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<document id="C3C22ED4FF9F613FB271AC1401B31309" ID="10.11646/zootaxa.4147.3.1" ID-DOI="10.11646/zootaxa.4147.3.1" ID-GBIF-Dataset="9640ef3e-e587-4bd9-8e59-9f19075f4ac4" ID-ISSN="1175-5326" ID-Zenodo-Dep="261264" ID-ZooBank="B54FCD54-7514-408D-9B6F-62BF3DE4DA66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470291641489" checkinUser="plazi" docAuthor="Kment, Petr, Jindra, Zdeněk &amp; Berchi, Gavril Marius" docDate="2016" docId="03BD2655FFB0D20CFF7CFF71A65FE51F" docLanguage="en" docName="zootaxa.4147.3.1.pdf" docOrigin="Zootaxa 4147 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebrus syriacus Horvath 1896" docType="treatment" docVersion="7" lastPageNumber="219" masterDocId="FF845E2DFFA0D21EFFEBFFE6A464E147" masterDocTitle="Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera)" masterLastPageNumber="239" masterPageNumber="201" pageNumber="217" updateTime="1698366881363" updateUser="plazi">
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<mods:title id="B2E2053190DB0FE6DC818D81D53AD12A">Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera)</mods:title>
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<treatment id="03BD2655FFB0D20CFF7CFF71A65FE51F" ID-DOI="http://doi.org/10.5281/zenodo.6055920" ID-GBIF-Taxon="123514124" ID-Zenodo-Dep="6055920" LSID="urn:lsid:plazi:treatment:03BD2655FFB0D20CFF7CFF71A65FE51F" httpUri="http://treatment.plazi.org/id/03BD2655FFB0D20CFF7CFF71A65FE51F" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217">
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<taxonomicName id="4C14ECC0FFB0D20EFF7CFF71A672E1F5" ID-CoL="3JYFK" authority="Horvath, 1896" authorityName="Horvath" authorityYear="1896" box="[151,534,151,178]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB0D20EFF7CFF71A672E1F5" bold="true" box="[151,534,151,178]" pageId="16" pageNumber="217">
<emphasis id="B9604B51FFB0D20EFF7CFF71A53FE1F6" bold="true" box="[151,347,151,177]" italics="true" pageId="16" pageNumber="217">Hebrus syriacus</emphasis>
<bibRefCitation id="EF85EAB2FFB0D20EFE89FF7EA672E1F5" author="Horvath" box="[354,534,152,178]" pageId="16" pageNumber="217" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth, 1896</bibRefCitation>
</emphasis>
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<paragraph id="8BAB9743FFB0D20EFF7CFF5CA5E6E194" blockId="16.[151,534,151,211]" box="[151,386,186,211]" pageId="16" pageNumber="217">
(
<figureCitation id="132F8BC6FFB0D20EFF74FF5CA498E194" box="[159,252,186,211]" captionStart="FIGURES 23 28. 23 24" captionStartId="10.[151,264,1381,1404]" captionTargetBox="[151,1436,193,1359]" captionTargetId="figure@10.[151,1436,193,1360]" captionTargetPageId="10" captionText="FIGURES 23 28. 23 24 — Hebrus jeanneli djaneti Poisson, 1953: 23 — female, Algeria: near Tamanrasset (body length 2.04 mm); 24 — female, Morocco: Qued Reraïa (1.98 mm). 25 — H. montanus Kolenati, 1847, male, Morocco: Asni (1.92 mm). 26 — H. syriacus Horváth, 1896, female, Cyprus: Kyrenis (1.62 mm). 27 — H. vaillanti Poisson, 1953, male, Morocco: Massa River (1.88 mm). 28 — H. acutiscutatus Brown, 1951, Israel: Ein Gedi (1.90 mm)." httpUri="https://zenodo.org/record/261269/files/figure.png" pageId="16" pageNumber="217">Figs. 26</figureCitation>
,
<figureCitation id="132F8BC6FFB0D20EFEE3FF5DA535E194" box="[264,337,187,211]" captionStart-0="FIGURES 29 35" captionStart-1="FIGURES 36 39" captionStartId-0="16.[151,264,1747,1770]" captionStartId-1="17.[151,264,1446,1469]" captionTargetBox-0="[166,1417,411,1723]" captionTargetBox-1="[151,1436,193,1425]" captionTargetId-0="figure@16.[166,1418,411,1723]" captionTargetId-1="figure@17.[151,1436,193,1426]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 29 35. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 29 — body, dorsal view (magnification: 65 ×); 30 — body, lateral view (70 ×); 31 — head, dorsal view (180 ×); 32 — head, lateral view (200 ×); 33 — antenna, dorsal view (170 ×); 34 — segments IVa and IVb (350 ×); 35 — detail of membranous joint between segments IVa and IVb (1300 ×). Abbreviations: IVa, IVb—segments IVa and IVb; mj—membranous junction between segments IVa and IVb. Scale bars: 29 30 — 0.5 mm; 31 33 — 0.2 mm; 34 — 0.1 mm; 35 — 0.02 mm." captionText-1="FIGURES 36 39. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 36 — thorax, dorsal view (130 ×); 37 — metanotal elevation, dorsal view (320 ×); 38 — hemelytron, dorsal view (250 ×); 39 — hind leg (190 ×). Abbreviations: cl—clavus; co—corium. Scale bars: 36, 38 39 — 0.2 mm; 37 — 0.1 mm." httpUri-0="https://zenodo.org/record/261270/files/figure.png" httpUri-1="https://zenodo.org/record/261271/files/figure.png" pageId="16" pageNumber="217">2939</figureCitation>
,
<figureCitation id="132F8BC6FFB0D20EFEB6FF5CA51EE194" box="[349,378,186,211]" captionStart="FIGURE 67" captionStartId="25.[151,250,1028,1050]" captionTargetBox="[153,1431,196,1004]" captionTargetId="figure@25.[151,1435,193,1006]" captionTargetPageId="25" captionText="FIGURE 67. Distribution of Hebrus kiritshenkoi Kanyukova, 1997 (red) and H. syriacus Horváth, 1896 (green)." httpUri="https://zenodo.org/record/261278/files/figure.png" pageId="16" pageNumber="217">67</figureCitation>
)
</paragraph>
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<materialsCitation id="3B7C9D1EFFB0D20EFF7CFF1CA16BE057" ID-GBIF-Occurrence="1290989273" box="[151,1295,250,272]" collectionCode="HNHM" country="Israel" location="Haifa" pageId="16" pageNumber="217" specimenCount="1" specimenCount-female="1" stateProvince="Haifa" typeStatus="lectotype">
<treatmentCitationGroup id="AB04B06DFFB0D20EFF7CFF1CA16BE057" box="[151,1295,250,272]" pageId="16" pageNumber="217">
<taxonomicName id="4C14ECC0FFB0D20EFF7CFF1CA66FE057" ID-CoL="3JYFK" authority="Horvath, 1896: 326" authorityName="Horvath" authorityPageNumber="326" authorityYear="1896" box="[151,523,250,272]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB0D20EFF7CFF1CA559E057" box="[151,317,250,272]" italics="true" pageId="16" pageNumber="217">Hebrus syriacus</emphasis>
<treatmentCitation id="0AB5B152FFB0D20EFEA9FF1CA66FE057" author="Horvath" box="[322,523,250,272]" page="326" pageId="16" pageNumber="217" year="1896">
<bibRefCitation id="EF85EAB2FFB0D20EFEA9FF1CA5BCE057" author="Horvath" box="[322,472,250,272]" pageId="16" pageNumber="217" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth, 1896</bibRefCitation>
: 326
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54AF29E1FFB0D20EFDFFFF1DA61DE057" box="[532,633,251,272]" pageId="16" pageNumber="217" type="lectotype">Lectotype</typeStatus>
(designated by
<treatmentCitation id="0AB5B152FFB0D20EFCF0FF1CA791E057" author="Linnavuori" box="[795,1013,250,272]" page="94" pageId="16" pageNumber="217" year="1994">
<bibRefCitation id="EF85EAB2FFB0D20EFCF0FF1CA7ABE057" author="Linnavuori" box="[795,975,250,272]" pageId="16" pageNumber="217" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori, 1994</bibRefCitation>
: 94
</treatmentCitation>
):
<specimenCount id="9D125CCAFFB0D20EFBECFF1DA07EE057" box="[1031,1050,251,272]" pageId="16" pageNumber="217" type="female"></specimenCount>
,
<collectingCountry id="F303D7D3FFB0D20EFBCCFF1CA006E057" box="[1063,1122,250,272]" name="Israel" pageId="16" pageNumber="217">Israel</collectingCountry>
,
<collectingRegion id="49D059A1FFB0D20EFB80FF1CA0C7E057" box="[1131,1187,250,272]" country="Israel" name="Haifa" pageId="16" pageNumber="217">Haifa</collectingRegion>
(
<collectionCode id="ED050F86FFB0D20EFB5AFF1CA160E057" box="[1201,1284,250,272]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="217">HNHM</collectionCode>
).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30EC4C8FFB0D20FFF7CFED1A65EE797" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217" type="materials_examined">
<paragraph id="8BAB9743FFB0D20EFF7CFED1A564E036" blockId="16.[151,1437,311,369]" pageId="16" pageNumber="217">
<materialsCitation id="3B7C9D1EFFB0D20EFF7CFED1A564E036" ID-GBIF-Occurrence="1290989291" collectingDate="1939-07-07" collectionCode="NMPC" collectorName="H. Lindberg &amp; P. Kment" country="Cyprus" county="Ammos" location="Material" municipality="Kyrenis" pageId="16" pageNumber="217" specimenCount="1" specimenCount-female="1">
<emphasis id="B9604B51FFB0D20EFF7CFED1A669E017" bold="true" box="[151,525,311,336]" pageId="16" pageNumber="217">
<location id="8ECBC198FFB0D20EFF7CFED1A49BE017" LSID="urn:lsid:plazi:treatment:03BD2655FFB0D20CFF7CFF71A65FE51F:8ECBC198FFB0D20EFF7CFED1A49BE017" box="[151,255,311,336]" country="Cyprus" county="Ammos" municipality="Kyrenis" name="Material" pageId="16" pageNumber="217">Material</location>
examined.
<collectingCountry id="F303D7D3FFB0D20EFE79FED1A66CE017" box="[402,520,311,336]" name="Cyprus" pageId="16" pageNumber="217">CYPRUS</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BCF0D39FFB0D20EFDF2FED1A611E017" box="[537,629,311,336]" pageId="16" pageNumber="217">Kyrenis</collectingMunicipality>
,
<collectingCounty id="62CAEFCFFFB0D20EFD6EFEDEA685E017" box="[645,737,312,336]" pageId="16" pageNumber="217">Ammos</collectingCounty>
,
<collectingDate id="EFEE486BFFB0D20EFD1AFED1A701E017" box="[753,869,311,336]" pageId="16" pageNumber="217" value="1939-07-07">
<date id="FFAAB183FFB0D20EFD1AFED1A701E017" box="[753,869,311,336]" pageId="16" pageNumber="217" value="1939-07-07">7.vii.1939</date>
</collectingDate>
,
<specimenCount id="9D125CCAFFB0D20EFC9DFEDEA7C2E008" box="[886,934,312,336]" pageId="16" pageNumber="217" type="female">1 ♀</specimenCount>
,
<collectorName id="26E1F295FFB0D20EFC52FEDEA023E017" box="[953,1095,311,336]" pageId="16" pageNumber="217">H. Lindberg</collectorName>
lgt. &amp; det.,
<collectorName id="26E1F295FFB0D20EFB08FEDEA12BE008" box="[1251,1359,312,335]" pageId="16" pageNumber="217">P. Kment</collectorName>
revid. (
<collectionCode id="ED050F86FFB0D20EFF74FEBFA491E036" box="[159,245,345,369]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="217">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF6BC7CBFFB0D20EFF7CF935A7F7E623" httpUri="https://zenodo.org/record/261270/files/figure.png" pageId="16" pageNumber="217" targetBox="[166,1417,411,1723]" targetPageId="16">
<paragraph id="8BAB9743FFB0D20EFF7CF935A7F7E623" blockId="16.[151,1436,1747,1892]" pageId="16" pageNumber="217">
<emphasis id="B9604B51FFB0D20EFF7CF935A529E7AE" bold="true" box="[151,333,1747,1770]" pageId="16" pageNumber="217">FIGURES 2935</emphasis>
.
<taxonomicName id="4C14ECC0FFB0D20EFEB1F932A6FEE7AD" authority="Horvath, 1896" authorityName="Horvath" authorityYear="1896" box="[346,666,1748,1770]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB0D20EFEB1F932A59BE7AD" box="[346,511,1748,1770]" italics="true" pageId="16" pageNumber="217">Hebrus syriacus</emphasis>
Horváth, 1896
</taxonomicName>
, female, Cyprus: Kyrenis, scanning electron micrographs: 29—body, dorsal view (magnification: 65×); 30—body, lateral view (70×); 31—head, dorsal view (180×); 32—head, lateral view (200×); 33— antenna, dorsal view (170×); 34—segments IVa and IVb (350×); 35—detail of membranous joint between segments IVa and IVb (1300×). Abbreviations: IVa, IVb—segments IVa and IVb; mj—membranous junction between segments IVa and IVb. Scale bars: 2930—0.5 mm; 3133—0.2 mm; 34—0.1 mm; 35—0.02 mm.
</paragraph>
</caption>
<caption id="DF6BC7CBFFB1D20FFF7CFA40A06CE4BD" httpUri="https://zenodo.org/record/261271/files/figure.png" pageId="17" pageNumber="218" targetBox="[151,1436,193,1425]" targetPageId="17">
<paragraph id="8BAB9743FFB1D20FFF7CFA40A06CE4BD" blockId="17.[151,1436,1446,1530]" pageId="17" pageNumber="218">
<emphasis id="B9604B51FFB1D20FFF7CFA40A53CE4FB" bold="true" box="[151,344,1446,1469]" pageId="17" pageNumber="218">FIGURES 3639.</emphasis>
<taxonomicName id="4C14ECC0FFB1D20FFE88FA41A6D4E4FA" authority="Horvath, 1896" authorityName="Horvath" authorityYear="1896" box="[355,688,1447,1469]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB1D20FFE88FA41A668E4FA" box="[355,524,1447,1469]" italics="true" pageId="17" pageNumber="218">Hebrus syriacus</emphasis>
Horváth, 1896
</taxonomicName>
, female, Cyprus: Kyrenis, scanning electron micrographs: 36—thorax, dorsal view (130×); 37—metanotal elevation, dorsal view (320×); 38—hemelytron, dorsal view (250×); 39—hind leg (190×). Abbreviations: cl—clavus; co—corium. Scale bars: 36, 3839—0.2 mm; 37—0.1 mm.
</paragraph>
</caption>
<paragraph id="8BAB9743FFB1D20FFF2CF9C1A65EE797" blockId="17.[151,1437,1575,2032]" pageId="17" pageNumber="218">
<emphasis id="B9604B51FFB1D20FFF2CF9C1A5C5E707" bold="true" box="[199,417,1575,1600]" pageId="17" pageNumber="218">Descriptive notes.</emphasis>
The female examined (
<figureCitation id="132F8BC6FFB1D20FFD2FF9C1A743E707" box="[708,807,1575,1600]" captionStart="FIGURES 23 28. 23 24" captionStartId="10.[151,264,1381,1404]" captionTargetBox="[151,1436,193,1359]" captionTargetId="figure@10.[151,1436,193,1360]" captionTargetPageId="10" captionText="FIGURES 23 28. 23 24 — Hebrus jeanneli djaneti Poisson, 1953: 23 — female, Algeria: near Tamanrasset (body length 2.04 mm); 24 — female, Morocco: Qued Reraïa (1.98 mm). 25 — H. montanus Kolenati, 1847, male, Morocco: Asni (1.92 mm). 26 — H. syriacus Horváth, 1896, female, Cyprus: Kyrenis (1.62 mm). 27 — H. vaillanti Poisson, 1953, male, Morocco: Massa River (1.88 mm). 28 — H. acutiscutatus Brown, 1951, Israel: Ein Gedi (1.90 mm)." httpUri="https://zenodo.org/record/261269/files/figure.png" pageId="17" pageNumber="218">Figs. 26</figureCitation>
,
<figureCitation id="132F8BC6FFB1D20FFCD1F9C1A7E5E778" box="[826,897,1575,1599]" captionStart-0="FIGURES 29 35" captionStart-1="FIGURES 36 39" captionStartId-0="16.[151,264,1747,1770]" captionStartId-1="17.[151,264,1446,1469]" captionTargetBox-0="[166,1417,411,1723]" captionTargetBox-1="[151,1436,193,1425]" captionTargetId-0="figure@16.[166,1418,411,1723]" captionTargetId-1="figure@17.[151,1436,193,1426]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 29 35. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 29 — body, dorsal view (magnification: 65 ×); 30 — body, lateral view (70 ×); 31 — head, dorsal view (180 ×); 32 — head, lateral view (200 ×); 33 — antenna, dorsal view (170 ×); 34 — segments IVa and IVb (350 ×); 35 — detail of membranous joint between segments IVa and IVb (1300 ×). Abbreviations: IVa, IVb—segments IVa and IVb; mj—membranous junction between segments IVa and IVb. Scale bars: 29 30 — 0.5 mm; 31 33 — 0.2 mm; 34 — 0.1 mm; 35 — 0.02 mm." captionText-1="FIGURES 36 39. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 36 — thorax, dorsal view (130 ×); 37 — metanotal elevation, dorsal view (320 ×); 38 — hemelytron, dorsal view (250 ×); 39 — hind leg (190 ×). Abbreviations: cl—clavus; co—corium. Scale bars: 36, 38 39 — 0.2 mm; 37 — 0.1 mm." httpUri-0="https://zenodo.org/record/261270/files/figure.png" httpUri-1="https://zenodo.org/record/261271/files/figure.png" pageId="17" pageNumber="218">2939</figureCitation>
) fits the redescription by
<bibRefCitation id="EF85EAB2FFB1D20FFB2BF9C1A1F3E778" author="Linnavuori" box="[1216,1431,1575,1600]" pageId="17" pageNumber="218" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori (1994)</bibRefCitation>
, although the scutellum seems less bifid apically (see
<figureCitation id="132F8BC6FFB1D20FFD02F9ADA70BE723" box="[745,879,1611,1636]" captionStart="FIGURES 36 39" captionStartId="17.[151,264,1446,1469]" captionTargetBox="[151,1436,193,1425]" captionTargetId="figure@17.[151,1436,193,1426]" captionTargetPageId="17" captionText="FIGURES 36 39. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 36 — thorax, dorsal view (130 ×); 37 — metanotal elevation, dorsal view (320 ×); 38 — hemelytron, dorsal view (250 ×); 39 — hind leg (190 ×). Abbreviations: cl—clavus; co—corium. Scale bars: 36, 38 39 — 0.2 mm; 37 — 0.1 mm." httpUri="https://zenodo.org/record/261271/files/figure.png" pageId="17" pageNumber="218">Figs. 3637</figureCitation>
) and the membrane of the hemelytra is uniformly milky-brown and of the same colour as the posterior portion of the clavus (
<figureCitation id="132F8BC6FFB1D20FFC0FF989A05CE7CF" box="[996,1080,1647,1672]" captionStart="FIGURES 23 28. 23 24" captionStartId="10.[151,264,1381,1404]" captionTargetBox="[151,1436,193,1359]" captionTargetId="figure@10.[151,1436,193,1360]" captionTargetPageId="10" captionText="FIGURES 23 28. 23 24 — Hebrus jeanneli djaneti Poisson, 1953: 23 — female, Algeria: near Tamanrasset (body length 2.04 mm); 24 — female, Morocco: Qued Reraïa (1.98 mm). 25 — H. montanus Kolenati, 1847, male, Morocco: Asni (1.92 mm). 26 — H. syriacus Horváth, 1896, female, Cyprus: Kyrenis (1.62 mm). 27 — H. vaillanti Poisson, 1953, male, Morocco: Massa River (1.88 mm). 28 — H. acutiscutatus Brown, 1951, Israel: Ein Gedi (1.90 mm)." httpUri="https://zenodo.org/record/261269/files/figure.png" pageId="17" pageNumber="218">Fig. 26</figureCitation>
). The clavus has sparse pilosity and corium relatively dense pilosity (
<figureCitation id="132F8BC6FFB1D20FFDDDF975A6EDE7EB" box="[566,649,1683,1708]" captionStart="FIGURES 36 39" captionStartId="17.[151,264,1446,1469]" captionTargetBox="[151,1436,193,1425]" captionTargetId="figure@17.[151,1436,193,1426]" captionTargetPageId="17" captionText="FIGURES 36 39. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 36 — thorax, dorsal view (130 ×); 37 — metanotal elevation, dorsal view (320 ×); 38 — hemelytron, dorsal view (250 ×); 39 — hind leg (190 ×). Abbreviations: cl—clavus; co—corium. Scale bars: 36, 38 39 — 0.2 mm; 37 — 0.1 mm." httpUri="https://zenodo.org/record/261271/files/figure.png" pageId="17" pageNumber="218">Fig. 38</figureCitation>
). The membrane reaches the posterior 2/3 of mediotergite VII, leaving the apex of the dorsum free (
<figureCitation id="132F8BC6FFB1D20FFE32F951A64AE797" box="[473,558,1719,1744]" captionStart="FIGURES 23 28. 23 24" captionStartId="10.[151,264,1381,1404]" captionTargetBox="[151,1436,193,1359]" captionTargetId="figure@10.[151,1436,193,1360]" captionTargetPageId="10" captionText="FIGURES 23 28. 23 24 — Hebrus jeanneli djaneti Poisson, 1953: 23 — female, Algeria: near Tamanrasset (body length 2.04 mm); 24 — female, Morocco: Qued Reraïa (1.98 mm). 25 — H. montanus Kolenati, 1847, male, Morocco: Asni (1.92 mm). 26 — H. syriacus Horváth, 1896, female, Cyprus: Kyrenis (1.62 mm). 27 — H. vaillanti Poisson, 1953, male, Morocco: Massa River (1.88 mm). 28 — H. acutiscutatus Brown, 1951, Israel: Ein Gedi (1.90 mm)." httpUri="https://zenodo.org/record/261269/files/figure.png" pageId="17" pageNumber="218">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30EC4C8FFB1D20FFF2CF93AA7E3E67B" pageId="17" pageNumber="218" type="description">
<paragraph id="8BAB9743FFB1D20FFF2CF93AA7E3E67B" blockId="17.[151,1437,1575,2032]" pageId="17" pageNumber="218">
<emphasis id="B9604B51FFB1D20FFF2CF93AA514E7B4" bold="true" box="[199,368,1756,1779]" italics="true" pageId="17" pageNumber="218">Measurements</emphasis>
(mm): Body length: 1.66. Head width (across eyes): 0.4. Pronotum: median length: 0.38, anterior width: 0.38, humeral width: 0.69. Lengths of antennal segments: I: 0.18, II: 0.14, III: 0.20, IVa: 0.08, IVb: 0.24. Length of metafemur: 0.51, metatibia: 0.63, metatarsus: 0.22.
</paragraph>
</subSubSection>
<subSubSection id="C30EC4C8FFB1D20CFF2CF8A1A020E193" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="distribution">
<paragraph id="8BAB9743FFB1D20FFF2CF8A1A0CDE6C3" blockId="17.[151,1437,1575,2032]" pageId="17" pageNumber="218">
<emphasis id="B9604B51FFB1D20FFF2CF8A1A53DE627" bold="true" box="[199,345,1863,1888]" pageId="17" pageNumber="218">Distribution</emphasis>
(
<figureCitation id="132F8BC6FFB1D20FFE81F8A1A5DBE627" box="[362,447,1863,1888]" captionStart="FIGURE 67" captionStartId="25.[151,250,1028,1050]" captionTargetBox="[153,1431,196,1004]" captionTargetId="figure@25.[151,1435,193,1006]" captionTargetPageId="25" captionText="FIGURE 67. Distribution of Hebrus kiritshenkoi Kanyukova, 1997 (red) and H. syriacus Horváth, 1896 (green)." httpUri="https://zenodo.org/record/261278/files/figure.png" pageId="17" pageNumber="218">Fig. 67</figureCitation>
).
<emphasis id="B9604B51FFB1D20FFE3FF8A1A617E627" bold="true" box="[468,627,1863,1888]" pageId="17" pageNumber="218">
Asia:
<collectingCountry id="F303D7D3FFB1D20FFDF1F8A1A617E627" box="[538,627,1863,1888]" name="Cyprus" pageId="17" pageNumber="218">Cyprus</collectingCountry>
</emphasis>
(new record),
<emphasis id="B9604B51FFB1D20FFCF7F8A1A704E627" bold="true" box="[796,864,1863,1888]" pageId="17" pageNumber="218">
<collectingCountry id="F303D7D3FFB1D20FFCF7F8A1A704E627" box="[796,864,1863,1888]" name="Israel" pageId="17" pageNumber="218">Israel</collectingCountry>
</emphasis>
(
<bibRefCitation id="EF85EAB2FFB1D20FFC9BF8A1A074E627" author="Horvath" box="[880,1040,1863,1888]" pageId="17" pageNumber="218" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth 1896</bibRefCitation>
, as
<collectingCountry id="F303D7D3FFB1D20FFBD6F8A1A018E627" box="[1085,1148,1863,1888]" name="Syria" pageId="17" pageNumber="218">Syria</collectingCountry>
; Linnavuori 1961, 1994),
<emphasis id="B9604B51FFB1D20FFF7CF88DA488E6C3" bold="true" box="[151,236,1899,1924]" pageId="17" pageNumber="218">
<collectingCountry id="F303D7D3FFB1D20FFF7CF88DA488E6C3" box="[151,236,1899,1924]" name="Jordan" pageId="17" pageNumber="218">Jordan</collectingCountry>
</emphasis>
(Linnavuori 1961),
<emphasis id="B9604B51FFB1D20FFE24F88DA679E6C3" bold="true" box="[463,541,1899,1924]" pageId="17" pageNumber="218">
?
<collectingCountry id="F303D7D3FFB1D20FFE30F88DA679E6C3" box="[475,541,1899,1924]" name="Syria" pageId="17" pageNumber="218">Syria</collectingCountry>
</emphasis>
(
<bibRefCitation id="EF85EAB2FFB1D20FFDC7F88DA6D9E6C3" author="Stichel" box="[556,701,1899,1924]" pageId="17" pageNumber="218" refString="Stichel, W. (1956) Liste der palaarktischen Hemiptera-Heteroptera. In: Stichel, W. (Ed.), (1955 - 1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa (Hemiptera-Heteroptera Europae). Vol. 1. W. Stichel, Berlin-Hermsdorf, pp. 157 - 168." type="book chapter" year="1956">Stichel 1956</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB1D20FFD23F88DA712E6C3" author="Andersen" box="[712,886,1899,1924]" pageId="17" pageNumber="218" refString="Andersen, N. M. (1995) Infraorder Gerromorpha Popov, 1971 - semiaquatic bugs, pp. 17 - 114. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. The Netherlands Entomological Society, Amsterdam, xxvi + 222 pp." type="book chapter" year="1995">Andersen 1995</bibRefCitation>
).
<emphasis id="B9604B51FFB1D20FFC62F88DA0CDE6C3" bold="true" box="[905,1193,1899,1924]" pageId="17" pageNumber="218">
New species for
<collectingCountry id="F303D7D3FFB1D20FFBA2F88DA0C0E6C3" box="[1097,1188,1899,1924]" name="Cyprus" pageId="17" pageNumber="218">Cyprus</collectingCountry>
.
</emphasis>
</paragraph>
<paragraph id="8BAB9743FFB1D20CFF2CF869A020E193" blockId="17.[151,1437,1575,2032]" lastBlockId="18.[151,1437,151,1112]" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">
The
<typeStatus id="54AF29E1FFB1D20FFF11F876A54EE6EF" box="[250,298,1936,1960]" pageId="17" pageNumber="218">type</typeStatus>
locality in the original description was given as
<collectingCountry id="F303D7D3FFB1D20FFCBEF869A7FDE6EF" box="[853,921,1935,1960]" name="Syria" pageId="17" pageNumber="218">Syria</collectingCountry>
: Kaiffa (Reitter) (
<bibRefCitation id="EF85EAB2FFB1D20FFB81F869A16FE6EF" author="Horvath" box="[1130,1291,1935,1960]" pageId="17" pageNumber="218" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth 1896</bibRefCitation>
). Linnavuori (1961) listed four more specimens, one from
<collectingCountry id="F303D7D3FFB1D20FFD5DF855A765E68B" box="[694,769,1971,1996]" name="Jordan" pageId="17" pageNumber="218">Jordan</collectingCountry>
and three from Judea, all from J. Sahlbergs collection.
<bibRefCitation id="EF85EAB2FFB1D20FFF7CF831A500E6A8" author="Linnavuori" box="[151,356,2007,2032]" pageId="17" pageNumber="218" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori (1994)</bibRefCitation>
interpreted the
<typeStatus id="54AF29E1FFB1D20FFDF8F83EA627E6B7" box="[531,579,2008,2032]" pageId="17" pageNumber="218">type</typeStatus>
locality as
<collectingCountry id="F303D7D3FFB1D20FFD2CF831A768E6B7" box="[711,780,2007,2032]" name="Israel" pageId="17" pageNumber="218">Israel</collectingCountry>
, Haifa and speculated that since all new finds from
<collectingCountry id="F303D7D3FFB1D20FFAB5F831A1F8E6B7" box="[1374,1436,2007,2032]" name="Israel" pageId="17" pageNumber="218">Israel</collectingCountry>
belong to
<taxonomicName id="4C14ECC0FFB2D20CFEECFF7FA6ECE1F7" authority="Fallen, 1807" authorityName="Fallen" authorityYear="1807" box="[263,648,151,176]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="subSpecies" species="pusillus" subSpecies="pusillus">
<emphasis id="B9604B51FFB2D20CFEECFF7FA5BBE1F7" box="[263,479,151,176]" italics="true" pageId="18" pageNumber="219">H. pusillus pusillus</emphasis>
(
<bibRefCitation id="EF85EAB2FFB2D20CFE04FF71A6E5E1F7" author="Fallen" box="[495,641,151,176]" pageId="18" pageNumber="219" refString="Fallen, C. F. (1807) Monographia Cimicum Sveciae. C. G. Proft, Hafniae, 123 pp." type="book" year="1807">Fallén, 1807</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C14ECC0FFB2D20CFD7FFF7FA770E1F7" box="[660,788,153,176]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFD7FFF7FA770E1F7" box="[660,788,153,176]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
has possibly become extinct. Distribution of
<taxonomicName id="4C14ECC0FFB2D20CFAF7FF7FA1FFE1F7" box="[1308,1435,153,176]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFAF7FF7FA1FFE1F7" box="[1308,1435,153,176]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
in
<collectingCountry id="F303D7D3FFB2D20CFF5FFF5AA48BE192" box="[180,239,188,213]" name="Syria" pageId="18" pageNumber="219">Syria</collectingCountry>
requires confirmation as it seems to be based on the above mentioned error.
</paragraph>
</subSubSection>
<subSubSection id="C30EC4C8FFB2D20CFF2CFF39A65FE51F" pageId="18" pageNumber="219" type="discussion">
<paragraph id="8BAB9743FFB2D20CFF2CFF39A04BE0CF" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="219">
<materialsCitation id="3B7C9D1EFFB2D20CFF2CFF39A04FE0CF" ID-GBIF-Occurrence="1290989306" collectionCode="ICZN" country="Syria" county="Article" location="Andersen" municipality="This" pageId="18" pageNumber="219" specimenCount="5" typeStatus="syntype">
<emphasis id="B9604B51FFB2D20CFF2CFF39A52BE1BF" bold="true" box="[199,335,223,248]" pageId="18" pageNumber="219">Comments.</emphasis>
<bibRefCitation id="EF85EAB2FFB2D20CFEBCFF39A662E1BF" author="Horvath" box="[343,518,223,248]" pageId="18" pageNumber="219" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth (1896)</bibRefCitation>
described the species based on the macropterous female, but did not specify the number of specimens, although there was probably only one.
<bibRefCitation id="EF85EAB2FFB2D20CFCA6FEE2A06FE05A" author="Andersen" box="[845,1035,260,285]" pageId="18" pageNumber="219" refString="Andersen, N. M. (1995) Infraorder Gerromorpha Popov, 1971 - semiaquatic bugs, pp. 17 - 114. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. The Netherlands Entomological Society, Amsterdam, xxvi + 222 pp." type="book chapter" year="1995">
<location id="8ECBC198FFB2D20CFCA6FEE2A7DCE05A" LSID="urn:lsid:plazi:treatment:03BD2655FFB0D20CFF7CFF71A65FE51F:8ECBC198FFB2D20CFCA6FEE2A7DCE05A" box="[845,952,260,285]" country="Syria" county="Article" municipality="This" name="Andersen" pageId="18" pageNumber="219">Andersen</location>
(1995)
</bibRefCitation>
listed the
<typeStatus id="54AF29E1FFB2D20CFB6BFEE3A0D4E05A" box="[1152,1200,261,285]" pageId="18" pageNumber="219">type</typeStatus>
material as
<typeStatus id="54AF29E1FFB2D20CFAD1FEE3A1F8E05A" box="[1338,1436,261,285]" pageId="18" pageNumber="219" type="syntype">syntypes</typeStatus>
from
<collectingCountry id="F303D7D3FFB2D20CFF33FEC1A573E007" box="[216,279,295,320]" name="Syria" pageId="18" pageNumber="219">Syria</collectingCountry>
.
<bibRefCitation id="EF85EAB2FFB2D20CFECEFEC1A593E007" author="Linnavuori" box="[293,503,295,320]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">
<location id="8ECBC198FFB2D20CFECEFEC1A5C6E007" LSID="urn:lsid:plazi:treatment:03BD2655FFB0D20CFF7CFF71A65FE51F:8ECBC198FFB2D20CFECEFEC1A5C6E007" box="[293,418,295,320]" country="Syria" county="Article" municipality="This" name="Linnavuori" pageId="18" pageNumber="219">Linnavuori</location>
(1994)
</bibRefCitation>
redescribed the species based on a single female, presenting it as a type, and explicitely stated:
<location id="8ECBC198FFB2D20CFE81FEAAA5FEE022" LSID="urn:lsid:plazi:treatment:03BD2655FFB0D20CFF7CFF71A65FE51F:8ECBC198FFB2D20CFE81FEAAA5FEE022" box="[362,410,332,357]" country="Syria" county="Article" municipality="This" name="The" pageId="18" pageNumber="219">The</location>
<typeStatus id="54AF29E1FFB2D20CFE49FEAAA660E022" box="[418,516,332,357]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
is the only known specimen.
<collectingMunicipality id="6BCF0D39FFB2D20CFCB5FEAAA7F4E022" box="[862,912,332,357]" pageId="18" pageNumber="219">This</collectingMunicipality>
statement fulfilled the requirements of
<collectingCounty id="62CAEFCFFFB2D20CFAA5FEAAA1F8E022" box="[1358,1436,332,357]" pageId="18" pageNumber="219">Article</collectingCounty>
74.6 of the
<bibRefCitation id="EF85EAB2FFB2D20CFEFDFE96A5CDE0CF" author="ICZN" box="[278,425,368,392]" pageId="18" pageNumber="219" refString="ICZN (International Comission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
<collectionCode id="ED050F86FFB2D20CFEFDFE96A533E0CF" box="[278,343,368,392]" pageId="18" pageNumber="219">ICZN</collectionCode>
(1999)
</bibRefCitation>
for fixation of the
<typeStatus id="54AF29E1FFB2D20CFD96FE89A680E0CF" box="[637,740,367,392]" pageId="18" pageNumber="219" type="lectotype">lectotype</typeStatus>
by inference of the
<typeStatus id="54AF29E1FFB2D20CFC2EFE89A04FE0CF" box="[965,1067,367,392]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB9743FFB2D20CFF2CFE72A6F3E3B7" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="219">
<bibRefCitation id="EF85EAB2FFB2D20CFF2CFE72A510E0EA" author="Horvath" box="[199,372,404,429]" pageId="18" pageNumber="219" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth (1896)</bibRefCitation>
compared the new species to
<taxonomicName id="4C14ECC0FFB2D20CFD2EFE73A726E0EB" box="[709,834,404,428]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="B9604B51FFB2D20CFD2EFE73A726E0EB" box="[709,834,404,428]" italics="true" pageId="18" pageNumber="219">H. ruficeps</emphasis>
</taxonomicName>
: Similar to
<taxonomicName id="4C14ECC0FFB2D20CFC3AFE73A029E0EB" box="[977,1101,404,428]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="B9604B51FFB2D20CFC3AFE73A029E0EB" box="[977,1101,404,428]" italics="true" pageId="18" pageNumber="219">H. ruficeps</emphasis>
</taxonomicName>
, easily distinguished by body smaller and narrower, colouration of body pale, pronotum anteriorly not declivous, and posteriorly less dilated. Length
<quantity id="4CEC3AA6FFB2D20CFF04FE3AA52AE0B3" box="[239,334,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="219" unit="mm" value="1.5">1.5 mm</quantity>
.
<bibRefCitation id="EF85EAB2FFB2D20CFEB4FE3AA64BE0B3" author="Linnavuori" box="[351,559,476,501]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori (1994)</bibRefCitation>
redescribed the species based on the holotype and provided illustrations of the head in lateral view and thorax and hemelytron in dorsal view, and provided the following differential diagnosis:
<taxonomicName id="4C14ECC0FFB2D20CFEF3FDC3A5F3E37B" box="[280,407,549,572]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFEF3FDC3A5F3E37B" box="[280,407,549,572]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C14ECC0FFB2D20CFD0FFDC3A738E37A" box="[740,860,548,573]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B9604B51FFB2D20CFD0FFDC3A738E37A" box="[740,860,548,573]" italics="true" pageId="18" pageNumber="219">H. pusillus</emphasis>
</taxonomicName>
and the related species by the smaller and narrower body, much narrower and more parallel-sided pronotum and, especially, the apically bifid scutellum. Neither
<bibRefCitation id="EF85EAB2FFB2D20CFF7CFD8AA520E3C2" author="Horvath" box="[151,324,620,645]" pageId="18" pageNumber="219" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth (1896)</bibRefCitation>
nor
<bibRefCitation id="EF85EAB2FFB2D20CFE9CFD8AA621E3C3" author="Linnavuori" box="[375,581,620,645]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori (1994)</bibRefCitation>
described apical antennal segments.
<bibRefCitation id="EF85EAB2FFB2D20CFC08FD8AA01BE3C2" author="Jordan" box="[995,1151,620,645]" pageId="18" pageNumber="219" refString="Jordan, K. H. C. (1954) Uber einen neuen Naeogeus aus Sudfrankreich (Heteroptera: Hebridae). Beitrage zur Entomologie, 4, 601 - 604." type="journal article" year="1954">
<collectingCountry id="F303D7D3FFB2D20CFC08FD8AA04AE3C2" box="[995,1070,620,645]" name="Jordan" pageId="18" pageNumber="219">Jordan</collectingCountry>
(1954)
</bibRefCitation>
included
<taxonomicName id="4C14ECC0FFB2D20CFB06FD8BA108E3C3" box="[1261,1388,621,644]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFB06FD8BA108E3C3" box="[1261,1388,621,644]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
in a comparative table of seven European
<taxonomicName id="4C14ECC0FFB2D20CFDA1FD69A6F8E3EF" box="[586,668,655,680]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9604B51FFB2D20CFDA1FD69A6F8E3EF" box="[586,668,655,680]" italics="true" pageId="18" pageNumber="219">Hebrus</emphasis>
</taxonomicName>
species, and provided lengths of the antennal segments (mm): I— 0.18, II—0.16, III—0.20, IVa—0.10, IVb—0.24. However, the material
<bibRefCitation id="EF85EAB2FFB2D20CFC3DFD52A010E38A" author="Jordan" box="[982,1140,692,717]" pageId="18" pageNumber="219" refString="Jordan, K. H. C. (1954) Uber einen neuen Naeogeus aus Sudfrankreich (Heteroptera: Hebridae). Beitrage zur Entomologie, 4, 601 - 604." type="journal article" year="1954">
<collectingCountry id="F303D7D3FFB2D20CFC3DFD52A045E38A" box="[982,1057,692,717]" name="Jordan" pageId="18" pageNumber="219">Jordan</collectingCountry>
(1954)
</bibRefCitation>
used for comparison was not listed in that paper and remains unknown.
</paragraph>
<paragraph id="8BAB9743FFB2D20CFF2CFD1AA65FE51F" blockId="18.[151,1437,151,1112]" pageId="18" pageNumber="219">
<bibRefCitation id="EF85EAB2FFB2D20CFF2CFD1AA5FEE253" author="Linnavuori" box="[199,410,764,789]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori (1994)</bibRefCitation>
did not assign
<taxonomicName id="4C14ECC0FFB2D20CFDB2FD1BA6BAE253" box="[601,734,765,788]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFDB2FD1BA6BAE253" box="[601,734,765,788]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
to any of the subgenera; however, the metanotal elevation apically truncated with a wide V-shaped incision (
<bibRefCitation id="EF85EAB2FFB2D20CFD29FCF9A7BAE27F" author="Linnavuori" box="[706,990,799,824]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994" yearSuffix="b">
Linnavuori 1994:
<figureCitation id="132F8BC6FFB2D20CFC67FCF9A7AAE27F" box="[908,974,799,824]" captionStart="FIGURES 4 12" captionStartId="5.[151,264,1916,1939]" captionTargetBox="[184,1395,193,1886]" captionTargetId="figure@5.[184,1396,193,1887]" captionTargetPageId="5" captionText="FIGURES 4 12. Hebrus atlas sp. nov., male, paratype, Morocco: Ifrane env., scanning electron micrographs: 4 — body, dorsal view (magnification: 65 ×); 5 — body, lateral view (60 ×); 6 — head, dorsal view (200 ×); 7 — head, lateral view (180 ×); 8 — thorax, dorsal view (130 ×); 9 — metanotal elevation, dorsal view (320 ×); 10 — hemelytron, dorsal view (350 ×); 11 — hind leg (140 ×); 12 — metatarsus (370 ×). Scale bars: 4 5 — 0.5 mm; 6 8, 11 — 0.2 mm; 9 10, 12 — 0.1 mm." httpUri="https://zenodo.org/record/261266/files/figure.png" pageId="18" pageNumber="219">Fig. 4</figureCitation>
b
</bibRefCitation>
), antennal segments I and II subequal in length (
<bibRefCitation id="EF85EAB2FFB2D20CFF1BFCA2A5E0E21A" author="Stichel" box="[240,388,836,861]" pageId="18" pageNumber="219" refString="Stichel, W. (1955) Fam. Hebridae Fb. In: Stichel, W. (Ed.) (1955 - 1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa (Hemiptera-Heteroptera Europae). Vol. 1. W. Stichel, Berlin-Hermsdorf, pp. 150 - 155." type="book chapter" year="1955">Stichel 1955</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB2D20CFE7FFCA2A63CE21B" author="Linnavuori" box="[404,600,836,861]" pageId="18" pageNumber="219" refString="Linnavuori, R. E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87 - 95." type="journal article" year="1994">Linnavuori 1994</bibRefCitation>
), antennal segment IVb more than twice as long as IVa (
<bibRefCitation id="EF85EAB2FFB2D20CFB17FCA2A1F4E21A" author="Jordan" box="[1276,1424,836,861]" pageId="18" pageNumber="219" refString="Jordan, K. H. C. (1954) Uber einen neuen Naeogeus aus Sudfrankreich (Heteroptera: Hebridae). Beitrage zur Entomologie, 4, 601 - 604." type="journal article" year="1954">
<collectingCountry id="F303D7D3FFB2D20CFB17FCA2A12CE21A" box="[1276,1352,836,861]" name="Jordan" pageId="18" pageNumber="219">Jordan</collectingCountry>
1954
</bibRefCitation>
), antennal segment I little longer than eye diameter (
<bibRefCitation id="EF85EAB2FFB2D20CFD3AFC81A714E2C7" author="Horvath" box="[721,880,871,896]" pageId="18" pageNumber="219" refString="Horvath, G. (1896) Hemiptera nova palaearctica. Termeszetrajzi Fuzetek, 19, 322 - 329." type="journal article" year="1896">Horváth 1896</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB2D20CFC90FC81A06FE2C7" author="Stichel" box="[891,1035,871,896]" pageId="18" pageNumber="219" refString="Stichel, W. (1955) Fam. Hebridae Fb. In: Stichel, W. (Ed.) (1955 - 1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa (Hemiptera-Heteroptera Europae). Vol. 1. W. Stichel, Berlin-Hermsdorf, pp. 150 - 155." type="book chapter" year="1955">Stichel 1955</bibRefCitation>
), and its small size (
<quantity id="4CEC3AA6FFB2D20CFB04FC81A1F0E2C7" box="[1263,1428,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.66" metricValueMin="1.5" pageId="18" pageNumber="219" unit="mm" value="1.58" valueMax="1.66" valueMin="1.5">1.501.66 mm</quantity>
) suggest placement of
<taxonomicName id="4C14ECC0FFB2D20CFE7DFC6BA67DE2E3" box="[406,537,909,932]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B9604B51FFB2D20CFE7DFC6BA67DE2E3" box="[406,537,909,932]" italics="true" pageId="18" pageNumber="219">H. syriacus</emphasis>
</taxonomicName>
in
<taxonomicName id="4C14ECC0FFB2D20CFDADFC6AA6A6E2E2" box="[582,706,908,933]" class="Insecta" family="Hebridae" genus="Hebrusella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9604B51FFB2D20CFDADFC6AA6A6E2E2" box="[582,706,908,933]" italics="true" pageId="18" pageNumber="219">Hebrusella</emphasis>
</taxonomicName>
. (Body length of Euro-Mediterranean
<taxonomicName id="4C14ECC0FFB2D20CFB6CFC6AA160E2E2" box="[1159,1284,908,933]" class="Insecta" family="Hebridae" genus="Hebrusella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9604B51FFB2D20CFB6CFC6AA160E2E2" box="[1159,1284,908,933]" italics="true" pageId="18" pageNumber="219">Hebrusella</emphasis>
</taxonomicName>
is 1.301.80 (exceptionally 1.93) mm, in
<taxonomicName id="4C14ECC0FFB2D20CFE03FC49A65EE28F" box="[488,570,943,968]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9604B51FFB2D20CFE03FC49A65EE28F" box="[488,570,943,968]" italics="true" pageId="18" pageNumber="219">Hebrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F62AFFB2D20CFDACFC56A6E2E280" box="[583,646,944,967]" pageId="18" pageNumber="219" sensu="stricto">s. str.</taxonomicNameLabel>
&gt; 1.70 (usually 1.80) mm—see
<bibRefCitation id="EF85EAB2FFB2D20CFBE6FC49A0CEE28F" author="Poisson" box="[1037,1194,943,968]" pageId="18" pageNumber="219" refString="Poisson, R. (1944) Contribution a la connaissance des especes africaines du genre Hebrus Curtis 1833 [Hemiptera Gymnocerata] (Missions de Ch. Alluaud et R. Jeannel en Afrique orientale). Revue Francaise d'Entomologie, Nouvelle Serie, 10 (3 - 4) [1943], 89 - 112." type="journal article" year="1944">Poisson 1944</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB2D20CFB51FC49A091E28F" author="Poisson" box="[1210,1269,943,968]" pageId="18" pageNumber="219" refString="Poisson, R. (1953) Les Hydrocorises du Tassili des Ajjer (Sahara central). In: Bernard, F. (Ed.), Mission scientifique au Tassili des Ajjers (1949). I. Recherches Zoologiques et Medicales. Institut des Recherches sahariennes de l'Universite d'Alger, Alger, pp. 65 - 85." type="journal article" year="1953">1953</bibRefCitation>
;
<bibRefCitation id="EF85EAB2FFB2D20CFAEDFC49A1F3E28F" author="Jordan" box="[1286,1431,943,968]" pageId="18" pageNumber="219" refString="Jordan, K. H. C. (1954) Uber einen neuen Naeogeus aus Sudfrankreich (Heteroptera: Hebridae). Beitrage zur Entomologie, 4, 601 - 604." type="journal article" year="1954">
<collectingCountry id="F303D7D3FFB2D20CFAEDFC49A135E28F" box="[1286,1361,943,968]" name="Jordan" pageId="18" pageNumber="219">Jordan</collectingCountry>
1954
</bibRefCitation>
;
<bibRefCitation id="EF85EAB2FFB2D20CFF7CFC33A54AE2AA" author="Wagner" box="[151,302,980,1005]" pageId="18" pageNumber="219" refString="Wagner, E. (1954) Une espece d' Hebrus du midi de la France jusqu'alors meconnu (Hemiptera Heteroptera, Hebridae). Cahiers des Naturalistes (Nouvelle Serie), 9, 67 - 69." type="journal article" year="1954">Wagner 1954</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB2D20CFED1FC32A511E2AA" author="Wagner" box="[314,373,980,1005]" pageId="18" pageNumber="219" refString="Wagner, E. (1957) Neogaeus (= Hebrus) franzi n. sp. Memorie della Societa Entomologica Italiana, 36, 201 - 202." type="journal article" year="1957">1957</bibRefCitation>
;
<bibRefCitation id="EF85EAB2FFB2D20CFE6AFC32A627E2AB" author="Kanyukova" box="[385,579,980,1005]" pageId="18" pageNumber="219" refString="Kanyukova, E. V. (1997) Hebridae of Russia and adjacent countries (Heteroptera). Zoosystematica Rossica, 6, 223 - 236." type="journal article" year="1997">Kanyukova 1997</bibRefCitation>
,
<bibRefCitation id="EF85EAB2FFB2D20CFDA4FC32A6EFE2AA" author="Kanyukova" box="[591,651,980,1005]" pageId="18" pageNumber="219" refString="Kanyukova, E. V. (2006) Vodnye poluzhestkokrylye nasekomye (Heteroptera: Nepomorpha, Gerromorpha) fauny Rossii i sopredelnykh stran. [Aquatic and semiaquatic (Heteroptera: Nepomorpha, Gerromorpha) of the fauna of Russia and neighbouring countries]. Dal'nauka, Vladivostok, 296 pp. [in Russian, English abstract]" type="book" year="2006">2006</bibRefCitation>
)). On the other hand, examination of the antenna by scanning electron microscope revealed that segment IV is subdivided only by a membranous joint (
<figureCitation id="132F8BC6FFB2D20CFBDBFC11A0DDE557" box="[1072,1209,1015,1040]" captionStart="FIGURES 29 35" captionStartId="16.[151,264,1747,1770]" captionTargetBox="[166,1417,411,1723]" captionTargetId="figure@16.[166,1418,411,1723]" captionTargetPageId="16" captionText="FIGURES 29 35. Hebrus syriacus Horváth, 1896, female, Cyprus: Kyrenis, scanning electron micrographs: 29 — body, dorsal view (magnification: 65 ×); 30 — body, lateral view (70 ×); 31 — head, dorsal view (180 ×); 32 — head, lateral view (200 ×); 33 — antenna, dorsal view (170 ×); 34 — segments IVa and IVb (350 ×); 35 — detail of membranous joint between segments IVa and IVb (1300 ×). Abbreviations: IVa, IVb—segments IVa and IVb; mj—membranous junction between segments IVa and IVb. Scale bars: 29 30 — 0.5 mm; 31 33 — 0.2 mm; 34 — 0.1 mm; 35 — 0.02 mm." httpUri="https://zenodo.org/record/261270/files/figure.png" pageId="18" pageNumber="219">Figs. 3435</figureCitation>
), which supports its placement in
<taxonomicName id="4C14ECC0FFB2D20CFEC4FBFAA5E6E572" box="[303,386,1052,1077]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9604B51FFB2D20CFEC4FBFAA5E6E572" box="[303,386,1052,1077]" italics="true" pageId="18" pageNumber="219">Hebrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253F62AFFB2D20CFE62FBFBA5A0E573" box="[393,452,1053,1076]" pageId="18" pageNumber="219" sensu="stricto">s. str.</taxonomicNameLabel>
sensu
<bibRefCitation id="EF85EAB2FFB2D20CFDFAFBFAA6D9E573" author="Poisson" box="[529,701,1052,1077]" pageId="18" pageNumber="219" refString="Poisson, R. (1944) Contribution a la connaissance des especes africaines du genre Hebrus Curtis 1833 [Hemiptera Gymnocerata] (Missions de Ch. Alluaud et R. Jeannel en Afrique orientale). Revue Francaise d'Entomologie, Nouvelle Serie, 10 (3 - 4) [1943], 89 - 112." type="journal article" year="1944">Poisson (1944)</bibRefCitation>
. However, the subgeneric classification of
<taxonomicName id="4C14ECC0FFB2D20CFB49FBFAA090E572" box="[1186,1268,1052,1077]" class="Insecta" family="Hebridae" genus="Hebrus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9604B51FFB2D20CFB49FBFAA090E572" box="[1186,1268,1052,1077]" italics="true" pageId="18" pageNumber="219">Hebrus</emphasis>
</taxonomicName>
is problematic and requires revision (see Summary).
</paragraph>
</subSubSection>
</treatment>
</document>