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<document id="DFE651E9E2787703B507B68380AD4877" ID-DOI="10.5252/z2014n1a2" ID-ISSN="1638-9387" ID-Zenodo-Dep="4539480" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613160692788" checkinUser="felipe" docAuthor="Nguyen Duy-Jacquemin, Monique" docDate="2014" docId="03ABB9399120E34EFBDDFB04FEE0589F" docLanguage="en" docName="Zoosystema.36.1.29-39.pdf.imf" docOrigin="Zoosystema 36 (1)" docSource="http://dx.doi.org/10.5252/z2014n1a2" docStyle="DocumentStyle:BCA1AC1CA2CB189AA8340D8E5C61F956.3:Zoosystema.2013-2014.journal_article" docStyleId="BCA1AC1CA2CB189AA8340D8E5C61F956" docStyleName="Zoosystema.2013-2014.journal_article" docStyleVersion="3" docTitle="Lophoturus humphreysi Duy-Jacquemin 2014, n. sp." docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="FF92C1419126E347FB01FF8AFF8D5E3C" masterDocTitle="Two new species of Lophoturus (Diplopoda, Penicillata, Lophoproctidae) from caves in Christmas Island, Australia, including the second troglomorph in Penicillata" masterLastPageNumber="39" masterPageNumber="29" pageNumber="35" updateTime="1698872000494" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="FF3BE760759BC7D528E6B8F48A04BFC6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="119AAC202638DA22A5DA58BFF3F8FFCF">
<mods:title id="B5918065FBB2C9579FDF30C00557CBB3">Two new species of Lophoturus (Diplopoda, Penicillata, Lophoproctidae) from caves in Christmas Island, Australia, including the second troglomorph in Penicillata</mods:title>
</mods:titleInfo>
<mods:name id="729D16460451A866A744A4786274D261" type="personal">
<mods:role id="BD4F9FD08FEE11C4CEB3981923F0DBFF">
<mods:roleTerm id="8D001F3973E77C664081C22E0B40F6EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD52A6C5249D3406582AFA7BCCFBD5CD">Nguyen Duy-Jacquemin, Monique</mods:namePart>
<mods:affiliation id="B51643FFE2308B4A8828FC00AE0C8C68">Muséum national d’Histoire naturelle, Département Systématique et Évolution, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) monguyen @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="B35048638F3D5E9D6F8E2A335E6B1D1B" type="email">monguyen@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="B2721FAC09C846C8C6120BE610EAD54A">text</mods:typeOfResource>
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<mods:title id="91AE9B8BEC4EE77EADC6017F8C286982">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="1E09F636931627F00581FE8807861429">
<mods:date id="E466722DC96B5C7F7DCB0F00863BFBFE">2014</mods:date>
<mods:detail id="19FF21925C95FB5F119150D21ACA226B" type="pubDate">
<mods:number id="C92C618ADEB13918C6EA4FB7B99E6085">2014-03-28</mods:number>
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<mods:detail id="DD82651D1A67C5C6D477A0F2365E87A2" type="volume">
<mods:number id="ACC6D94AA1F42CF3136BA75540364FC6">36</mods:number>
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<mods:number id="50F3BEB6FAD9E0C570759670D2C48692">1</mods:number>
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<mods:end id="AC55D25EF456C6788E43D3C66234C4A3">39</mods:end>
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<mods:location id="398FCAB9E5350A965177307F2DCC7263">
<mods:url id="A3DD58DA0B32644F0599E0870883C6C9">http://dx.doi.org/10.5252/z2014n1a2</mods:url>
</mods:location>
<mods:classification id="0B0CCDB93F992D4ED9B4DE63163D5CBA">journal article</mods:classification>
<mods:identifier id="3AB7B1E483515E2E01B245504BAE1698" type="DOI">10.5252/z2014n1a2</mods:identifier>
<mods:identifier id="8D06F0D29A4D906B993F345AFADD7B33" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="46F1077F31B381FE2CD38F47A5E1EA04" type="Zenodo-Dep">4539480</mods:identifier>
</mods:mods>
<treatment id="03ABB9399120E34EFBDDFB04FEE0589F" LSID="urn:lsid:plazi:treatment:03ABB9399120E34EFBDDFB04FEE0589F" httpUri="http://treatment.plazi.org/id/03ABB9399120E34EFBDDFB04FEE0589F" lastPageId="9" lastPageNumber="38" pageId="6" pageNumber="35">
<subSubSection id="C3185BA49120E341FBDDFB04FD905A95" box="[220,541,1166,1193]" pageId="6" pageNumber="35" type="nomenclature">
<paragraph id="8BBD082F9120E341FBDDFB04FD905A95" blockId="6.[220,541,1166,1224]" box="[220,541,1166,1193]" pageId="6" pageNumber="35">
<heading id="D0F5BF439120E341FBDDFB04FD905A95" box="[220,541,1166,1193]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="35" reason="2">
<taxonomicName id="4C0273AC9120E341FBDDFB04FE565A95" authority="Duy-Jacquemin, 2014" authorityName="Duy-Jacquemin" authorityYear="2014" box="[220,475,1166,1193]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="humphreysi" status="sp. nov.">
<emphasis id="B976D43D9120E341FBDDFB04FE565A95" bold="true" box="[220,475,1166,1193]" italics="true" pageId="6" pageNumber="35">Lophoturus humphreysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24569469120E341FAE3FB05FD905A95" box="[482,541,1167,1193]" pageId="6" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3185BA49120E341FA44FB24FE3E5AF4" box="[325,435,1198,1224]" pageId="6" pageNumber="35" type="description">
<paragraph id="8BBD082F9120E341FA44FB24FE3E5AF4" blockId="6.[220,541,1166,1224]" box="[325,435,1198,1224]" pageId="6" pageNumber="35">
(
<figureCitation id="133914AA9120E341FA4DFB24FE025AF4" box="[332,399,1198,1224]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="6" pageNumber="35">Figs 3</figureCitation>
,
<figureCitation id="133914AA9120E341FA9DFB24FE265AF4" box="[412,427,1198,1224]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="6" pageNumber="35">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3185BA49120E341FB70FB6DFD085B72" pageId="6" pageNumber="35" type="materials_examined">
<paragraph id="8BBD082F9120E341FB70FB6DFD085B72" blockId="6.[111,647,1255,1358]" pageId="6" pageNumber="35">
<materialsCitation id="3B6A02729120E341FB70FB6DFD085B72" collectingDate="1998-03-30" collectionCode="TYPE, MATERIAL, WAM" collectorName="W. F. Humphreys" country="Australia" location="Hole cave" pageId="6" pageNumber="35" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED1390EA9120E341FB70FB6DFE885AC1" box="[113,261,1255,1278]" pageId="6" pageNumber="35">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="54B9B68D9120E341FA2FFB6DFE015AC3" box="[302,396,1255,1279]" pageId="6" pageNumber="35" type="holotype">Holotype</typeStatus>
: adult male from
<collectingCountry id="F31548BF9120E341F92CFB6DFD095AC3" box="[557,644,1255,1279]" name="Australia" pageId="6" pageNumber="35">Australia</collectingCountry>
,
<collectingCountry id="F31548BF9120E341FB70FA8BFE955B25" box="[113,280,1281,1305]" name="Christmas Island" pageId="6" pageNumber="35">Christmas Island</collectingCountry>
, 19th
<location id="8EDD5EF49120E341FA55FA8BFE3B5B25" LSID="urn:lsid:plazi:treatment:03ABB9399120E34EFBDDFB04FEE0589F:8EDD5EF49120E341FA55FA8BFE3B5B25" box="[340,438,1281,1305]" country="Australia" name="Hole cave" pageId="6" pageNumber="35">Hole cave</location>
, located at – 10.425°, 105.701°:anchialine cave, t° 27-27.4°, humidity 98-99 %, BES-5921,
<date id="FFBC2EEF9120E341FBE1FABCFED95B72" box="[224,340,1334,1358]" pageId="6" pageNumber="35" value="1998-03-30">
<collectingDate id="EFF8D7079120E341FBE1FABCFED95B72" box="[224,340,1334,1358]" pageId="6" pageNumber="35" value="1998-03-30">30.III.1998</collectingDate>
</date>
,
<collectorName id="26F76DF99120E341FA5CFABDFD8A5B72" box="[349,519,1334,1358]" pageId="6" pageNumber="35">W. F. Humphreys</collectorName>
coll.,
<collectionCode id="ED1390EA9120E341F940FABCFD0F5B72" box="[577,642,1334,1358]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="6" pageNumber="35" type="Museum">WAM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3185BA49120E341FB70FAECFF585B8F" pageId="6" pageNumber="35" type="etymology">
<paragraph id="8BBD082F9120E341FB70FAECFF585B8F" blockId="6.[112,647,1382,1459]" pageId="6" pageNumber="35">ETYMOLOGY. — This species is named after its collector, Dr W. F. Humphreys (Senior Curator, Western Australian Museum).</paragraph>
</subSubSection>
<subSubSection id="C3185BA49120E341FB70FA61FC5B5A55" pageId="6" pageNumber="35" type="diagnosis">
<paragraph id="8BBD082F9120E341FB70FA61FF6A583F" blockId="6.[113,648,1515,1699]" box="[113,231,1515,1541]" pageId="6" pageNumber="35">DIAGNOSIS</paragraph>
<paragraph id="8BBD082F9120E341FB70F980FC5B5A55" blockId="6.[113,648,1515,1699]" lastBlockId="6.[675,1211,818,1129]" pageId="6" pageNumber="35">Anterior sensilla basiconica of antennal articles VI and VII shorter and slightly thinner than others. Labrum clothed with small cuticular setae and 2+3 lamellar processes on anteromedian margin. Trichomes of each tergite grouped into two, separate, oval clusters with an additional posterior row barely subdivided in its middle, except on collum and tergite X; distance between clusters much greater than length of each cluster. Pubescent oval setae on leg articles, except tibia and tarsus. Spine of tarsus II subequal in length to the telotarsus on posterior legs. Claw with one small sternal denticle. Apical part of tarsus II with 2 rows of cuticular setae that are longer and more spaced than the other setae.</paragraph>
</subSubSection>
<subSubSection id="C3185BA49120E34EF9A4FB05FEE0589F" lastPageId="9" lastPageNumber="38" pageId="6" pageNumber="35" type="description">
<paragraph id="8BBD082F9120E341F9A4FB05FC6E5A9B" blockId="6.[675,1211,1167,1319]" box="[677,995,1167,1193]" pageId="6" pageNumber="35">DESCRIPTION OF ADULT MALE</paragraph>
<paragraph id="8BBD082F9120E341F9A4FB25FCBB5AF4" blockId="6.[675,1211,1167,1319]" box="[677,822,1199,1224]" pageId="6" pageNumber="35">
<emphasis id="B976D43D9120E341F9A4FB25FCBB5AF4" box="[677,822,1199,1224]" italics="true" pageId="6" pageNumber="35">Measurements</emphasis>
</paragraph>
<paragraph id="8BBD082F9120E341F9A4FB47FBC55B1B" blockId="6.[675,1211,1167,1319]" pageId="6" pageNumber="35">
Body length (without caudal pencil):
<quantity id="4CFAA5CA9120E341FF4AFB47FB3B5AD4" box="[1099,1206,1229,1256]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="35" unit="mm" value="3.6">3.60 mm</quantity>
. Length of caudal penicil
<quantity id="4CFAA5CA9120E341F8ABFB67FB825B34" box="[938,1039,1261,1288]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="35" unit="mm" value="0.6">0.60 mm</quantity>
.Tarsus II length of 1st leg
<quantity id="4CFAA5CA9120E341F80DFA87FCE95B1B" box="[780,868,1293,1319]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="35" unit="mm" value="180.0">180 µm</quantity>
, of 13th leg
<quantity id="4CFAA5CA9120E341F8EAFA87FBC95B1B" box="[1003,1092,1293,1319]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="35" unit="mm" value="240.0">240 µm</quantity>
.
</paragraph>
<paragraph id="8BBD082F9120E341F9A4FAC6FD505B5A" blockId="6.[675,1210,1356,1700]" box="[677,733,1356,1382]" pageId="6" pageNumber="35">
<emphasis id="B976D43D9120E341F9A4FAC6FD505B5A" box="[677,733,1356,1382]" italics="true" pageId="6" pageNumber="35">Head</emphasis>
</paragraph>
<paragraph id="8BBD082F9120E341F9A4FAE6FC9C5818" blockId="6.[675,1210,1356,1700]" pageId="6" pageNumber="35">
Anophthalmic. Vertex with a pair of posterior tufts, each arranged in 2 rows, anterior right row of 23 and left of 22 trichomes; posterior right row of 2 and left of 3 trichomes. Distance between each tuft about equal to their length (
<figureCitation id="133914AA9120E341F9ACF980FC895818" box="[685,772,1546,1572]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="6" pageNumber="35">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="8BBD082F9120E34FF9BAF9A0FCF45858" blockId="6.[675,1210,1356,1700]" lastBlockId="8.[675,1211,1515,1700]" lastPageId="8" lastPageNumber="37" pageId="6" pageNumber="35">
Length and diameter of the antennal articles as represented in
<figureCitation id="133914AA9120E341F866F9C0FC305858" box="[871,957,1610,1636]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="6" pageNumber="35">Fig. 3A</figureCitation>
; length/diameter ratio of article VI about 1.90 (
<tableCitation id="C6803D949120E341F8CFF9E3FBA758B8" box="[974,1066,1641,1668]" captionStart="TABLE" captionStartId="6.[115,124,218,236]" captionText="TABLE 2. — Antennal article ratios (L/d) in the species of Lophoturus Brolemann, 1931 and Lophoproctus Pocock, 1894. Symbol: *, Data after Condé (1982: 315)." pageId="6" pageNumber="35">Table 2</tableCitation>
). Article VI with 3 sensilla basiconica (
<figureCitation id="133914AA9120E341F8D5F903FBDE5898" box="[980,1107,1673,1700]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="6" pageNumber="35">Fig. 3C, D</figureCitation>
), the an- terior (a) more distal, shorter and slightly thinner than the 2 others, intermediate (i) slightly longer than the posterior (p); one sensillum coeloconicum (c) near the posterior sensillum (p), but more distal. Antennal article VII with 2 dorsal sensilla basiconica, the anterior sensil- lum, more distal slightly thinner and shorter (1/2) than the posterior; sensillum coeloconicum (c) near the posterior sensillum (p), but more distal (
<figureCitation id="133914AA912EE34FF9F3F9C0FCE15858" box="[754,876,1610,1636]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="8" pageNumber="37">Fig. 3B, E</figureCitation>
).
</paragraph>
<caption id="DF7D58A79121E340FB70F9BFFEE8589E" ID-DOI="http://doi.org/10.5281/zenodo.4539488" ID-Zenodo-Dep="4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="7" pageNumber="36" startId="7.[113,124,1589,1607]" targetBox="[143,1172,215,1520]" targetPageId="7">
<paragraph id="8BBD082F9121E340FB70F9BFFEE8589E" blockId="7.[113,1211,1589,1698]" pageId="7" pageNumber="36">
FIG. 3. —
<taxonomicName id="4C0273AC9121E340FBBEF9BFFE0E587B" authority="Duy-Jacquemin, 2014" authorityName="Duy-Jacquemin" authorityYear="2014" box="[191,387,1589,1607]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="humphreysi" status="sp. nov.">
<emphasis id="B976D43D9121E340FBBEF9BFFE0E587B" box="[191,387,1589,1607]" italics="true" pageId="7" pageNumber="36">Lophoturus humphreysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24569469121E340FA89F9BFFE34587B" box="[392,441,1589,1607]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
, male holotype:
<emphasis id="B976D43D9121E340F93FF9BFFDC6587B" bold="true" box="[574,587,1589,1607]" pageId="7" pageNumber="36">A</emphasis>
, right antenna;
<emphasis id="B976D43D9121E340F9CAF9BFFD55587B" bold="true" box="[715,728,1589,1607]" pageId="7" pageNumber="36">B</emphasis>
, detail of sensilla on article VII of right antenna;
<emphasis id="B976D43D9121E340FF61F9BFFBE3587B" bold="true" box="[1120,1134,1589,1607]" pageId="7" pageNumber="36">C</emphasis>
, detail of sensilla on article VI of right antenna;
<emphasis id="B976D43D9121E340FAA9F9C6FE3B5862" bold="true" box="[424,438,1612,1630]" pageId="7" pageNumber="36">D</emphasis>
, detail of sensilla on article VII of left antenna;
<emphasis id="B976D43D9121E340F837F9C6FCCF5862" bold="true" box="[822,834,1612,1630]" pageId="7" pageNumber="36">E</emphasis>
, detail of sensilla on article VI of left antenna;
<emphasis id="B976D43D9121E340FB70F9E8FFF15848" bold="true" box="[113,124,1634,1652]" pageId="7" pageNumber="36">F</emphasis>
, trichomes of posterior tufts and right trichobothria on vertex;
<emphasis id="B976D43D9121E340F984F9E8FD1E5848" bold="true" box="[645,659,1634,1652]" pageId="7" pageNumber="36">G</emphasis>
, detail of right anterior (
<emphasis id="B976D43D9121E340F85AF9E8FCE15848" bold="true" box="[859,876,1634,1652]" pageId="7" pageNumber="36">at</emphasis>
) and postero-internal (
<emphasis id="B976D43D9121E340FF28F9E8FBCD5848" bold="true" box="[1065,1088,1634,1652]" pageId="7" pageNumber="36">pit</emphasis>
) trichobothria;
<emphasis id="B976D43D9121E340FB70F9F3FFF258B7" bold="true" box="[113,127,1657,1675]" pageId="7" pageNumber="36">H</emphasis>
, left palpus of gnathochilarium;
<emphasis id="B976D43D9121E340FA96F9F3FE1058B7" bold="true" box="[407,413,1657,1675]" pageId="7" pageNumber="36">I</emphasis>
, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm.
</paragraph>
</caption>
<caption id="DF7D58A7912EE34FFB70FAD4FBBD5BA2" ID-DOI="http://doi.org/10.5281/zenodo.4539492" ID-Zenodo-Dep="4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="8" pageNumber="37" startId="8.[113,124,1374,1392]" targetBox="[126,1187,222,1309]" targetPageId="8">
<paragraph id="8BBD082F912EE34FFB70FAD4FBBD5BA2" blockId="8.[113,1210,1374,1438]" pageId="8" pageNumber="37">
FIG. 4.—
<taxonomicName id="4C0273AC912EE34FFBBCFAD4FEF35B4C" authority="Duy-Jacquemin, 2014" authorityName="Duy-Jacquemin" authorityYear="2014" box="[189,382,1374,1392]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="humphreysi" status="sp. nov.">
<emphasis id="B976D43D912EE34FFBBCFAD4FEF35B4C" box="[189,382,1374,1392]" italics="true" pageId="8" pageNumber="37">Lophoturus humphreysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2456946912EE34FFA83FAD4FE3E5B4C" box="[386,435,1374,1392]" pageId="8" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
, male holotype:
<emphasis id="B976D43D912EE34FF935FAD4FDCC5B4C" bold="true" box="[564,577,1374,1392]" pageId="8" pageNumber="37">A</emphasis>
, insertions of trichomes on collum and right lateral protuberance;
<emphasis id="B976D43D912EE34FFF55FAD4FBEC5B4C" bold="true" box="[1108,1121,1374,1392]" pageId="8" pageNumber="37">B</emphasis>
, insertions of trichomes on tergite IV;
<emphasis id="B976D43D912EE34FFA4BFAFFFED55BBB" bold="true" box="[330,344,1397,1415]" pageId="8" pageNumber="37">C</emphasis>
, right leg II and penis;
<emphasis id="B976D43D912EE34FF913FAFFFDAD5BBB" bold="true" box="[530,544,1397,1415]" pageId="8" pageNumber="37">D</emphasis>
, coxal seta of right leg I;
<emphasis id="B976D43D912EE34FF9EFFAFFFD775BBB" bold="true" box="[750,762,1397,1415]" pageId="8" pageNumber="37">E</emphasis>
, seta of trochanter of right leg I;
<emphasis id="B976D43D912EE34FFF07FAFFFB9C5BBB" bold="true" box="[1030,1041,1397,1415]" pageId="8" pageNumber="37">F</emphasis>
, tarsal spine of right leg VII;
<emphasis id="B976D43D912EE34FFBAEFA01FF305BA1" bold="true" box="[175,189,1419,1437]" pageId="8" pageNumber="37">G</emphasis>
, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm.
</paragraph>
</caption>
<paragraph id="8BBD082F912EE34EF9BAF9E3FEE35F2E" blockId="8.[675,1211,1515,1700]" lastBlockId="9.[111,648,215,464]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
Three trichobothria arranged triangularly, apex of anterior one slightly dilated and ending in a short spine-hair (
<figureCitation id="133914AA912FE34EFA2CFF52FEE05ECE" box="[301,365,216,242]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="9" pageNumber="38">Fig. 3</figureCitation>
[at]); the other 2 are very elongated (
<figureCitation id="133914AA912FE34EFBE9FF7DFEAA5F2E" box="[232,295,247,274]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="9" pageNumber="38">Fig. 3</figureCitation>
[pit]).
</paragraph>
<paragraph id="8BBD082F912FE34EFB89FE9DFE155FAC" blockId="9.[111,648,215,464]" pageId="9" pageNumber="38">
Labrum clothed with some small cuticular setae and 2+3 lamellar processes on antero-median margin (
<figureCitation id="133914AA912FE34EFB76FEDCFF4C5F4D" box="[119,193,342,369]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="9" pageNumber="38">Fig. 3I</figureCitation>
); clypeo-labrum with fourteen short setae along the posterior margin.
</paragraph>
<paragraph id="8BBD082F912FE34EFB89FE1CFE055FF3" blockId="9.[111,648,215,464]" pageId="9" pageNumber="38">
Left palpus of gnathochilarium with 59 (
<figureCitation id="133914AA912FE34EF92AFE1CFDF05F8C" box="[555,637,406,432]" captionStart="FIG" captionStartId="7.[113,124,1589,1607]" captionTargetBox="[143,1172,215,1520]" captionTargetId="figure-130@7.[142,1179,198,1536]" captionTargetPageId="7" captionText="FIG. 3. — Lophoturus humphreysi n. sp., male holotype: A, right antenna; B, detail of sensilla on article VII of right antenna; C, detail of sensilla on article VI of right antenna; D, detail of sensilla on article VII of left antenna; E, detail of sensilla on article VI of left antenna; F, trichomes of posterior tufts and right trichobothria on vertex; G, detail of right anterior (at) and postero-internal (pit) trichobothria; H, left palpus of gnathochilarium; I, labrum, left side only partially drawn. Abbreviations: see Material and methods. Scale bars: A, F-H, 50 µm; B-E, I, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539488" httpUri="https://zenodo.org/record/4539488/files/figure.png" pageId="9" pageNumber="38">Fig. 3H</figureCitation>
) and right with 56 sensilla.
</paragraph>
<paragraph id="8BBD082F912FE34EFB70FE7FFF3F5C33" blockId="9.[111,648,501,939]" box="[113,178,501,527]" pageId="9" pageNumber="38">
<emphasis id="B976D43D912FE34EFB70FE7FFF3F5C33" box="[113,178,501,527]" italics="true" pageId="9" pageNumber="38">Trunk</emphasis>
</paragraph>
<paragraph id="8BBD082F912FE34EFB70FD9FFEB35D97" blockId="9.[111,648,501,939]" pageId="9" pageNumber="38">
Trichomes of each tergite grouped into two, separate, oval clusters with an additional posterior row barely subdivided in its middle (
<figureCitation id="133914AA912FE34EFA75FDDEFE4F5C52" box="[372,450,596,622]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4B</figureCitation>
), except on collum (
<figureCitation id="133914AA912FE34EFB76FDFEFF475CB2" box="[119,202,628,654]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4A</figureCitation>
) and tergite X; distance between clusters much greater than the length of each cluster. Collum with left tuft of 85 trichomes and right tuft of 89 trichomes; lateral left protuberance of collum with 11 trichomes and right with 12 trichomes (
<figureCitation id="133914AA912FE34EFB7AFC98FF585D10" box="[123,213,786,812]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4A</figureCitation>
). Each oval cluster of tergites II to VI with 43 to 54 trichomes and VII to IX with 36 to 41 trichomes; posterior row with 41 to 55 trichomes. Oval clusters of tergite X with 56 trichomes each (no posterior row).
</paragraph>
<paragraph id="8BBD082F912FE34EFB70FC5BFF115DD6" blockId="9.[112,649,977,1510]" box="[113,156,977,1002]" pageId="9" pageNumber="38">
<emphasis id="B976D43D912FE34EFB70FC5BFF115DD6" box="[113,156,977,1002]" italics="true" pageId="9" pageNumber="38">Legs</emphasis>
</paragraph>
<paragraph id="8BBD082F912FE34EFB70FC7AFE1F5BD9" blockId="9.[112,649,977,1510]" pageId="9" pageNumber="38">
Each leg article, except for tibia and tarsus, bearing pubescent oval setae extended by an acute process (
<figureCitation id="133914AA912FE34EFB78FBA5FF625A76" box="[121,239,1071,1098]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4E, D</figureCitation>
); coxae II to XIII bearing 4 to 6 setae (
<figureCitation id="133914AA912FE34EFB78FBC5FF635A55" box="[121,238,1103,1129]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4C, D</figureCitation>
), coxae I with 3 setae; trochanters with 2 setae (
<figureCitation id="133914AA912FE34EFBCBFBE4FE875AB5" box="[202,266,1134,1161]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4</figureCitation>
C-E) (except 1 seta on trochanters of legs I and V); prefemora with 2 setae (except 1 seta on leg I and III, 3 setae on right legs VII and X and left legs X and XIII); femora with 1 seta, very rarely 2 (on VIII femora); 1 small seta only on tibia of leg I. Telotarsus with 2 latero-dorsal, subequal denticles (ldd); a small, frayed, basal denticle (bd) is visible ventrally (
<figureCitation id="133914AA912FE34EFAE2FAC6FDB35B5A" box="[483,574,1356,1382]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4G</figureCitation>
), as in all
<taxonomicName id="4C0273AC912FE34EFB97FAE6FECA5BBA" authorityName="Silvestri" authorityYear="1897" box="[150,327,1388,1414]" class="Diplopoda" family="Lophoproctidae" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="38" phylum="Arthropoda" rank="family">Lophoproctidae</taxonomicName>
. Claw with one small sternal denticle (smd). Apical part of tarsus II with 2 rows of cuticular setae that are longer and more spaced than other setae (
<figureCitation id="133914AA912FE34EFA2CFA41FE085BD9" box="[301,389,1483,1509]" captionStart="FIG" captionStartId="8.[113,124,1374,1392]" captionTargetBox="[126,1187,222,1309]" captionTargetId="figure-184@8.[113,1209,215,1317]" captionTargetPageId="8" captionText="FIG. 4.— Lophoturus humphreysi n. sp., male holotype:A, insertions of trichomes on collum and right lateral protuberance; B, insertions of trichomes on tergite IV; C, right leg II and penis; D, coxal seta of right leg I; E, seta of trochanter of right leg I; F, tarsal spine of right leg VII; G, telotarsus of right leg VII. Abbreviations: see Material and methods. Scale bars: A-C, 50 µm; D-G, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.4539492" httpUri="https://zenodo.org/record/4539492/files/figure.png" pageId="9" pageNumber="38">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BBD082F912FE34EFB70F980FF3D5818" blockId="9.[112,646,1546,1699]" box="[113,176,1546,1572]" pageId="9" pageNumber="38">
<taxonomicName id="4C0273AC912FE34EFB70F980FF3D5818" authorityName="Pearse" authorityYear="1952" box="[113,176,1546,1572]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="9" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B976D43D912FE34EFB70F980FF3D5818" box="[113,176,1546,1572]" italics="true" pageId="9" pageNumber="38">Telson</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BBD082F912FE34EFB70F9A0FEE0589F" blockId="9.[112,646,1546,1699]" pageId="9" pageNumber="38">Seven and 8 trichomes a, of which 6 and 7 placed on internal side and 2 on external side of the group of trichomes b-c1-c3. Trichomes of caudal pencil generally with 3 hooks.</paragraph>
</subSubSection>
</treatment>
</document>
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