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<document id="47B5C77161C12ED0A9A1E43930C4130F" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C19FF8CFF3051A264F927E7" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena (Hydraenopsis) palawanensis Freitag &amp; Jach, sp. n." docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="26" updateTime="1698221773297" updateUser="plazi">
<mods:mods id="A2601A2A1AA4E1F9D57D54D7AEEF1542" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2C89259C04B9C563CE88D026D3DDCB70">
<mods:title id="EE5040961DDD230EBF6C3479DCCBA5E0">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
</mods:titleInfo>
<mods:name id="296342F05DFDC4F3D468D81B0671D7B3" type="personal">
<mods:role id="418223C65B360D1878DDB040D8268152">
<mods:roleTerm id="B3E81C11A5A692615BD7D46ED2C94F26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E916BED8C6EDF0B2228E52C903FC4339">Freitag, Hendrik</mods:namePart>
</mods:name>
<mods:name id="FD7390A6A482ED42D50EA5FAC588B6E8" type="personal">
<mods:role id="D6BF87FEFF3A8A489E91C19F5FDC7FC9">
<mods:roleTerm id="FD5F88F0EDBE2769703196725E0E8958">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9DA5E187ACB5674720123AB20CB232A">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E82FE99F03739E63F3B2A731A517B9A4">text</mods:typeOfResource>
<mods:relatedItem id="CBD4490B8CB700C2BB4E42AA82D9AD7F" type="host">
<mods:titleInfo id="F07A6EF10B37B7C725BD778DDE85848C">
<mods:title id="CCEF45E182DA4DDD8A1030A26867BC92">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6EBBC813F792E171B5C2ABB58B631F31">
<mods:date id="EE5197AE63B3221C257E7A5C9C26941D">2007</mods:date>
<mods:detail id="CCEB8BC34B7856A6D30DFD0B311E1521" type="volume">
<mods:number id="A66D563410AE856C0E722FE8CAC395C2">1431</mods:number>
</mods:detail>
<mods:extent id="D2F0DF3CDCA7AB14DD735D099AEC6672" unit="page">
<mods:start id="ACBB334BD1E0C5B0C9FA4C1D8950D99D">1</mods:start>
<mods:end id="30E4E85F3A055CB9DD17D3A28A6CB710">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="99CAB017903D1872C18D4423FD5D587E">journal article</mods:classification>
<mods:identifier id="642F64289AEAEDADAE3C02300C7A2025" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="E8A13AF343405D01C5C3C7FCFBBD58CC" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="5470FA5C7C6546D50E3173DEE73174AD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5674EF2A5703DFF500ACFD16B30D0EFE" type="Zenodo-Dep">175790</mods:identifier>
</mods:mods>
<treatment id="03B9F24A7C19FF8CFF3051A264F927E7" ID-DOI="http://doi.org/10.5281/zenodo.6248085" ID-GBIF-Taxon="119355657" ID-Zenodo-Dep="6248085" LSID="urn:lsid:plazi:treatment:03B9F24A7C19FF8CFF3051A264F927E7" httpUri="http://treatment.plazi.org/id/03B9F24A7C19FF8CFF3051A264F927E7" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="C30A10D77C19FF89FF3051A265432391" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BAF435C7C19FF89FF3051A2679323F4" blockId="25.[151,911,1431,1492]" box="[151,911,1431,1457]" pageId="25" pageNumber="26">
<heading id="D0E7F4307C19FF89FF3051A2679323F4" bold="true" box="[151,911,1431,1457]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B9649F4E7C19FF89FF3051A2679323F4" bold="true" box="[151,911,1431,1457]" pageId="25" pageNumber="26">
<taxonomicName id="4C1038DF7C19FF89FF3051A2672323F4" ID-CoL="6MJBR" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,831,1431,1457]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9649F4E7C19FF89FF3051A2650C23F4" bold="true" box="[151,272,1431,1457]" italics="true" pageId="25" pageNumber="26">Hydraena</emphasis>
(
<emphasis id="B9649F4E7C19FF89FE8751A265DA23F4" bold="true" box="[288,454,1431,1457]" italics="true" pageId="25" pageNumber="26">Hydraenopsis</emphasis>
)
<emphasis id="B9649F4E7C19FF89FE7051A2666623F4" bold="true" box="[471,634,1431,1457]" italics="true" pageId="25" pageNumber="26">palawanensis</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<taxonomicNameLabel id="A25722357C19FF89FCEB51A2679323F4" box="[844,911,1431,1457]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF435C7C19FF89FF30518F65432391" blockId="25.[151,911,1431,1492]" box="[151,351,1466,1492]" pageId="25" pageNumber="26">(Figs. 10, 22a–h)</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C19FF8AFF30523F67A12577" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BAF435C7C19FF89FF30523F657E2009" blockId="25.[151,1437,1546,2012]" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF30523F65222061" bold="true" box="[151,318,1546,1572]" pageId="25" pageNumber="26">
<typeStatus id="54ABFDFE7C19FF89FF30523F64C92061" box="[151,213,1546,1572]" pageId="25" pageNumber="26">Type</typeStatus>
locality.
</emphasis>
Residual river pools, Mate, Municipality of Brooke’s Point,
<geoCoordinate id="EE24259B7C19FF89FBA8523F606C2061" box="[1039,1136,1546,1572]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="8.766666">08°46'N</geoCoordinate>
<geoCoordinate id="EE24259B7C19FF89FBDF523F60FA2061" box="[1144,1254,1546,1572]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="117.833336">117°50'E</geoCoordinate>
, southern Palawan,
<collectingCountry id="F30703CC7C19FF89FF70520765432009" box="[215,351,1586,1612]" name="Philippines" pageId="25" pageNumber="26">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BAF435C7C19FF8AFF60526F67A12577" blockId="25.[151,1437,1546,2012]" lastBlockId="26.[151,1437,151,2018]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF60526F65EB2031" bold="true" box="[199,503,1626,1652]" pageId="25" pageNumber="26">
<typeStatus id="54ABFDFE7C19FF89FF60526F65182031" box="[199,260,1626,1652]" pageId="25" pageNumber="26">Type</typeStatus>
material.
<typeStatus id="54ABFDFE7C19FF89FE22526F65EB2031" box="[389,503,1626,1652]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED01DB997C19FF89FD85526F66692031" box="[546,629,1626,1652]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C19FF89FD34526F66C22031" box="[659,734,1626,1652]" pageId="25" pageNumber="26">PHIL</collectionCode>
.: Palawan Brooke’s Point Mate,
<date id="FFAE659C7C19FF89FBFC526F60CC2031" box="[1115,1232,1626,1652]" pageId="25" pageNumber="26" value="1994-03-31">31.3.1994</date>
leg. Zettel (54)”, terminal parts of abdomen and aedeagus glued separately.
<emphasis id="B9649F4E7C19FF89FCE752B767DB20D9" bold="true" box="[832,967,1666,1692]" pageId="25" pageNumber="26">
<typeStatus id="54ABFDFE7C19FF89FCE752B767DE20D9" box="[832,962,1666,1692]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
:
</emphasis>
44 ɗɗ (37 exs.
<collectionCode id="ED01DB997C19FF89FB2452B760D320D9" box="[1155,1231,1666,1692]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
, 3 exs. mutilated; 2 exs.
<collectionCode id="ED01DB997C19FF89FF42529F652E2081" box="[229,306,1706,1732]" pageId="25" pageNumber="26">IRML</collectionCode>
; 2 ex.
<collectionCode id="ED01DB997C19FF89FE24529F65CE2081" box="[387,466,1706,1732]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="25" pageNumber="26">UPLB</collectionCode>
; 2 ex. WPU; 1 ex. CZW), 23 ΨΨ (21 exs.
<collectionCode id="ED01DB997C19FF89FC77529F60022081" box="[976,1054,1706,1732]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
; 1ex.
<collectionCode id="ED01DB997C19FF89FBC2529F60A82081" box="[1125,1204,1706,1732]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="25" pageNumber="26">UPLB</collectionCode>
; 1 ex. WPU): same locality data as
<typeStatus id="54ABFDFE7C19FF89FEEB52E765AA20A9" box="[332,438,1746,1772]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
; 13 ɗɗ (
<collectionCode id="ED01DB997C19FF89FDBB52E7667520A9" box="[540,617,1746,1772]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
, 2 exs. mutilated, aedeagus of 1 ex. lacking), 9 ΨΨ (
<collectionCode id="ED01DB997C19FF89FB6252E7610E20A9" box="[1221,1298,1746,1772]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
, 1 ex. mutilated): “
<collectionCode id="ED01DB997C19FF89FF5E52CF65582151" box="[249,324,1786,1812]" pageId="25" pageNumber="26">PHIL</collectionCode>
.: Palawan, Taytay; Lake
<taxonomicName id="4C1038DF7C19FF89FDDE52CF66FB2151" box="[633,743,1786,1812]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
tributary, Alipuran Stream, semi-prim. forest c.
<quantity id="4CE8EEB97C19FF89FA9752CF61742151" box="[1328,1384,1786,1812]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="m" value="40.0">40m</quantity>
asl,
<geoCoordinate id="EE24259B7C19FF89FF30531764E42179" box="[151,248,1826,1852]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="10.75">10°45'N</geoCoordinate>
<geoCoordinate id="EE24259B7C19FF89FEA7531765772179" box="[256,363,1826,1852]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="119.53333">
119°32'
<geoCoordinate id="EE24259B7C19FF89FEFF531765772179" box="[344,363,1826,1852]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="5555" value="29.4">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C19FF89FED3531765F12179" box="[372,493,1826,1852]" pageId="25" pageNumber="26" value="1995-04-29">29.4.1995</date>
, leg. Freitag (15)M”; 6 ɗɗ (
<collectionCode id="ED01DB997C19FF89FCE7531767932179" box="[832,911,1826,1852]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
; 1 ex.
<collectionCode id="ED01DB997C19FF89FC47531760322179" box="[992,1070,1826,1852]" pageId="25" pageNumber="26">IRML</collectionCode>
), 5 ΨΨ (
<collectionCode id="ED01DB997C19FF89FB34531760F92179" box="[1171,1253,1826,1852]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C19FF89FAA3531761532179" box="[1284,1359,1826,1852]" pageId="25" pageNumber="26">PHIL</collectionCode>
.: Palawan, P.Princesa; S. Vincente Taranaban Riv;
<quantity id="4CE8EEB97C19FF89FD0E537F66EF2121" box="[681,755,1866,1892]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="km" value="0.5">0.5km</quantity>
N of Highway, mountain riv. resid. pool,
<quantity id="4CE8EEB97C19FF89FB7F537F61172121" box="[1240,1291,1866,1892]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="10.0">10m</quantity>
asl
<geoCoordinate id="EE24259B7C19FF89FA9B537F61802121" box="[1340,1436,1866,1892]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="10.016666">10°01'N</geoCoordinate>
<geoCoordinate id="EE24259B7C19FF89FF305347651821C9" box="[151,260,1906,1932]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="119.01667">119°01'E</geoCoordinate>
,
<date id="FFAE659C7C19FF89FEB05347659021C9" box="[279,396,1906,1932]" pageId="25" pageNumber="26" value="1995-04-30">30.4.1995</date>
leg. Freitag (16)M”; 1 ɗ (
<collectionCode id="ED01DB997C19FF89FD405347672521C9" box="[743,825,1906,1932]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C19FF89FCF9534767B521C9" box="[862,937,1906,1932]" pageId="25" pageNumber="26">PHIL</collectionCode>
.: Palawan, P.Princesa; Luzvimin-da,Villa Bukit, garden/farmland creek
<quantity id="4CE8EEB97C19FF89FE5B53AF662C21F1" box="[508,560,1946,1972]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="km" value="8.0">8km</quantity>
SSW PPC;
<quantity id="4CE8EEB97C19FF89FD6353AF66E421F1" box="[708,760,1946,1972]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="20.0">20m</quantity>
asl
<geoCoordinate id="EE24259B7C19FF89FC8C53AF676121F1" box="[811,893,1946,1972]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="9.666667">9°40'N</geoCoordinate>
<geoCoordinate id="EE24259B7C19FF89FC2153AF67E821F1" box="[902,1012,1946,1972]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="118.71667">118°43'E</geoCoordinate>
,
<date id="FFAE659C7C19FF89FBA553AF606B21F1" box="[1026,1143,1946,1972]" pageId="25" pageNumber="26" value="1995-03-25">25.3.1995</date>
leg. Freitag(21)M”; 1 ɗ (
<collectionCode id="ED01DB997C19FF89FF3853F764EE2199" box="[159,242,1986,2012]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C19FF89FEB453F765422199" box="[275,350,1986,2012]" pageId="25" pageNumber="26">PHIL</collectionCode>
.: Palawan, P.Princesa; Luzviminda, forest creek, small fall,
<quantity id="4CE8EEB97C19FF89FB9453F760742199" box="[1075,1128,1986,2012]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="25" pageNumber="26" unit="km" value="9.0">9km</quantity>
SSW PPC; sandy banks, pool, c.
<quantity id="4CE8EEB97C1AFF8AFF4C54AD653F26F7" box="[235,291,152,178]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="50.0">50m</quantity>
asl
<geoCoordinate id="EE24259B7C1AFF8AFEF354AD65BB26F7" box="[340,423,152,178]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="9.65">9°39'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFE0854AD660B26F7" box="[431,535,152,178]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="118.63333">118°38'E</geoCoordinate>
,
<date id="FFAE659C7C1AFF8AFDBB54AD66B626F7" box="[540,682,152,178]" pageId="26" pageNumber="27" value="1994-12-14">14.12.1994</date>
,leg.Freitag(17b)M”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFC7B54AD603226F7" box="[988,1070,152,178]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFBE954AD608526F7" box="[1102,1177,152,178]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P.Princesa; Luzviminda, forest creek, small fall,
<quantity id="4CE8EEB97C1AFF8AFD3654F566D9269F" box="[657,709,192,218]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="27" unit="km" value="9.0">9km</quantity>
SSW PPC; side pools w/ leaves, c.
<quantity id="4CE8EEB97C1AFF8AFB6254F560E1269F" box="[1221,1277,192,218]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="50.0">50m</quantity>
asl
<geoCoordinate id="EE24259B7C1AFF8AFAED54F56180269F" box="[1354,1436,192,218]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="9.65">9°39'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFF3054DD64F32747" box="[151,239,232,258]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="18.633333">18°38'E</geoCoordinate>
,
<date id="FFAE659C7C1AFF8AFF5354DD65982747" box="[244,388,232,258]" pageId="26" pageNumber="27" value="1994-12-14">14.12.1994</date>
,leg.Freitag(17c)M”; 2 ɗɗ (
<collectionCode id="ED01DB997C1AFF8AFD7354DD673D2747" box="[724,801,232,258]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
, 1 ex. mutilated): “
<collectionCode id="ED01DB997C1AFF8AFBB354DD60432747" box="[1044,1119,232,258]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa SE Manturon, Cabayugan R.
<geoCoordinate id="EE24259B7C1AFF8AFE6E55256641276F" box="[457,605,272,298]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="10.1581">10°09.486'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFDC155256718276F" box="[614,772,272,298]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="118.89155">
118°53.493'
<geoCoordinate id="EE24259B7C1AFF8AFD5655256718276F" box="[753,772,272,298]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="3.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C1AFF8AFCAB55256769276F" box="[780,885,272,298]" pageId="26" pageNumber="27" value="2001-03-03">3.3.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFBBD55256028276F" box="[1050,1076,272,298]" pageId="26" pageNumber="27">i4</date>
)”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFB34552560F9276F" box="[1171,1253,272,298]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFAA25525614C276F" box="[1285,1360,272,298]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa S Manturon, Cabayugan R.
<geoCoordinate id="EE24259B7C1AFF8AFD18550D674F2717" box="[703,851,312,338]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="10.154317">10°09.259'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFCF9550D67E12717" box="[862,1021,312,338]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="118.87505">
118°52.503'
<geoCoordinate id="EE24259B7C1AFF8AFC4D550D67E12717" box="[1002,1021,312,338]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="31.7">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C1AFF8AFBAF550D609C2717" box="[1032,1152,312,338]" pageId="26" pageNumber="27" value="2001-07-31">31.7.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFA89550D615A2717" box="[1326,1350,312,338]" pageId="26" pageNumber="27">i3</date>
)”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFF38555564EE273F" box="[159,242,352,378]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFEA855556546273F" box="[271,346,352,378]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa SE Manturon, Karst spring LS4R
<geoCoordinate id="EE24259B7C1AFF8AFC4D5555606E273F" box="[1002,1138,352,378]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.158055">10°09'29''N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFBDE55556110273F" box="[1145,1292,352,378]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="118.89166">118°53'30''E</geoCoordinate>
<date id="FFAE659C7C1AFF8AFAB45555618B273F" box="[1299,1431,352,378]" pageId="26" pageNumber="27" value="2001-04-21">21.IV.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFE8C55BD654F27E7" box="[299,339,392,418]" pageId="26" pageNumber="27" value="1934-01">i34</date>
) E”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFE6F55BD660727E7" box="[456,539,392,418]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFD9F55BD669F27E7" box="[568,643,392,418]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa SE Manturon, Karst spring LS4R
<geoCoordinate id="EE24259B7C1AFF8AFAB455BD618027E7" box="[1299,1436,392,418]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.158055">10°09'29''N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFF3055856531278F" box="[151,301,432,458]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="118.89166">118°53'30''E</geoCoordinate>
;
<date id="FFAE659C7C1AFF8AFE9A558565DD278F" box="[317,449,432,458]" pageId="26" pageNumber="27" value="2001-04-21">21.IV.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFDC85585668A278F" box="[623,662,432,458]" pageId="26" pageNumber="27" value="1935-01">i35</date>
) E”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFCBA55856773278F" box="[797,879,432,458]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFC35558567C1278F" box="[914,989,432,458]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa SE Manturon, Karst spring LS4R
<geoCoordinate id="EE24259B7C1AFF8AFEDE55ED661D27B7" box="[377,513,472,498]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.158055">10°09'29''N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFDAE55ED668727B7" box="[521,667,472,498]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="118.89166">118°53'30''E</geoCoordinate>
<date id="FFAE659C7C1AFF8AFD0455ED673527B7" box="[675,809,472,498]" pageId="26" pageNumber="27" value="2001-03-18">18.III.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFC6D55ED67ED27B7" box="[970,1009,472,498]" pageId="26" pageNumber="27" value="1951-01">i51</date>
) E”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFBC055ED60A527B7" box="[1127,1209,472,498]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFB7F55ED613F27B7" box="[1240,1315,472,498]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa SE Manturon, Karst spring LS4R
<geoCoordinate id="EE24259B7C1AFF8AFD1456356727245F" box="[691,827,512,538]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.158055">10°09'29''N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFCE3563567CA245F" box="[836,982,512,538]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="118.89166">118°53'30''E</geoCoordinate>
<date id="FFAE659C7C1AFF8AFC7856356079245F" box="[991,1125,512,538]" pageId="26" pageNumber="27" value="2001-03-18">18.III.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFAAD5635612E245F" box="[1290,1330,512,538]" pageId="26" pageNumber="27" value="1952-01">i52</date>
) E”; 1 ɗ (WPU): “
<collectionCode id="ED01DB997C1AFF8AFEAF561D654F2407" box="[264,339,552,578]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge
<geoCoordinate id="EE24259B7C1AFF8AFB60561D61342407" box="[1223,1320,552,578]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="10.166667">10°10'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFA96561D61802407" box="[1329,1436,552,578]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="118.88333">
118°53'
<geoCoordinate id="EE24259B7C1AFF8AFA2F561D61802407" box="[1416,1436,552,578]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="27.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C1AFF8AFF3056656513242F" box="[151,271,592,618]" pageId="26" pageNumber="27" value="2001-03-27">27.3.2001</date>
, leg. Freitag (5)M”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFDE056656685242F" box="[583,665,592,618]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="26" pageNumber="27">UPLB</collectionCode>
): “
<collectionCode id="ED01DB997C1AFF8AFD1E56656718242F" box="[697,772,592,618]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge
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118°53'
<geoCoordinate id="EE24259B7C1AFF8AFD8D564D662224D7" box="[554,574,632,658]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="27.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C1AFF8AFDE2564D66A124D7" box="[581,701,632,658]" pageId="26" pageNumber="27" value="2001-03-27">27.3.2001</date>
, leg. Freitag (2)M”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFC4D564D602B24D7" box="[1002,1079,632,658]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
, mutilated): “
<collectionCode id="ED01DB997C1AFF8AFB70564D613E24D7" box="[1239,1314,632,658]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge
<geoCoordinate id="EE24259B7C1AFF8AFC045695601F24FF" box="[931,1027,672,698]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="10.166667">10°10'N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFBB75695606724FF" box="[1040,1147,672,698]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="118.88333">
118°53'
<geoCoordinate id="EE24259B7C1AFF8AFBC05695606724FF" box="[1127,1147,672,698]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5555" value="27.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C1AFF8AFB2F5695611C24FF" box="[1160,1280,672,698]" pageId="26" pageNumber="27" value="2001-03-27">27.3.2001</date>
, leg. Freitag (3)M”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFE9456FD659C24A7" box="[307,384,712,738]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
, mutilated): “
<collectionCode id="ED01DB997C1AFF8AFD8156FD666D24A7" box="[550,625,712,738]" pageId="26" pageNumber="27">PHIL</collectionCode>
.: Palawan, P. Princesa S Manturon, Cabayugan R.CR4
<geoCoordinate id="EE24259B7C1AFF8AFAB456FD618024A7" box="[1299,1436,712,738]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="10.157778">10°09'28''N</geoCoordinate>
<geoCoordinate id="EE24259B7C1AFF8AFF3056C56535254F" box="[151,297,752,778]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="118.89056">118°53'26''E</geoCoordinate>
<date id="FFAE659C7C1AFF8AFE9456C565A6254F" box="[307,442,752,778]" pageId="26" pageNumber="27" value="2001-03-18">18.III.2001</date>
, leg. Freitag (
<date id="FFAE659C7C1AFF8AFDC456C56697254F" box="[611,651,752,778]" pageId="26" pageNumber="27" value="1941-01">i41</date>
)E”; 1 ɗ (
<collectionCode id="ED01DB997C1AFF8AFCA656C5674E254F" box="[769,850,752,778]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
), 5 ΨΨ (
<collectionCode id="ED01DB997C1AFF8AFC1F56C56019254F" box="[952,1029,752,778]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
, 1 ex. mutilated): “
<collectingCountry id="F30703CC7C1AFF8AFB4C56C56187254F" box="[1259,1435,752,778]" name="Philippines" pageId="26" pageNumber="27">PHILIPPINES</collectingCountry>
Sabang, N. of Mt. St. Paul, Palawan July, 11–13, 1977 M. Satô leg.”.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C1AFF8BFF60570A660521C0" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BAF435C7C1AFF8AFF60570A670625EF" blockId="26.[151,1437,151,2018]" pageId="26" pageNumber="27">
<emphasis id="B9649F4E7C1AFF8AFF60570A6543251C" bold="true" box="[199,351,831,857]" pageId="26" pageNumber="27">Description.</emphasis>
Body
<quantity id="4CE8EEB97C1AFF8AFE0957756646251F" box="[430,602,832,858]" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="26" pageNumber="27" unit="mm" value="1.175" valueMax="1.25" valueMin="1.1">1.10–1.25 mm</quantity>
long. Habitus as in Fig. 10. Elytra brown; rectangular spot in the middle of pronotal disc and frons (particularly around eyes) darker brown; pronotal margins, legs, maxillary palpi and sometimes posterior third of elytra distinctly paler.
</paragraph>
<paragraph id="8BAF435C7C1AFF8AFF60578D66A122DF" blockId="26.[151,1437,151,2018]" pageId="26" pageNumber="27">Pronotum wider than long, broadest at the middle; entire pronotum moderately densely punctate, sometimes appearing bristled; punctures very small and shallowly impressed, interstices glabrous; foveae slightly impressed or inconspicuous; anterior margin slightly concave; anterior angles rounded; lateral rim inconspicuously denticulate; lateral margins anteriorly convergent, posteriorly sinuously convergent; pronoto-elytral angle obtuse; posterior margin slightly biconcave; lateral hypomeron posteriorly slightly broader than profemur; mesal hypomeron narrow, inconspicuous.</paragraph>
<paragraph id="8BAF435C7C1AFF8AFF60509D652C23CF" blockId="26.[151,1437,151,2018]" pageId="26" pageNumber="27">Elytra elongately oval, usually not conjointly rounded apically, with inconspicuously small acute notch, with ca. 14 longitudinal, more or less regular, inconspicuous rows of punctures (approximately eight between suture and shoulder); punctures very small and very shallowly impressed; interstices and intervals almost flat, almost glabrous; lateral portion narrow, reaching ca. posterior 0.3; inflexed lateral portion moderately wide (ca. as broad as metafemur), very short (reaching ca. posterior 0.3); pseudepipleuron without conspicuous row of punctures.</paragraph>
<paragraph id="8BAF435C7C1AFF8AFF6051AD66A1203F" blockId="26.[151,1437,151,2018]" pageId="26" pageNumber="27">Mesoventrite very deeply impressed, with well developed longitudinal ridges; mesoventral intercoxal process very wide and long, broader than pseudepipleuron, reaching beyond posterior margin of mesocoxa, firmly connected with metaventrite. Metaventral disc slightly impressed; metaventral plaques moderately widely separated (as wide as inflexed lateral portion of elytron), slightly convergent anteriorly, moderately narrow, glabrous; intercoxal sternite moderately large, approximately as broad as metafemur; metaventrite and abdominal sternites moderately pubescent.</paragraph>
<paragraph id="8BAF435C7C1AFF8AFF6052BD67C420E7" blockId="26.[151,1437,151,2018]" box="[199,984,1672,1698]" pageId="26" pageNumber="27">
Femora almost as in
<taxonomicName id="4C1038DF7C1AFF8AFE1C52BD66A220E4" box="[443,702,1672,1697]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="nielshaggei">
<emphasis id="B9649F4E7C1AFF8AFE1C52BD66A220E4" box="[443,702,1672,1697]" italics="true" pageId="26" pageNumber="27">Hydraena nielshaggei</emphasis>
</taxonomicName>
. Protibiae slightly bent.
</paragraph>
<paragraph id="8BAF435C7C1AFF8AFF605285666E21A7" blockId="26.[151,1437,151,2018]" pageId="26" pageNumber="27">
Aedeagus (
<figureCitation id="132B5FD97C1AFF8AFEEC528565B7208F" box="[331,427,1712,1738]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="26" pageNumber="27">Figs. 22</figureCitation>
a–c) basally slender, strongly widening in apical 0.7 (dorsal/ventral view). Main piece without setae (five micropores present), right margin with distinct emargination near insertion of right paramere, apex sinuous and slender; phallobase almost symmetric. Distal lobe intricately shaped, ventrally with strongly sclerotized, bisinuous structure, medioapically with tapered appendix. Parameres articulately connected with main piece; left paramere very small, inserted ventro-lateral, at about basal 0.2 of aedeagus, apically with five short setae; right paramere distinctly longer and broader, inserted at about basal 0.25 of aedeagus, less sclerotized, with inconspicuous tiny apical bristles and with irregular ventro-median longitudinal row of ca. six moderately long setae.
</paragraph>
<caption id="DF6F13D47C1BFF8BFF3050906799229A" httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="27" pageNumber="28" targetBox="[269,1321,197,1154]" targetPageId="27">
<paragraph id="8BAF435C7C1BFF8BFF3050906799229A" blockId="27.[151,1436,1189,1247]" pageId="27" pageNumber="28">
<emphasis id="B9649F4E7C1BFF8BFF305090653A22F8" bold="true" box="[151,294,1189,1213]" pageId="27" pageNumber="28">FIGURE 22.</emphasis>
<taxonomicName id="4C1038DF7C1BFF8BFE8B5093663222F8" box="[300,558,1190,1213]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B9649F4E7C1BFF8BFE8B5093663222F8" box="[300,558,1190,1213]" italics="true" pageId="27" pageNumber="28">Hydraena palawanensis</emphasis>
</taxonomicName>
<emphasis id="B9649F4E7C1BFF8BFD935090666C22F8" bold="true" box="[564,624,1189,1213]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A25722357C1BFF8BFD935090666C22F8" box="[564,624,1189,1213]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a–c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f–g) spermatheca; h) female tergite X.
</paragraph>
</caption>
<paragraph id="8BAF435C7C1BFF8BFF60512665AD23E0" blockId="27.[151,1437,1299,2005]" pageId="27" pageNumber="28">
Gonocoxite (
<figureCitation id="132B5FD97C1BFF8BFEC5512665A62368" box="[354,442,1299,1325]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
e) subtriangular; apical area of ventral plate very short; margin between apical and basal area not clearly defined; setae of subapical tufts almost straight; basal area of ventral plate sparsely setose apically; condyles moderately large, distinctly retracted; dorsal plate not surpassing outer plate, with one moderately large cavity.
</paragraph>
<paragraph id="8BAF435C7C1BFF8BFF605186654723B0" blockId="27.[151,1437,1299,2005]" pageId="27" pageNumber="28">Spermatheca (Figs. 8f–g). Proximal portion crescentic, with a pair of tiny appendages; distal portion more or less discoidal.</paragraph>
<paragraph id="8BAF435C7C1BFF8BFF60523667D620D0" blockId="27.[151,1437,1299,2005]" pageId="27" pageNumber="28">
Secondary sexual characters. Basoventral corner of male femur more acute than in female (dissection of leg recommended). Male terminal sternite pyriform (
<figureCitation id="132B5FD97C1BFF8BFCBB521E67642000" box="[796,888,1579,1605]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
d). Male tergite X distinctly excised apically. Female tergite X (
<figureCitation id="132B5FD97C1BFF8BFED7526665DA2028" box="[368,454,1619,1645]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="27" pageNumber="28">Fig. 22</figureCitation>
h) subsemicircular; disc moderately densely setose; subapical fringe setae trichoid, not arranged in regular row; hyaline apical margin with three notches.
</paragraph>
<paragraph id="8BAF435C7C1BFF8BFF605296660521C0" blockId="27.[151,1437,1299,2005]" pageId="27" pageNumber="28">
<emphasis id="B9649F4E7C1BFF8BFF60529665C120F8" bold="true" box="[199,477,1699,1725]" pageId="27" pageNumber="28">Differential diagnosis.</emphasis>
The external habitus of this species is similar to
<taxonomicName id="4C1038DF7C1BFF8BFB9E5296615C20F9" box="[1081,1344,1699,1724]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nielshaggei">
<emphasis id="B9649F4E7C1BFF8BFB9E5296615C20F9" box="[1081,1344,1699,1724]" italics="true" pageId="27" pageNumber="28">Hydraena nielshaggei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1038DF7C1BFF8BFAD85296653C20A1" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jojoorculloi">
<emphasis id="B9649F4E7C1BFF8BFAD85296653C20A1" italics="true" pageId="27" pageNumber="28">H. jojoorculloi</emphasis>
</taxonomicName>
, from which it can be distinguished externally by the subquadrate pronotum with inconspicuous lateral denticulation, the small and shallowly impressed elytral and pronotal punctures, the bent protibiae, and unusual mesoventral process. Furthermore, its male genital characters (aedeagus strongly widened apically, distal lobe intricately shaped, distinctly unequal parameres articulately connected to main piece) allow to distinguish it from all other species.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C1BFF8CFF6053A666A92717" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BAF435C7C1BFF8BFF6053A6619721E8" blockId="27.[151,1437,1299,2005]" box="[199,1419,1939,1965]" pageId="27" pageNumber="28">
<emphasis id="B9649F4E7C1BFF8BFF6053A6657C21E8" bold="true" box="[199,352,1939,1965]" pageId="27" pageNumber="28">Distribution</emphasis>
(
<figureCitation id="132B5FD97C1BFF8BFED753A665DB21E8" box="[368,455,1939,1965]" captionStart="FIGURE 31" captionStartId="42.[151,255,1170,1194]" captionTargetBox="[267,1314,194,1138]" captionTargetId="figure@42.[264,1323,194,1146]" captionTargetPageId="42" captionText="FIGURE 31. Collecting sites and geographical distribution of Hydraena species in Palawan and Busuanga, including enlarged map of St. Paul National Park: Hydraena palawanensis sp. n., H. pseudopalawanensis sp. n., H. spp. ΨΨ cf. palawanensis / pseudopalawanensis, Hydraena sp. 1, H. manguao sp. n., H. hosiwergi sp. n., H. busuanga sp. n., Hydraena sp. 2, Hydraena sp. 3." httpUri="https://zenodo.org/record/175809/files/figure.png" pageId="27" pageNumber="28">Fig. 31</figureCitation>
).
<taxonomicName id="4C1038DF7C1BFF8BFE7B53A666E421E9" box="[476,760,1939,1964]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B9649F4E7C1BFF8BFE7B53A666E421E9" box="[476,760,1939,1964]" italics="true" pageId="27" pageNumber="28">Hydraena palawanensis</emphasis>
</taxonomicName>
was so far collected in central and in southern Palawan.
</paragraph>
<paragraph id="8BAF435C7C1BFF8CFF60538E66A92717" blockId="27.[151,1437,1299,2005]" lastBlockId="28.[151,1437,152,418]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B9649F4E7C1BFF8BFF60538E652C2190" bold="true" box="[199,304,1979,2005]" pageId="27" pageNumber="28">Ecology.</emphasis>
This species is encountered in undisturbed pristine forests (e.g. LS4) and in moderately disturbed rural areas (CR3;
<bibRefCitation id="EF813EAD7C1CFF8CFE6754AD664026F7" author="Freitag" box="[448,604,152,178]" pageId="28" pageNumber="43" refString="Freitag, H. (2005) Longitudinal zonation patterns and determinants in Decapoda (Crustacea) in rivers of Palawan Island, the Philippines. Archiv fur Hydrobiologie, Supplement, 151, 243 - 268." type="journal article" year="2005">Freitag 2005</bibRefCitation>
), which suggests a rather high ecological potency. Most specimens of
<taxonomicName id="4C1038DF7C1CFF8CFF3054F565AB269C" box="[151,439,192,217]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B9649F4E7C1CFF8CFF3054F565AB269C" box="[151,439,192,217]" italics="true" pageId="28" pageNumber="29">Hydraena palawanensis</emphasis>
</taxonomicName>
were collected in residual pools of a small stream, and between CPOM in sandy slack-water zones of small lowland streams. Additional material was retrieved from emergence traps positioned over pool sections of first to fourth order streams. Therefore it is likely that the species inhabits the detritus intersticial and avoids strong currents.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C1CFF8CFF60556A64F927E7" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BAF435C7C1CFF8CFF60556A64F927E7" blockId="28.[151,1437,152,418]" pageId="28" pageNumber="29">
<emphasis id="B9649F4E7C1CFF8CFF60556A654F273C" bold="true" box="[199,339,351,377]" pageId="28" pageNumber="29">Etymology.</emphasis>
This species is named after Palawan, in reference to its comparably wide distribution on this island.
</paragraph>
</subSubSection>
</treatment>
</document>
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