treatments-xml/data/03/AB/2F/03AB2F70EE678710A1AB0352FC20FB08.xml

<document id="40A91CA23214E6C3F90A394209F9751F" ID-DOI="10.11646/zootaxa.4759.2.8" ID-GBIF-Dataset="550b757f-332d-4fea-a732-45df5e304ce2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3737497" ID-ZooBank="B70895F2-A6E4-4FEE-8FD4-AE79EA79557B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585803918549" checkinUser="plazi" docAuthor="He, Liang &amp; Zhou, Hong-Zhang" docDate="2020" docId="03AB2F70EE678710A1AB0352FC20FB08" docLanguage="en" docName="zootaxa.4759.2.8.pdf" docOrigin="Zootaxa 4759 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyanocypus He &amp; Zhou 2020, gen. nov." docType="treatment" docVersion="5" lastPageNumber="264" masterDocId="FF925708EE668713A13C017CFF87FFB2" masterDocTitle="Cyanocypus gen. nov., a new rove beetle genus of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae) with one new species from China" masterLastPageNumber="268" masterPageNumber="261" pageNumber="262" updateTime="1698734744615" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9320F9BCE4BD3FE75675E27933E5FA17">Cyanocypus gen. nov., a new rove beetle genus of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae) with one new species from China</mods:title>
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<mods:namePart id="58A34F583E7CDC7D4CF61C1ABC17A099">He, Liang</mods:namePart>
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<mods:namePart id="5E8E6A427783597B815594EA2F9615D8">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="03AB2F70EE678710A1AB0352FC20FB08" ID-DOI="http://doi.org/10.5281/zenodo.3811435" ID-GBIF-Taxon="163420907" ID-Zenodo-Dep="3811435" LSID="urn:lsid:plazi:treatment:03AB2F70EE678710A1AB0352FC20FB08" httpUri="http://treatment.plazi.org/id/03AB2F70EE678710A1AB0352FC20FB08" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262">
<subSubSection id="C318CDEDEE678712A1AB0352FE18FDFB" box="[151,415,558,585]" pageId="1" pageNumber="262" type="nomenclature">
<paragraph id="8BBD9E66EE678712A1AB0352FE18FDFB" blockId="1.[151,415,558,619]" box="[151,415,558,585]" pageId="1" pageNumber="262">
<heading id="D0F5290AEE678712A1AB0352FE18FDFB" bold="true" box="[151,415,558,585]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
<emphasis id="B9764274EE678712A1AB0352FE18FDFB" bold="true" box="[151,415,558,585]" pageId="1" pageNumber="262">
<taxonomicName id="4C02E5E5EE678712A1AB0352FEADFDFA" ID-CoL="8XKPH" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[151,298,558,584]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE678712A1AB0352FEADFDFA" bold="true" box="[151,298,558,584]" italics="true" pageId="1" pageNumber="262">Cyanocypus</emphasis>
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<taxonomicNameLabel id="A245FF0FEE678712A00D0353FE18FDFB" box="[305,415,559,585]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C318CDEDEE678712A1AB032DFEA9FDD9" box="[151,302,593,619]" pageId="1" pageNumber="262" type="description">
<paragraph id="8BBD9E66EE678712A1AB032DFEA9FDD9" blockId="1.[151,415,558,619]" box="[151,302,593,619]" pageId="1" pageNumber="262">
(
<figureCitation id="133982E3EE678712A1A3032DFF6AFDD9" box="[159,237,593,619]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[159,1413,188,1930]" captionTargetId="figure@2.[151,1435,179,1940]" captionTargetPageId="2" captionText="FIGURE 1. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. habitus, dorsal view (2.0 mm); B. habitus, ventral view (2.0 mm); C. left mandible (0.5 mm); D. right mandible (0.5 mm). Scales in parentheses." figureDoi="http://doi.org/10.5281/zenodo.3737499" httpUri="https://zenodo.org/record/3737499/files/figure.png" pageId="1" pageNumber="262">Figs. 1</figureCitation>
, 
<figureCitation id="133982E3EE678712A1C7032DFE8DFDD9" box="[251,266,593,619]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1986]" captionTargetBox="[179,1425,321,1927]" captionTargetId="figure@4.[151,1435,309,1936]" captionTargetPageId="4" captionText="FIGURE 3. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. head, ventral view; B. head, dorsal view; C. aedeagus, lateral view; D. aedeagus, ventral view; E. underside of apical portion of paramere. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737503" httpUri="https://zenodo.org/record/3737503/files/figure.png" pageId="1" pageNumber="262">3</figureCitation>
, 
<figureCitation id="133982E3EE678712A02B032DFEA1FDD9" box="[279,294,593,619]" captionStart="FIGURE 4" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[208,1380,181,1919]" captionTargetId="figure@5.[208,1380,181,1920]" captionTargetPageId="5" captionText="FIGURE 4. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. underside of apical portion of paramere; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. male tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. male sternite 9; H. male sternite 8. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737505" httpUri="https://zenodo.org/record/3737505/files/figure.png" pageId="1" pageNumber="262">4</figureCitation>
)
</paragraph>
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<paragraph id="8BBD9E66EE678712A1AB03EAFCA0FD03" blockId="1.[151,1437,661,2041]" box="[151,807,661,689]" pageId="1" pageNumber="262">
<emphasis id="B9764274EE678712A1AB03EAFEACFD02" bold="true" box="[151,299,661,688]" pageId="1" pageNumber="262">
<typeStatus id="54B920C4EE678712A1AB03EAFF55FD02" box="[151,210,662,688]" pageId="1" pageNumber="262">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="4C02E5E5EE678712A00503EBFD96FD02" ID-CoL="8XZ28" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[313,529,662,689]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="leukos" status="sp. nov.">
<emphasis id="B9764274EE678712A00503EBFD96FD02" box="[313,529,662,689]" italics="true" pageId="1" pageNumber="262">Cyanocypus leukos</emphasis>
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<emphasis id="B9764274EE678712A32403EAFDF6FD02" bold="true" box="[536,625,662,688]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A32403EAFDF6FD02" box="[536,625,662,688]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C318CDEDEE678712A1FB03C4FDDAFC33" pageId="1" pageNumber="262" type="diagnosis">
<paragraph id="8BBD9E66EE678712A1FB03C4FDDAFC33" blockId="1.[151,1437,661,2041]" pageId="1" pageNumber="262">
<emphasis id="B9764274EE678712A1FB03C4FEC5FD61" bold="true" box="[199,322,696,723]" pageId="1" pageNumber="262">Diagnosis.</emphasis>
The new genus 
<taxonomicName id="4C02E5E5EE678712A33D03C5FD0DFD61" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[513,650,697,723]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE678712A33D03C5FD0DFD61" box="[513,650,697,723]" italics="true" pageId="1" pageNumber="262">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE678712A3AF03C5FD79FD61" bold="true" box="[659,766,697,723]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A3AF03C5FD79FD61" box="[659,766,697,723]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from other members of the “ 
<taxonomicName id="4C02E5E5EE678712A59F03C5FAA1FD61" authorityName="Linne" authorityYear="1758" box="[1187,1318,697,723]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A59F03C5FAA1FD61" box="[1187,1318,697,723]" italics="true" pageId="1" pageNumber="262">Staphylinus</emphasis>
</taxonomicName>
-complex” by the following characteristics: mandibles each with one simple tooth in middle of medial margin and with a deep subbasal emargination (
<figureCitation id="133982E3EE678712A09E0382FDB1FCAB" box="[418,566,766,793]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[159,1413,188,1930]" captionTargetId="figure@2.[151,1435,179,1940]" captionTargetPageId="2" captionText="FIGURE 1. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. habitus, dorsal view (2.0 mm); B. habitus, ventral view (2.0 mm); C. left mandible (0.5 mm); D. right mandible (0.5 mm). Scales in parentheses." figureDoi="http://doi.org/10.5281/zenodo.3737499" httpUri="https://zenodo.org/record/3737499/files/figure.png" pageId="1" pageNumber="262">Figs. 1 C, D</figureCitation>
); last segment of maxillary palpus narrowly fusiform, with narrowly rounded apex, asetose; last segment of labial palpus setose, widely fusiform, with widely obliquely truncate apex (
<figureCitation id="133982E3EE678712A40C025DFA08FC89" box="[1328,1423,801,827]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1986]" captionTargetBox="[179,1425,321,1927]" captionTargetId="figure@4.[151,1435,309,1936]" captionTargetPageId="4" captionText="FIGURE 3. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. head, ventral view; B. head, dorsal view; C. aedeagus, lateral view; D. aedeagus, ventral view; E. underside of apical portion of paramere. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737503" httpUri="https://zenodo.org/record/3737503/files/figure.png" pageId="1" pageNumber="262">Fig. 3 A</figureCitation>
); abdominal tergites with oblique accessory basal ridges; sensory peg setae numerous, distributed from apex to basal part of underside of paramere (
<figureCitation id="133982E3EE678712A0CE021AFDD7FC33" box="[498,592,870,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1986]" captionTargetBox="[179,1425,321,1927]" captionTargetId="figure@4.[151,1435,309,1936]" captionTargetPageId="4" captionText="FIGURE 3. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. head, ventral view; B. head, dorsal view; C. aedeagus, lateral view; D. aedeagus, ventral view; E. underside of apical portion of paramere. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737503" httpUri="https://zenodo.org/record/3737503/files/figure.png" pageId="1" pageNumber="262">Fig. 3 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C318CDEDEE678710A1FB02F4FAD4FBC0" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262" type="description">
<paragraph id="8BBD9E66EE678712A1FB02F4FAEDFA19" blockId="1.[151,1437,661,2041]" pageId="1" pageNumber="262">
<emphasis id="B9764274EE678712A1FB02F4FEDEFC11" bold="true" box="[199,345,904,931]" pageId="1" pageNumber="262">Description.</emphasis>
Head, pronotum and elytra with shining blue metallic luster. Mandibles moderately long, each on medial margin with subbasal emargination, and with one simple tooth in middle (
<figureCitation id="133982E3EE678712A55B02D7FA83FC74" box="[1127,1284,939,966]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[159,1413,188,1930]" captionTargetId="figure@2.[151,1435,179,1940]" captionTargetPageId="2" captionText="FIGURE 1. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. habitus, dorsal view (2.0 mm); B. habitus, ventral view (2.0 mm); C. left mandible (0.5 mm); D. right mandible (0.5 mm). Scales in parentheses." figureDoi="http://doi.org/10.5281/zenodo.3737499" httpUri="https://zenodo.org/record/3737499/files/figure.png" pageId="1" pageNumber="262">Figs. 1 C, D</figureCitation>
). Mandibular prostheca with supporting structure long and lanceolate, with ciliae along medial margin short near basal part, but gradually becoming longer towards apex. Gular sutures subcontiguous in middle section. Maxillary palpus moderately long, last segment narrowly fusiform, with narrowly rounded apex, asetose. Labial palpus with last segment setose, widely fusiform, with widely obliquely truncate apex. Pronotal epimeron absent. Pronotal hypomeron glabrous. Superior and inferior lines of pronotal hypomeron join together at about anterior third of pronotal length; deflected portion of pronotal disc largely visible in ventral view at anterior angles of pronotum. Prosternum with sternacostal ridge bisinuate. Mesosternum short; intercoxal process narrow, quite acute apically. Elytra moderately long, not markedly depressed along base; wings fully developed. Metasternum long; anterior projection of metasternum broadly rounded, with acutely converging margins. Dorsolateral face of front tibia with a few spines on apical part; long spines on dorsolateral face of middle tibia robust, quite numerous; dorsolateral face of hind tibia with long spines only on apical half, distinctly shorter and fewer than those on middle tibia. Empodial setae moderately long, distinctly not reaching apex of claws. First three visible abdominal tergites with oblique basal lines extending posteriad from anterior basal line, base of tergite between extensions markedly transversely impressed; lateral parts of first three visible tergites and almost all the surface of visible tergite 5 covered with long silver pubescence.
</paragraph>
<paragraph id="8BBD9E66EE678712A1FB04CFFBDEF984" blockId="1.[151,1437,661,2041]" pageId="1" pageNumber="262">Male genital segment with tergite 8 moderately wide, with obtusely triangular medioapical emargination; sternite 9 with tapered basal portion moderately long, apical portion with wide and deeply rounded medioapical emargination; tergite 10 subtriangular, obtusely angular at apex; aedeagus with sensory peg setae on underside of paramere extremely numerous, arranged from apical to basal part of apical portion of paramere.</paragraph>
<paragraph id="8BBD9E66EE678712A1FB0741FB1DF84B" blockId="1.[151,1437,661,2041]" pageId="1" pageNumber="262">
<emphasis id="B9764274EE678712A1FB0741FD95F9EA" bold="true" box="[199,530,1597,1624]" pageId="1" pageNumber="262">Recognition and comments.</emphasis>
The new genus 
<taxonomicName id="4C02E5E5EE678712A3F60743FCD4F9EB" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[714,851,1599,1625]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE678712A3F60743FCD4F9EB" box="[714,851,1599,1625]" italics="true" pageId="1" pageNumber="262">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE678712A2670742FC43F9EA" bold="true" box="[859,964,1598,1624]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A2670742FC43F9EA" box="[859,964,1598,1624]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, at present, contains only the 
<typeStatus id="54B920C4EE678712A4280743FAC3F9EB" box="[1300,1348,1599,1625]" pageId="1" pageNumber="262">type</typeStatus>
species 
<taxonomicName id="4C02E5E5EE678712A1AB071DFEE9F9C9" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[151,366,1633,1659]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="leukos" status="sp. nov.">
<emphasis id="B9764274EE678712A1AB071DFEE9F9C9" box="[151,366,1633,1659]" italics="true" pageId="1" pageNumber="262">Cyanocypus leukos</emphasis>
</taxonomicName>
<emphasis id="B9764274EE678712A048071DFE4AF9C9" bold="true" box="[372,461,1633,1659]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A048071DFE4AF9C9" box="[372,461,1633,1659]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It can be easily distinguished from other members of the “ 
<taxonomicName id="4C02E5E5EE678712A564071DFB5CF9C9" authorityName="Linne" authorityYear="1758" box="[1112,1243,1633,1659]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A564071DFB5CF9C9" box="[1112,1243,1633,1659]" italics="true" pageId="1" pageNumber="262">Staphylinus</emphasis>
</taxonomicName>
-complex” by the characteristics listed above under Diagnosis, especially the mandibular characters. Mandible morphology is now considered as an important character system for distinguishing the genera of the “ 
<taxonomicName id="4C02E5E5EE678712A51707DAFB29F972" authorityName="Linne" authorityYear="1758" box="[1067,1198,1702,1728]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A51707DAFB29F972" box="[1067,1198,1702,1728]" italics="true" pageId="1" pageNumber="262">Staphylinus</emphasis>
</taxonomicName>
-complex” (
<bibRefCitation id="EF93E397EE678712A40F07DAFF53F951" author="Smetana, A." pageId="1" pageNumber="262" pagination="57 - 136" refId="ref3491" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 136." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
). 
<bibRefCitation id="EF93E397EE678712A1DA07B5FE10F951" author="Smetana, A." box="[230,407,1737,1763]" pageId="1" pageNumber="262" pagination="57 - 136" refId="ref3491" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 136." type="journal article" year="2003">Smetana (2003)</bibRefCitation>
provided a summary of the 
<typeStatus id="54B920C4EE678712A3F007B5FC80F951" box="[716,775,1737,1763]" pageId="1" pageNumber="262">types</typeStatus>
of mandible shape in the “ 
<taxonomicName id="4C02E5E5EE678712A51307B5FB35F951" authorityName="Linne" authorityYear="1758" box="[1071,1202,1737,1763]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A51307B5FB35F951" box="[1071,1202,1737,1763]" italics="true" pageId="1" pageNumber="262">Staphylinus</emphasis>
</taxonomicName>
-complex”; however, the mandible morphology of 
<taxonomicName id="4C02E5E5EE678712A0DC0790FDEEF8B4" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[480,617,1772,1798]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE678712A0DC0790FDEEF8B4" box="[480,617,1772,1798]" italics="true" pageId="1" pageNumber="262">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE678712A34C0790FD5DF8B4" bold="true" box="[624,730,1772,1798]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A34C0790FD5DF8B4" box="[624,730,1772,1798]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
does not belong to any of those 
<typeStatus id="54B920C4EE678712A5760790FB0EF8B4" box="[1098,1161,1772,1798]" pageId="1" pageNumber="262">types</typeStatus>
. In this new genus, each mandible has a wide subbasal indentation and one simple tooth in the middle, easily distinguishing it from 
<taxonomicName id="4C02E5E5EE678712A4740673FA1BF89B" box="[1352,1436,1807,1833]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A4740673FA1BF89B" box="[1352,1436,1807,1833]" italics="true" pageId="1" pageNumber="262">Ocypus</emphasis>
</taxonomicName>
and similar genera. A similar kind of wide subbasal indentation occurs in the species of 
<taxonomicName id="4C02E5E5EE678712A55B064EFB62F8F9" box="[1127,1253,1842,1867]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A55B064EFB62F8F9" box="[1127,1253,1842,1867]" italics="true" pageId="1" pageNumber="262">Protocypus</emphasis>
</taxonomicName>
, but the tooth in the middle of the mandibles of 
<taxonomicName id="4C02E5E5EE678712A0C90628FDF3F8DF" box="[501,628,1876,1901]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A0C90628FDF3F8DF" box="[501,628,1876,1901]" italics="true" pageId="1" pageNumber="262">Protocypus</emphasis>
</taxonomicName>
is bidentate (
<figureCitation id="133982E3EE678712A235062FFCE5F8DC" box="[777,866,1875,1902]" captionStart="FIGURE 2" captionStartId="3.[151,250,752,777]" captionTargetBox="[151,1435,188,726]" captionTargetId="figure@3.[151,1435,181,727]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal view of head:A. Cyanocypus leukos He &amp; Zhou, sp. nov.; B. Sphaerobulbus yunnanus Smetana; C. Protocypus taibaiensis He &amp; Zhou. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737501" httpUri="https://zenodo.org/record/3737501/files/figure.png" pageId="1" pageNumber="262">Fig. 2C</figureCitation>
). The new genus shares the modification of the abdominal tergites (first three visible abdominal tergites with accessory basal lines, base of tergite between these lines transversely impressed) with the genus 
<taxonomicName id="4C02E5E5EE678712A36806E5FCD6F801" authority="J. Muller" authorityName="J. Muller" box="[596,849,1945,1971]" class="Insecta" family="Staphylinidae" genus="Aulacocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A36806E5FD64F801" box="[596,739,1945,1971]" italics="true" pageId="1" pageNumber="262">Aulacocypus</emphasis>
J. Müller
</taxonomicName>
, the “ 
<emphasis id="B9764274EE678712A2AF06E5FB81F801" box="[915,1030,1945,1971]" italics="true" pageId="1" pageNumber="262">bisinuatus</emphasis>
-group” of the genus 
<taxonomicName id="4C02E5E5EE678712A5C906E5FF7DF864" authority="Smetana" authorityName="Smetana" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A5C906E5FA1BF801" box="[1269,1436,1945,1971]" italics="true" pageId="1" pageNumber="262">Sphaerobulbus</emphasis>
Smetana
</taxonomicName>
, and the genera 
<taxonomicName id="4C02E5E5EE678712A08506C0FD41F864" authority="Smetana" authorityName="Smetana" authorityYear="2006" box="[441,710,1980,2006]" class="Insecta" family="Staphylinidae" genus="Nelmanwaslus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A08506C0FDDBF864" box="[441,604,1980,2006]" italics="true" pageId="1" pageNumber="262">Nelmanwaslus</emphasis>
Smetana
</taxonomicName>
and 
<taxonomicName id="4C02E5E5EE678712A23E06C0FBEBF864" authority="Bernhauer. From" authorityName="Bernhauer. From" authorityYear="1920" box="[770,1132,1979,2006]" class="Insecta" family="Staphylinidae" genus="Wasmannellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE678712A23E06C0FC24F864" box="[770,931,1980,2006]" italics="true" pageId="1" pageNumber="262">Wasmannellus</emphasis>
Bernhauer. From
</taxonomicName>
these genera, 
<taxonomicName id="4C02E5E5EE678712A42F06C0FA1BF864" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[1299,1436,1980,2006]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE678712A42F06C0FA1BF864" box="[1299,1436,1980,2006]" italics="true" pageId="1" pageNumber="262">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE678712A1AB06A2FE87F84A" bold="true" box="[151,256,2014,2040]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A245FF0FEE678712A1AB06A2FE87F84A" box="[151,256,2014,2040]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished by the subbasal mandibular indentation (
<figureCitation id="133982E3EE678712A53F06A2FB0AF84B" box="[1027,1165,2014,2041]" captionStart="FIGURE 2" captionStartId="3.[151,250,752,777]" captionTargetBox="[151,1435,188,726]" captionTargetId="figure@3.[151,1435,181,727]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal view of head:A. Cyanocypus leukos He &amp; Zhou, sp. nov.; B. Sphaerobulbus yunnanus Smetana; C. Protocypus taibaiensis He &amp; Zhou. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737501" httpUri="https://zenodo.org/record/3737501/files/figure.png" pageId="1" pageNumber="262">Figs. 2 A, B</figureCitation>
).
</paragraph>
<caption id="DF7DCEEEEE648711A1AB06D3FC14F85E" ID-DOI="http://doi.org/10.5281/zenodo.3737499" ID-Zenodo-Dep="3737499" httpUri="https://zenodo.org/record/3737499/files/figure.png" pageId="2" pageNumber="263" startId="2.[151,250,1967,1992]" targetBox="[159,1413,188,1930]" targetPageId="2">
<paragraph id="8BBD9E66EE648711A1AB06D3FC14F85E" blockId="2.[151,1437,1967,2028]" pageId="2" pageNumber="263">
<emphasis id="B9764274EE648711A1AB06D3FE94F87A" bold="true" box="[151,275,1967,1992]" pageId="2" pageNumber="263">FIGURE 1.</emphasis>
<taxonomicName id="4C02E5E5EE648711A02506CCFDD9F87A" authority="He &amp; Zhou" authorityName="He &amp; Zhou" authorityYear="2020" box="[281,606,1967,1992]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="leukos" status="sp. nov.">
<emphasis id="B9764274EE648711A02506CCFE5BF87A" box="[281,476,1968,1992]" italics="true" pageId="2" pageNumber="263">Cyanocypus leukos</emphasis>
<emphasis id="B9764274EE648711A0DE06CCFDD9F87A" bold="true" box="[482,606,1967,1992]" pageId="2" pageNumber="263">He &amp; Zhou</emphasis>
</taxonomicName>
<emphasis id="B9764274EE648711A36206CCFD3DF87A" bold="true" box="[606,698,1968,1992]" pageId="2" pageNumber="263">
, 
<taxonomicNameLabel id="A245FF0FEE648711A35506CCFD3DF87A" box="[617,698,1968,1992]" pageId="2" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. habitus, dorsal view (2.0 mm); B. habitus, ventral view (2.0 mm); C. left mandible (0.5 mm); D. right mandible (0.5 mm). Scales in parentheses.
</paragraph>
</caption>
<caption id="DF7DCEEEEE658710A1AB038CFD32FC9E" ID-DOI="http://doi.org/10.5281/zenodo.3737501" ID-Zenodo-Dep="3737501" httpUri="https://zenodo.org/record/3737501/files/figure.png" pageId="3" pageNumber="264" startId="3.[151,250,752,777]" targetBox="[151,1435,188,726]" targetPageId="3">
<paragraph id="8BBD9E66EE658710A1AB038CFD32FC9E" blockId="3.[151,1436,752,813]" pageId="3" pageNumber="264">
<emphasis id="B9764274EE658710A1AB038CFE95FCBB" bold="true" box="[151,274,752,777]" pageId="3" pageNumber="264">FIGURE 2.</emphasis>
Dorsal view of head: A. 
<taxonomicName id="4C02E5E5EE658710A332038DFCD6FCBB" authority="He &amp; Zhou" authorityName="He &amp; Zhou" authorityYear="2020" box="[526,849,752,777]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="leukos" status="sp. nov.">
<emphasis id="B9764274EE658710A332038DFD57FCBB" box="[526,720,753,777]" italics="true" pageId="3" pageNumber="264">Cyanocypus leukos</emphasis>
<emphasis id="B9764274EE658710A3EA038DFCD6FCBB" bold="true" box="[726,849,752,777]" pageId="3" pageNumber="264">He &amp; Zhou</emphasis>
</taxonomicName>
<emphasis id="B9764274EE658710A26D038DFC2BFCBB" bold="true" box="[849,940,753,777]" pageId="3" pageNumber="264">
, 
<taxonomicNameLabel id="A245FF0FEE658710A260038DFC2BFCBB" box="[860,940,753,777]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B. 
<taxonomicName id="4C02E5E5EE658710A2EA038DFAB2FCBB" authority="Smetana" authorityName="Smetana" authorityYear="2003" box="[982,1333,753,777]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B9764274EE658710A2EA038DFB53FCBB" box="[982,1236,753,777]" italics="true" pageId="3" pageNumber="264">Sphaerobulbus yunnanus</emphasis>
Smetana
</taxonomicName>
; C. 
<taxonomicName id="4C02E5E5EE658710A467038EFD8EFC9F" authority="He &amp; Zhou. Scale" authorityName="He &amp; Zhou. Scale" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="taibaiensis">
<emphasis id="B9764274EE658710A467038EFEC2FC9F" italics="true" pageId="3" pageNumber="264">Protocypus taibaiensis</emphasis>
He &amp; Zhou. Scale
</taxonomicName>
bars = 0.50 mm.
</paragraph>
</caption>
<paragraph id="8BBD9E66EE658710A1FB0227FCBEFB9B" blockId="3.[151,1437,859,1210]" pageId="3" pageNumber="264">
The numerous sensory peg setae as well as their arrangement on underside of the paramere of 
<taxonomicName id="4C02E5E5EE658710A5E00220FAE2FCC4" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[1244,1381,860,886]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE658710A5E00220FAE2FCC4" box="[1244,1381,860,886]" italics="true" pageId="3" pageNumber="264">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE658710A4570220FF4FFC28" bold="true" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A245FF0FEE658710A4570220FF4FFC28" pageId="3" pageNumber="264" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are also quite unique in the 
<taxonomicName id="4C02E5E5EE658710A32F0203FD11FC2B" authorityName="Linne" authorityYear="1758" box="[531,662,895,921]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE658710A32F0203FD11FC2B" box="[531,662,895,921]" italics="true" pageId="3" pageNumber="264">Staphylinus</emphasis>
</taxonomicName>
complex. In all other genera or species of the 
<taxonomicName id="4C02E5E5EE658710A58C0203FAB4FC2B" authorityName="Linne" authorityYear="1758" box="[1200,1331,895,921]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE658710A58C0203FAB4FC2B" box="[1200,1331,895,921]" italics="true" pageId="3" pageNumber="264">Staphylinus</emphasis>
</taxonomicName>
complex with sensory peg setae, they are mostly situated on apical to middle portion on the underside of paramere, never stretching far towards the base of paramere. In 
<taxonomicName id="4C02E5E5EE658710A38902B4FCB9FC50" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[693,830,968,994]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE658710A38902B4FCB9FC50" box="[693,830,968,994]" italics="true" pageId="3" pageNumber="264">Cyanocypus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245FF0FEE658710A27B02B4FC34FC50" box="[839,947,968,994]" pageId="3" pageNumber="264" rank="genus">
<emphasis id="B9764274EE658710A27B02B4FC2AFC50" bold="true" box="[839,941,968,994]" pageId="3" pageNumber="264">gen. nov</emphasis>
.
</taxonomicNameLabel>
the sensory peg setae are widely extended from apical to basal portion of the paramere (
<figureCitation id="133982E3EE658710A3B00297FD6FFBB4" box="[652,744,1003,1030]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1986]" captionTargetBox="[179,1425,321,1927]" captionTargetId="figure@4.[151,1435,309,1936]" captionTargetPageId="4" captionText="FIGURE 3. Cyanocypus leukos He &amp; Zhou, sp. nov.: A. head, ventral view; B. head, dorsal view; C. aedeagus, lateral view; D. aedeagus, ventral view; E. underside of apical portion of paramere. Scale bars = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3737503" httpUri="https://zenodo.org/record/3737503/files/figure.png" pageId="3" pageNumber="264">Fig. 3 E</figureCitation>
). This character state seems to be an autapomorphy of the new genus 
<taxonomicName id="4C02E5E5EE658710A1DC056CFEEEFB98" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[224,361,1040,1066]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE658710A1DC056CFEEEFB98" box="[224,361,1040,1066]" italics="true" pageId="3" pageNumber="264">Cyanocypus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245FF0FEE658710A053056CFE5EFB98" box="[367,473,1040,1066]" pageId="3" pageNumber="264" rank="genus">
<emphasis id="B9764274EE658710A053056CFE54FB98" bold="true" box="[367,467,1040,1066]" pageId="3" pageNumber="264">gen. nov</emphasis>
.
</taxonomicNameLabel>
, but this requires further study.
</paragraph>
<paragraph id="8BBD9E66EE658710A1FB054FFAD4FBC0" blockId="3.[151,1437,859,1210]" pageId="3" pageNumber="264">
<bibRefCitation id="EF93E397EE658710A1FB054FFE76FBFC" author="Brunke, A. J. &amp; Smetana, A." box="[199,497,1075,1102]" pageId="3" pageNumber="264" pagination="745 - 758" refId="ref3435" refString="Brunke, A. J. &amp; Smetana, A. (2019) A new genus of Staphylinina and a review of major lineages (Staphylinidae: Staphylininae: Staphylinini). Systematics and Biodiversity, 17 (8), 745 - 758. https: // doi. org / 10.1080 / 14772000.2019.1691082" type="journal article" year="2019">Brunke &amp; Smetana (2019)</bibRefCitation>
characterized the major lineages of subtribe 
<taxonomicName id="4C02E5E5EE658710A2D7054FFBFEFBFF" authorityName="Latreille" authorityYear="1802" box="[1003,1145,1075,1101]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
and according the abovementioned characters of 
<taxonomicName id="4C02E5E5EE658710A0970524FDB3FBC0" authority="He &amp; Zhou, 2020" authorityName="He &amp; Zhou" authorityYear="2020" box="[427,564,1112,1138]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9764274EE658710A0970524FDB3FBC0" box="[427,564,1112,1138]" italics="true" pageId="3" pageNumber="264">Cyanocypus</emphasis>
</taxonomicName>
<emphasis id="B9764274EE658710A3070524FD23FBC0" bold="true" box="[571,676,1112,1138]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A245FF0FEE658710A3070524FD23FBC0" box="[571,676,1112,1138]" pageId="3" pageNumber="264" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, it is possible to include this new genus in the 
<taxonomicName id="4C02E5E5EE658710A5930524FA84FBC0" box="[1199,1283,1112,1138]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE658710A5930524FA84FBC0" box="[1199,1283,1112,1138]" italics="true" pageId="3" pageNumber="264">Ocypus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C318CDEDEE658710A1FB0500FC20FB08" pageId="3" pageNumber="264" type="etymology">
<paragraph id="8BBD9E66EE658710A1FB0500FC20FB08" blockId="3.[151,1437,859,1210]" pageId="3" pageNumber="264">
<emphasis id="B9764274EE658710A1FB0500FEC9FB24" bold="true" box="[199,334,1148,1174]" pageId="3" pageNumber="264">Etymology.</emphasis>
The new genus name 
<taxonomicName id="4C02E5E5EE658710A37D0500FD4DFB24" authorityName="He &amp; Zhou" authorityYear="2020" box="[577,714,1148,1174]" class="Insecta" family="Staphylinidae" genus="Cyanocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE658710A37D0500FD4DFB24" box="[577,714,1148,1174]" italics="true" pageId="3" pageNumber="264">Cyanocypus</emphasis>
</taxonomicName>
(masc.) is derived from the combination of the Greek term 
<emphasis id="B9764274EE658710A4640500FA1BFB24" box="[1368,1436,1148,1174]" italics="true" pageId="3" pageNumber="264">Cyan-</emphasis>
(dark blue) and 
<taxonomicName id="4C02E5E5EE658710A07505DCFE1AFB08" box="[329,413,1184,1210]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B9764274EE658710A07505DCFE1AFB08" box="[329,413,1184,1210]" italics="true" pageId="3" pageNumber="264">Ocypus</emphasis>
</taxonomicName>
(the known genus name used for a long time).
</paragraph>
</subSubSection>
</treatment>
</document>