box="[199,345,1776,1802]" pageId="15" pageNumber="306">Description.</emphasis>
Morphometrics and meristics for the 
<typeStatus id="54AF883CFFAAE64A3B7CF902FCBAF8B0" box="[771,869,1776,1802]" pageId="15" pageNumber="306" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54AF883CFFAAE64A3BE3F903FBD8F8B1" box="[924,1031,1777,1803]" pageId="15" pageNumber="306" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C144D1DFFAAE64A3C51F902FAECF8B0" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[1070,1331,1776,1802]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="15" pageNumber="306" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFAAE64A3C51F902FAECF8B0" box="[1070,1331,1776,1802]" italics="true" pageId="15" pageNumber="306">Distichodus mbiniensis</emphasis>
</taxonomicName>
are summarized in table 6. 
<typeStatus id="54AF883CFFAAE64A3912F8E2FE0AF890" box="[365,469,1808,1834]" pageId="15" pageNumber="306" type="holotype">Holotype</typeStatus>
shown in figure 7 and the live color of the 
<typeStatus id="54AF883CFFAAE64A3BCBF8E2FBC9F890" box="[948,1046,1808,1834]" pageId="15" pageNumber="306" type="holotype">holotype</typeStatus>
and a 
<typeStatus id="54AF883CFFAAE64A3C21F8E3FB61F891" box="[1118,1214,1809,1835]" pageId="15" pageNumber="306" type="paratype">paratype</typeStatus>
are shown in figure 9. Maximum sized observed: 
<quantity id="4CEC9B7BFFAAE64A3992F8C2FDBDF8F1" box="[493,610,1840,1867]" metricMagnitude="-1" metricUnit="m" metricValue="1.2209999999999999" pageId="15" pageNumber="306" unit="mm" value="122.1">122.1 mm</quantity>
SL. Body laterally compressed and moderately elongate. Dorsal profile with a convex curve from tip of snout to the vertical of anterior margin of eye, weakly concave from the vertical of the eye to posterior margin of head, and convex to origin of dorsal fin. Profile weakly convex from origin of dorsal to adipose fin origin; weakly concave from adipose fin origin to caudal fin. Ventral profile broadly convex from snout to origin of anal fin; weakly concave from anal-fin terminus to caudal fin. Body covered in relatively large ctenoid scales; extending over the proximal half of adipose fin and caudal fin.
</paragraph>
<paragraph id="8BAB369EFFB5E65538B8FF65FBBEFE33" blockId="16.[151,1436,151,393]" pageId="16" pageNumber="307">Mouth subterminal; two rows of bicuspid teeth in the upper and lower jaws. Dorsal fin with convex margin; its origin just anterior to the vertical of the origin of the pelvic fins. Adipose fin two thirds the distance from dorsal-fin terminus to caudal fin origin; base entirely anterior to the vertical of anal-fin terminus; distal tip extending beyond vertical of anal-fin terminus. Caudal fin forked; upper and lower lobes slightly rounded with moderate point. Anal fin with concave margin; base nearly equal to longest ray. Pelvic-fin origin at mid-length of snout to caudal fin distance; nearly as long as head length. Pectoral-fin origin just posterior to vertical of posterior margin of subopercle; horizontal just inferior to inferior margin of subopercle; slightly longer than pelvic fin.</paragraph>
<caption id="DF6B6616FFB5E65538E8FC38FF2BFB90" ID-DOI="http://doi.org/10.5281/zenodo.4674092" ID-Zenodo-Dep="4674092" httpUri="https://zenodo.org/record/4674092/files/figure.png" pageId="16" pageNumber="307" startId="16.[151,250,970,994]" targetBox="[171,1414,441,919]" targetPageId="16">
<paragraph id="8BAB369EFFB5E65538E8FC38FF2BFB90" blockId="16.[151,1436,970,1066]" pageId="16" pageNumber="307">
<emphasis id="B960EA8CFFB5E65538E8FC38FECBFC58" bold="true" box="[151,276,970,994]" pageId="16" pageNumber="307">FIGURE 7.</emphasis>
<taxonomicName id="4C144D1DFFB5E6553964FC38FDD8FC58" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[283,519,970,994]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="307" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB5E6553964FC38FDD8FC58" box="[283,519,970,994]" italics="true" pageId="16" pageNumber="307">Distichodus mbiniensis</emphasis>
</taxonomicName>
, a new species, holotype, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by S. Raredon.
</paragraph>
</caption>
<caption id="DF6B6616FFB5E65538E8F9C9FAA0F9CB" ID-DOI="http://doi.org/10.5281/zenodo.4674094" ID-Zenodo-Dep="4674094" httpUri="https://zenodo.org/record/4674094/files/figure.png" pageId="16" pageNumber="307" startId="16.[151,250,1595,1619]" targetBox="[166,1436,1106,1569]" targetPageId="16">
<paragraph id="8BAB369EFFB5E65538E8F9C9FAA0F9CB" blockId="16.[151,1436,1595,1649]" pageId="16" pageNumber="307">
<emphasis id="B960EA8CFFB5E65538E8F9C9FECCF9E9" bold="true" box="[151,275,1595,1619]" pageId="16" pageNumber="307">FIGURE 8.</emphasis>
Lateral views of head region in 
<taxonomicName id="4C144D1DFFB5E6553A1FF9CEFCEEF9E9" authority="Schmidt &amp; Knobloch &amp; Barrientos, 2021" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[608,817,1596,1620]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="307" phylum="Chordata" rank="species" species="microps" status="sp. nov.">
<emphasis id="B960EA8CFFB5E6553A1FF9CEFCEEF9E9" box="[608,817,1596,1620]" italics="true" pageId="16" pageNumber="307">Distichodus microps</emphasis>
</taxonomicName>
<emphasis id="B960EA8CFFB5E6553B47F9CEFC56F9EE" bold="true" box="[824,905,1596,1620]" pageId="16" pageNumber="307">
<taxonomicNameLabel id="A25357F7FFB5E6553B47F9CEFC56F9EE" box="[824,905,1596,1620]" pageId="16" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A), 
<taxonomicName id="4C144D1DFFB5E6553BBEF9CEFB72F9EE" authority="Schmidt &amp; Knobloch &amp; Barrientos, 2021" authorityName="Schmidt &amp; Knobloch &amp; Barrientos" authorityYear="2021" box="[961,1197,1596,1620]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="307" phylum="Chordata" rank="species" species="mbiniensis" status="sp. nov.">
<emphasis id="B960EA8CFFB5E6553BBEF9CEFB72F9EE" box="[961,1197,1596,1620]" italics="true" pageId="16" pageNumber="307">Distichodus mbiniensis</emphasis>
</taxonomicName>
<emphasis id="B960EA8CFFB5E6553CCBF9CEFADAF9EE" bold="true" box="[1204,1285,1596,1620]" pageId="16" pageNumber="307">
<taxonomicNameLabel id="A25357F7FFB5E6553CCBF9CEFADAF9EE" box="[1204,1285,1596,1620]" pageId="16" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B), and topotypic 
<emphasis id="B960EA8CFFB5E65538AFF9ABFE3FF9CA" box="[208,480,1625,1649]" italics="true" pageId="16" pageNumber="307">
<taxonomicName id="4C144D1DFFB5E65538AFF9ABFE67F9CB" box="[208,440,1625,1649]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="307" phylum="Chordata" rank="species" species="notospilus">Distichodus notospilus</taxonomicName>
(C)
</emphasis>
. Same specimens and locality information as previous figures. Photographs by S. Raredon.
</paragraph>
</caption>
<paragraph id="8BAB369EFFB5E65438B8F96DFDD7FF43" blockId="16.[151,1437,1695,2010]" lastBlockId="17.[151,1437,151,249]" lastPageId="17" lastPageNumber="308" pageId="16" pageNumber="307">
<emphasis id="B960EA8CFFB5E65538B8F96DFE90F903" bold="true" box="[199,335,1695,1721]" pageId="16" pageNumber="307">Coloration.</emphasis>
Live coloration: in both large and small specimens the ground color is bronze with some yellow and gold along the sides of the head. Some dark melanophores on subopercle in larger specimens which appears to be absent in smaller specimens. Anal and pelvic fins bright red in smaller specimens; a darker brick-red in larger individuals (
<figureCitation id="132F2A1BFFB5E655395CF8F9FEB7F89F" box="[291,360,1803,1829]" captionStart="FIGURE 9" captionStartId="17.[151,250,850,874]" captionTargetBox="[151,1437,273,827]" captionTargetId="figure-62@17.[151,1437,273,827]" captionTargetPageId="17" captionText="FIGURE 9. Live coloration of Distichodus microps, USNM 451457, ALC 81.6 mm SL, voucher EqGui2017_0274, Equatorial Guinea, Litoral, Rio Mbia at Menzong, 32 m elev, 2.07529° N, 9.92378° E photograph by R.C. Schmidt (A); Distichodus mbiniensis, USNM 451319, ALC 61.3 mm SL, voucher EqGui2017_0130, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m, 1.29091° N, 10.9174° E, photograph by R. C. Schmidt (B); Topotypic Distichodus notospilus, OS 19544, ALC, 104.1 mm SL, Gabon, Province de l’Ogooué-Lolo, Right bank of Ogooué River at Doumé village, 0.84245° S, 12.96249° E, Photograph by B. Sidlauskas, included with permission (C); Holotype of Distichodus mbiniensis, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by R. C. Schmidt (D)." figureDoi="http://doi.org/10.5281/zenodo.4674096" httpUri="https://zenodo.org/record/4674096/files/figure.png" pageId="16" pageNumber="307">Fig. 9</figureCitation>
). Pectoral fins, adipose fin, and caudal fin clear to yellow. The dorsal fin with prominent black spot originating at the base of the third or fourth ray and extending anteriorly at an angle to the first ray. In small specimens this black marking extends to the distal tip of the dorsal fin but only extends distally about two-thirds in larger individuals (
<figureCitation id="132F2A1BFFB5E6553914F885FE67F828" box="[363,440,1911,1938]" captionStart="FIGURE 7" captionStartId="16.[151,250,970,994]" captionTargetBox="[171,1414,441,919]" captionTargetId="figure-155@16.[151,1437,417,947]" captionTargetPageId="16" captionText="FIGURE 7. Distichodus mbiniensis, a new species, holotype, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by S. Raredon." figureDoi="http://doi.org/10.5281/zenodo.4674092" httpUri="https://zenodo.org/record/4674092/files/figure.png" pageId="16" pageNumber="307">Figs. 7</figureCitation>
&amp; 
<figureCitation id="132F2A1BFFB5E65539A3F885FE33F82B" box="[476,492,1911,1937]" captionStart="FIGURE 9" captionStartId="17.[151,250,850,874]" captionTargetBox="[151,1437,273,827]" captionTargetId="figure-62@17.[151,1437,273,827]" captionTargetPageId="17" captionText="FIGURE 9. Live coloration of Distichodus microps, USNM 451457, ALC 81.6 mm SL, voucher EqGui2017_0274, Equatorial Guinea, Litoral, Rio Mbia at Menzong, 32 m elev, 2.07529° N, 9.92378° E photograph by R.C. Schmidt (A); Distichodus mbiniensis, USNM 451319, ALC 61.3 mm SL, voucher EqGui2017_0130, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m, 1.29091° N, 10.9174° E, photograph by R. C. Schmidt (B); Topotypic Distichodus notospilus, OS 19544, ALC, 104.1 mm SL, Gabon, Province de l’Ogooué-Lolo, Right bank of Ogooué River at Doumé village, 0.84245° S, 12.96249° E, Photograph by B. Sidlauskas, included with permission (C); Holotype of Distichodus mbiniensis, USNM 451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by R. C. Schmidt (D)." figureDoi="http://doi.org/10.5281/zenodo.4674096" httpUri="https://zenodo.org/record/4674096/files/figure.png" pageId="16" pageNumber="307">9</figureCitation>
). The last third of the dorsal fin is clear to yellow in smaller specimens and a darker red to yellow in larger specimens. Typical coloration after preservation is shown in figure 7. Specimens are medium to dark brown along the back and upper quarter of sides, lighter below. Scales along sides have darker spot on anterior and poster margin providing an effect of light horizontal stripes. Pigmentation on opercle and subopercle, and the spot a base of caudal fin more distinct than in live specimens. Fins beige with the dorsal fin with distinct black marking as described previously.
</paragraph>
</subSubSection>
<caption id="DF6B6616FFB4E65438E8FCA0FCF7FBDD" ID-DOI="http://doi.org/10.5281/zenodo.4674096" ID-Zenodo-Dep="4674096" httpUri="https://zenodo.org/record/4674096/files/figure.png" pageId="17" pageNumber="308" startId="17.[151,250,850,874]" targetBox="[151,1437,273,827]" targetPageId="17">
<paragraph id="8BAB369EFFB4E65438E8FCA0FCF7FBDD" blockId="17.[151,1437,850,1127]" pageId="17" pageNumber="308">
<emphasis id="B960EA8CFFB4E65438E8FCA0FECBFCD0" bold="true" box="[151,276,850,874]" pageId="17" pageNumber="308">FIGURE 9.</emphasis>
Live coloration of 
<taxonomicName id="4C144D1DFFB4E65439A1FCA0FCD9FCD1" authority=", USNM" authorityName="USNM" box="[478,774,850,875]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="microps">
<emphasis id="B960EA8CFFB4E65439A1FCA0FD6FFCD0" box="[478,688,850,874]" italics="true" pageId="17" pageNumber="308">Distichodus microps</emphasis>
, USNM
</taxonomicName>
451457, ALC 81.6 mm SL, voucher EqGui2017_0274, Equatorial Guinea, Litoral, Rio Mbia at Menzong, 32 m elev, 2.07529° N, 9.92378° E photograph by R.C. Schmidt (A); 
<taxonomicName id="4C144D1DFFB4E6543D41FC84FE58FC09" authority=", USNM" authorityName="USNM" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E6543D41FC84FEF0FC08" italics="true" pageId="17" pageNumber="308">Distichodus mbiniensis</emphasis>
, USNM
</taxonomicName>
451319, ALC 61.3 mm SL, voucher EqGui2017_0130, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m, 1.29091° N, 10.9174° E, photograph by R. C. Schmidt (B); Topotypic 
<taxonomicName id="4C144D1DFFB4E6543CD0FC4CFF68FC41" authority=", OS" authorityName="OS" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="notospilus">
<emphasis id="B960EA8CFFB4E6543CD0FC4CFA49FC6C" box="[1199,1430,958,982]" italics="true" pageId="17" pageNumber="308">Distichodus notospilus</emphasis>
, OS
</taxonomicName>
19544, ALC, 104.1 mm SL, Gabon, Province de l’Ogooué-Lolo, Right bank of Ogooué River at Doumé village, 0.84245° S, 12.96249° E, Photograph by B. Sidlauskas, included with permission (C); Holotype of 
<taxonomicName id="4C144D1DFFB4E6543C7BFBF4FA9BFBA5" authority=", USNM" authorityName="USNM" box="[1028,1348,1030,1055]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E6543C7BFBF4FB31FBA4" box="[1028,1262,1030,1054]" italics="true" pageId="17" pageNumber="308">Distichodus mbiniensis</emphasis>
, USNM
</taxonomicName>
451317, ALC, 99.9 mm SL, voucher EqGui2017_0127, Equatorial Guinea, Wele-Nzas, Rio Ntoro at Aconibe—Acurenam Rd., 668 m elev, 1.29091° N, 10.9174° E; Photograph by R. C. Schmidt (D).
</paragraph>
</caption>
<subSubSection id="C30E6515FFB4E65438B8FB67FB5AFB69" pageId="17" pageNumber="308" type="etymology">
<paragraph id="8BAB369EFFB4E65438B8FB67FB5AFB69" blockId="17.[151,1437,1173,1847]" pageId="17" pageNumber="308">
<emphasis id="B960EA8CFFB4E65438B8FB67FE91FB15" bold="true" box="[199,334,1173,1199]" pageId="17" pageNumber="308">Etymology.</emphasis>
The specific epithet refers to the 
<collectingCountry id="F303760EFFB4E6543AA9FB67FCC5FB15" box="[726,794,1173,1199]" name="Equatorial Guinea" pageId="17" pageNumber="308">Mbini</collectingCountry>
River which is the Ndowe name for the Rio Wele. The Ndowe are Bantu-speaking people that live along the coastal region in 
<collectingCountry id="F303760EFFB4E6543BCEFB4BFB5DFB69" box="[945,1154,1209,1235]" name="Equatorial Guinea" pageId="17" pageNumber="308">Equatorial Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6515FFB4E65438B8FB2EFEDBF88D" pageId="17" pageNumber="308" type="distribution">
<paragraph id="8BAB369EFFB4E65438B8FB2EFEC3FA49" blockId="17.[151,1437,1173,1847]" pageId="17" pageNumber="308">
<emphasis id="B960EA8CFFB4E65438B8FB2EFEBEFB4C" bold="true" box="[199,353,1244,1270]" pageId="17" pageNumber="308">Distribution.</emphasis>
This species is only known from the upper 
<collectingCountry id="F303760EFFB4E6543B3EFB2FFC5AFB4D" box="[833,901,1245,1271]" name="Equatorial Guinea" pageId="17" pageNumber="308">Mbini</collectingCountry>
River drainage (Rio Wele) in 
<collectingCountry id="F303760EFFB4E6543CAEFB2FFA43FB4D" box="[1233,1436,1245,1271]" name="Equatorial Guinea" pageId="17" pageNumber="308">Equatorial Guinea</collectingCountry>
but may also occur within this drainage in 
<collectingCountry id="F303760EFFB4E6543A13FAF3FD68FAA1" box="[620,695,1281,1307]" name="Gabon" pageId="17" pageNumber="308">Gabon</collectingCountry>
(
<figureCitation id="132F2A1BFFB4E6543ABAFAF3FCD6FAA1" box="[709,777,1281,1307]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1759]" captionTargetBox="[151,1436,211,1649]" captionTargetId="figure-19@7.[151,1436,211,1649]" captionTargetPageId="7" captionText="FIGURE 2. Phylogeny of Distichodus spp. inferred from 996 bp of cytochrome b (A). Phylogeny inferred from concatenated dataset of 996 bp CYTB, 729 bp MYH6, and 627 bp of CO1(B). Posterior probabilities from Bayesian inference above major branches; above 0.95 not shown and branches with support values &lt;70 were collapsed. Metadata and accession numbers for specimens included in the analyses shown in table 1 and average uncorrected p-distances among these different taxa shown in table 3." figureDoi="http://doi.org/10.5281/zenodo.4674082" httpUri="https://zenodo.org/record/4674082/files/figure.png" pageId="17" pageNumber="308">Fig. 2</figureCitation>
). Research expeditions in the Woleu R. (
<collectingCountry id="F303760EFFB4E6543CB6FAF3FAD0FAA1" box="[1225,1295,1281,1307]" name="Equatorial Guinea" pageId="17" pageNumber="308">Mbini</collectingCountry>
River drainage) in 
<collectingCountry id="F303760EFFB4E6543894FAD7FEE9FA85" box="[235,310,1317,1343]" name="Gabon" pageId="17" pageNumber="308">Gabon</collectingCountry>
discovered a new species of 
<taxonomicName id="4C144D1DFFB4E6543A05FAD7FD2AFA85" authorityName="G.Cuvier" authorityYear="1816" box="[634,757,1317,1343]" class="Actinopterygii" family="Mochokidae" genus="Synodontis" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="308" phylum="Chordata" rank="genus">
<emphasis id="B960EA8CFFB4E6543A05FAD7FD2AFA85" box="[634,757,1317,1343]" italics="true" pageId="17" pageNumber="308">Synodontis</emphasis>
</taxonomicName>
, but no specimens of 
<taxonomicName id="4C144D1DFFB4E6543B96FAD7FBB0FA85" authorityName="J.P.Muller &amp; Troschel" authorityYear="1844" box="[1001,1135,1317,1343]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="genus">
<emphasis id="B960EA8CFFB4E6543B96FAD7FBB0FA85" box="[1001,1135,1317,1343]" italics="true" pageId="17" pageNumber="308">Distichodus</emphasis>
</taxonomicName>
were collected at that time (
<bibRefCitation id="EF854B6FFFB4E65438E0FABBFE45FAD9" author="Friel, J. P. &amp; Sullivan J. P." box="[159,410,1353,1379]" pageId="17" pageNumber="308" pagination="3 - 12" refId="ref13049" refString="Friel, J. P. &amp; Sullivan J. P. (2008) Synodontis woleuensis (Siluriformes: Mochokidae), a new species of catfish from Gabon and Equatorial Guinea, Africa. Proceedings of the Academy of Natural Sciences of Philadelphia, 157, 3 - 12. https: // doi. org / 10.1635 / 0097 - 3157 (2008) 157 [3: SWSMAN] 2.0. CO; 2" type="journal article" year="2008">Friel &amp; Sullivan 2008</bibRefCitation>
). 
<taxonomicName id="4C144D1DFFB4E65439D3FABBFD6FFAD9" box="[428,688,1353,1379]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E65439D3FABBFD6FFAD9" box="[428,688,1353,1379]" italics="true" pageId="17" pageNumber="308">Distichodus mbiniensis</emphasis>
</taxonomicName>
was collected in relatively deep (~2 meters) pools with some flow. At the 
<typeStatus id="54AF883CFFB4E6543897FA9FFEC7FA3D" box="[232,280,1389,1415]" pageId="17" pageNumber="308">type</typeStatus>
locality they were collected in the outflow of a black-water reservoir impounded by the recent road construction. Additional sampling in the region is needed to determine how widespread this species is within the region. The expedition in 2017 was one of the first to collect fishes in the tributaries of the 
<collectingCountry id="F303760EFFB4E6543CE9FA47FB05FA75" box="[1174,1242,1461,1487]" name="Equatorial Guinea" pageId="17" pageNumber="308">Mbini</collectingCountry>
River in 
<collectingCountry id="F303760EFFB4E6543D3BFA47FEC6FA49" name="Equatorial Guinea" pageId="17" pageNumber="308">Equatorial Guinea</collectingCountry>
.
</paragraph>
<paragraph id="8BAB369EFFB4E65438B8FA0EFEDBF88D" blockId="17.[151,1437,1173,1847]" pageId="17" pageNumber="308">
<emphasis id="B960EA8CFFB4E65438B8FA0EFE1DF9AC" bold="true" box="[199,450,1532,1558]" pageId="17" pageNumber="308">Conservation Status.</emphasis>
<taxonomicName id="4C144D1DFFB4E65439B4FA0FFD0DF9AD" box="[459,722,1533,1559]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E65439B4FA0FFD0DF9AD" box="[459,722,1533,1559]" italics="true" pageId="17" pageNumber="308">Distichodus mbiniensis</emphasis>
</taxonomicName>
is only known to occur within the upper reaches of the 
<collectingCountry id="F303760EFFB4E6543D27FA0FFA43F9AD" box="[1368,1436,1533,1559]" name="Equatorial Guinea" pageId="17" pageNumber="308">Mbini</collectingCountry>
River drainage in 
<collectingCountry id="F303760EFFB4E654391FF9D3FDF3F981" box="[352,556,1569,1595]" name="Equatorial Guinea" pageId="17" pageNumber="308">Equatorial Guinea</collectingCountry>
but may also occur in this drainage in 
<collectingCountry id="F303760EFFB4E6543BA4F9D3FBF6F981" box="[987,1065,1569,1595]" name="Gabon" pageId="17" pageNumber="308">Gabon</collectingCountry>
. The extant of occurrence (EOO) for this species is estimated at 
<quantity id="4CEC9B7BFFB4E6543991F9B7FDB8F9E5" box="[494,615,1605,1632]" metricMagnitude="7" metricUnit="m" metricValue="1.9085" pageId="17" pageNumber="308" unit="km" value="19085.0">19,085 km</quantity>
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, determined in the IUCN Freshwater Mapping Application, and the species likely occurs in less than ten threat-based locations. These locations are the mainstrem of the 
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River and the major tributaries that flow into the 
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River. The habitats of 
<taxonomicName id="4C144D1DFFB4E6543BEEF97CFBF4F91D" box="[913,1067,1677,1703]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E6543BEEF97CFBF4F91D" box="[913,1067,1677,1703]" italics="true" pageId="17" pageNumber="308">D. mbiniensis</emphasis>
</taxonomicName>
are under threat due to pollution and sedimentation caused by ongoing road construction and logging activities in the region. Due to the restricted EOO, fewer than ten threat-based locations, and ongoing threats in the region that are negatively affecting the quality and extant of habitat for 
<taxonomicName id="4C144D1DFFB4E65439AEF908FDB4F8A9" box="[465,619,1785,1811]" class="Actinopterygii" family="Distichodontidae" genus="Distichodus" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="308" phylum="Chordata" rank="species" species="mbiniensis">
<emphasis id="B960EA8CFFB4E65439AEF908FDB4F8A9" box="[465,619,1785,1811]" italics="true" pageId="17" pageNumber="308">D. mbiniensis</emphasis>
</taxonomicName>
, this species is assessed as Vulnerable in the IUCN Red List (
<bibRefCitation id="EF854B6FFFB4E6543D5DF90BFF24F890" author="Schmidt, R. C. &amp; Knobloch, E. C. &amp; Barrientos, C." pageId="17" pageNumber="308" refId="ref14348" refString="Schmidt, R. C., Knobloch, E. C. &amp; Barrientos, C. (2020 b) Distichodus sp. nov. ' mbiniensis'. The IUCN Red List of Threatened Species, 2020, e. T 177754327 A 177754338. https: // doi. org / 10.2305 / IUCN. UK. 2020 - 3. RLTS. T 177754327 A 177754338. en" type="book" year="2020">
Schmidt 
<emphasis id="B960EA8CFFB4E6543DF7F908FF6BF88D" italics="true" pageId="17" pageNumber="308">et al.</emphasis>
2020 
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</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>