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<document id="23EEDACBF7BD43CA02BC30516ADCA46C" ID-DOI="10.1111/zoj.12295" ID-ISSN="0024-4082" ID-Zenodo-Dep="5339562" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630184374276" checkinUser="felipe" docAuthor="Santodomingo, Nadiezhda, Wallace, Carden C. &amp; Johnson, Kenneth G." docDate="2015" docId="03AB216FFFA3F655FEF8FF167802FA8C" docLanguage="en" docName="zoj12295.pdf" docOrigin="Zoological Journal of the Linnean Society 175 (4)" docSource="http://dx.doi.org/10.1111/zoj.12295" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Acropora renemai Santodomingo &amp; Wallace &amp; Johnson 2015, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="753" masterDocId="FF925917FFEFF619FFE0FFD07C56FFBA" masterDocTitle="Fossils reveal a high diversity of the staghorn coral genera Acropora and Isopora (Scleractinia: Acroporidae) in the Neogene of Indonesia" masterLastPageNumber="763" masterPageNumber="677" pageNumber="753" updateTime="1699178715059" updateUser="plazi">
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<mods:title id="0B30716686A8360F000EDD411E173DBB">Fossils reveal a high diversity of the staghorn coral genera Acropora and Isopora (Scleractinia: Acroporidae) in the Neogene of Indonesia</mods:title>
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<mods:namePart id="8CA71AAA522CDB06DDA4AF72E76C6006">Santodomingo, Nadiezhda</mods:namePart>
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<mods:namePart id="3817AB844C81261D0A01724A8743F39A">Wallace, Carden C.</mods:namePart>
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<mods:namePart id="6C223ED8A7046520BCD95A3B0A8DB0F0">Johnson, Kenneth G.</mods:namePart>
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<mods:title id="6CBDE4A7F41873C666BBC7C63C5F1DB4">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="ABC6CF79E9F4B83BFB273F62C90BFC52">2015</mods:date>
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<mods:number id="63132CBB550E8B821E1B5D0F4726FED1">2015-11-18</mods:number>
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<mods:classification id="335A1EEE8F38FF01F300196114B8944C">journal article</mods:classification>
<mods:identifier id="765B10D4746BDBE2D5B5B3C993AFE1DC" type="DOI">10.1111/zoj.12295</mods:identifier>
<mods:identifier id="557B4A8E18570E31EDEA55C842978505" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="895CD3113820BF6412EA52C803CFD1EA" type="Zenodo-Dep">5339562</mods:identifier>
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<treatment id="03AB216FFFA3F655FEF8FF167802FA8C" LSID="urn:lsid:plazi:treatment:03AB216FFFA3F655FEF8FF167802FA8C" httpUri="http://treatment.plazi.org/id/03AB216FFFA3F655FEF8FF167802FA8C" lastPageNumber="753" pageId="76" pageNumber="753">
<subSubSection id="C318C3F2FFA3F655FEF8FF167E38FF61" box="[280,622,198,219]" pageId="76" pageNumber="753" type="nomenclature">
<paragraph id="8BBD9079FFA3F655FEF8FF167E38FF61" blockId="76.[280,622,198,219]" box="[280,622,198,219]" pageId="76" pageNumber="753">
<heading id="D0F52715FFA3F655FEF8FF167E38FF61" box="[280,622,198,219]" centered="true" fontSize="9" level="2" pageId="76" pageNumber="753" reason="4">
<taxonomicName id="4C02EBFAFFA3F655FEF8FF167E5FFF60" ID-CoL="64NMH" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" box="[280,521,198,219]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="690" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">ACROPORA RENEMAI</taxonomicName>
<emphasis id="B9764C6BFFA3F655FDF2FF197E38FF61" bold="true" box="[530,622,198,219]" pageId="76" pageNumber="753">
<taxonomicNameLabel id="A245F110FFA3F655FDF2FF197E38FF61" box="[530,622,198,219]" pageId="76" pageNumber="753" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318C3F2FFA3F655FE67FF3D7802FA8C" pageId="76" pageNumber="753" type="description">
<paragraph id="8BBD9079FFA3F655FE67FF3D7DA9FEB8" blockId="76.[391,511,237,258]" box="[391,511,237,258]" pageId="76" pageNumber="753">
<figureCitation id="13398CFCFFA3F655FE67FF3D7DA9FEB8" box="[391,511,237,258]" captionStart="Figure 39" captionStartId="77.[164,243,1704,1723]" captionTargetBox="[194,1413,195,1675]" captionTargetId="figure-7@77.[638,976,686,1170]" captionTargetPageId="77" captionText="Figure 39. Acropora renemai sp. nov. Holotype, A, D, H, NHMUK PI AZ 7439, TF502, Bontang, East Kalimantan, Late Miocene (9.4–9.8 Ma). A, whole specimen. B, C, E–G, paratype, NHMUK PI AZ 9117, TF502, Bontang, East Kalimantan, Late Miocene (9.4–9.8 Ma): B, AZ 9117a–d anticlockwise, four specimens showing different parts of a colony. C, AZ 9117b, transverse section of a branch showing axial corallite and cylindrical branches. D, AZ 7439, holotype, electron micrograph of the branch tip showing immersed radial corallites. E, AZ 9117e, branch tip showing appressed tubular radial corallites. F, AZ 9117a, detail of appressed tubular radial corallite. G, AZ 9117c, detail of incipient branch and immersed radial corallites. H, AZ 7439, holotype, electron micrograph of dense reticulate coenosteum formed of elaborated spinules." figureDoi="http://doi.org/10.5281/zenodo.5339644" httpUri="https://zenodo.org/record/5339644/files/figure.png" pageId="76" pageNumber="753">FIGURE 39</figureCitation>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FEC47D56FE93" blockId="76.[144,759,276,419]" box="[144,256,276,297]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF70FEC47D56FE93" box="[144,256,276,297]" italics="true" pageId="76" pageNumber="753">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FEE27D41FE19" blockId="76.[144,759,276,419]" pageId="76" pageNumber="753">Colony probably corymbose; radial corallites immersed, evenly sized, scattered on the branches; coenosteum with densely packed elaborated spinules throughout.</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FE6F7D18FE6E" blockId="76.[144,759,447,775]" box="[144,334,447,468]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF70FE6F7D18FE6E" box="[144,334,447,468]" italics="true" pageId="76" pageNumber="753">Material studied</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FE0D7DAEFDF4" blockId="76.[144,759,447,775]" pageId="76" pageNumber="753">
<materialsCitation id="3B6A9A24FFA3F655FF70FE0D7DAEFDF4" collectingDate="2011-07-02" collectionCode="NHMUK, PI" collectorName="Tortonian &amp; Ma. Collector N. Santodomingo &amp; Collector N. Santodomingo" latitude="0.14130001" location="Bontang" longLatPrecision="1" longitude="117.431" pageId="76" pageNumber="753" specimenCode="AZ 7439, TF502" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
<emphasis id="B9764C6BFFA3F655FF70FE0D7CADFE48" box="[144,251,477,498]" italics="true" pageId="76" pageNumber="753">
<typeStatus id="54B92EDBFFA3F655FF70FE0D7CA1FE48" box="[144,247,477,498]" pageId="76" pageNumber="753" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED1308BCFFA3F655FEE3FE0E7D3CFE49" box="[259,362,478,499]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="76" pageNumber="753" type="Museum">NHMUK</collectionCode>
<collectionCode id="ED1308BCFFA3F655FE94FE0E7DD8FE49" box="[372,398,478,499]" country="Russia" name="Paleontological Institute" pageId="76" pageNumber="753">PI</collectionCode>
<specimenCode id="DBA43802FFA3F655FE77FE0E7DACFE49" box="[407,506,478,499]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="76" pageNumber="753">AZ 7439</specimenCode>
,
<specimenCount id="9D045BF0FFA3F655FDE8FE0E7EDBFE49" box="[520,653,477,499]" pageId="76" pageNumber="753" type="generic">1 specimen</specimenCount>
. Type locality:
<specimenCode id="DBA43802FFA3F655FF3BFE2C7D70FDA8" box="[219,294,508,530]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="76" pageNumber="753">TF502</specimenCode>
,
<location id="8EDDC6A2FFA3F655FED1FE2D7DC4FDA8" LSID="urn:lsid:plazi:treatment:03AB216FFFA3F655FEF8FF167802FA8C:8EDDC6A2FFA3F655FED1FE2D7DC4FDA8" box="[305,402,509,530]" latitude="0.14130001" longLatPrecision="1" longitude="117.431" name="Bontang" pageId="76" pageNumber="753" stateProvince="Kalimantan Timur">Bontang</location>
,
<collectingRegion id="49C65E9BFFA3F655FE7EFE2D7E36FDA8" box="[414,608,508,530]" country="Indonesia" name="Kalimantan Timur" pageId="76" pageNumber="753">East Kalimantan</collectingRegion>
,
<geoCoordinate id="EE36F6BEFFA3F655FD8CFE2C7EA4FDA8" box="[620,754,508,530]" degrees="0" direction="north" minutes="8" orientation="latitude" pageId="76" pageNumber="753" precision="1" seconds="28.68" value="0.14130001">0°8′28.68″N</geoCoordinate>
,
<geoCoordinate id="EE36F6BEFFA3F655FF70FDCB7D69FD8A" box="[144,319,538,561]" degrees="117" direction="east" minutes="25" orientation="longitude" pageId="76" pageNumber="753" precision="1" seconds="51.60" value="117.431">117°25′51.60″E</geoCoordinate>
.
<collectorName id="26F7F5AFFFA3F655FEADFDCB7DEAFD8B" box="[333,444,539,561]" pageId="76" pageNumber="753">Tortonian</collectorName>
age, 9.4–9.8
<collectorName id="26F7F5AFFFA3F655FDBBFDCB7D0CFDF5" pageId="76" pageNumber="753">Ma. Collector N. Santodomingo</collectorName>
,
<date id="FFBCB6B9FFA3F655FE88FDE97DA2FDF4" box="[360,500,569,591]" pageId="76" pageNumber="753" value="2011-07-02">
<collectingDate id="EFF84F51FFA3F655FE88FDE97DA2FDF4" box="[360,500,569,591]" pageId="76" pageNumber="753" value="2011-07-02">2 July 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF48FD897D49FCBC" blockId="76.[144,759,447,775]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF48FD897D40FDD4" box="[168,278,601,622]" italics="true" pageId="76" pageNumber="753">
<typeStatus id="54B92EDBFFA3F655FF48FD897D44FDD4" box="[168,274,601,622]" pageId="76" pageNumber="753" type="paratype">Paratype</typeStatus>
.
</emphasis>
NHMUK PI AZ 9117, 69 fragments recovered together with the
<typeStatus id="54B92EDBFFA3F655FE4BFDA67E5FFD36" box="[427,521,630,652]" pageId="76" pageNumber="753" type="holotype">holotype</typeStatus>
, uncertain if all specimens belong to the same colony. Type locality:
<materialsCitation id="3B6A9A24FFA3F655FD46FD457D4AFCBC" collectingDate="2011-07-02" collectionCode="TF" collectorName="Tortonian &amp; Ma. Collector N. Santodomingo &amp; Collector N. Santodomingo" latitude="0.14130001" location="Bontang" longLatPrecision="1" longitude="117.431" pageId="76" pageNumber="753" specimenCode="TF502" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<specimenCode id="DBA43802FFA3F655FD46FD457EA5FD11" box="[678,755,661,683]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="76" pageNumber="753">TF502</specimenCode>
,
<location id="8EDDC6A2FFA3F655FF70FD647CB8FD73" LSID="urn:lsid:plazi:treatment:03AB216FFFA3F655FEF8FF167802FA8C:8EDDC6A2FFA3F655FF70FD647CB8FD73" box="[144,238,692,713]" latitude="0.14130001" longLatPrecision="1" longitude="117.431" name="Bontang" pageId="76" pageNumber="753" stateProvince="Kalimantan Timur">Bontang</location>
,
<collectingRegion id="49C65E9BFFA3F655FF19FD647DE1FD70" box="[249,439,692,714]" country="Indonesia" name="Kalimantan Timur" pageId="76" pageNumber="753">East Kalimantan</collectingRegion>
,
<geoCoordinate id="EE36F6BEFFA3F655FE21FD647E15FD73" box="[449,579,691,714]" degrees="0" direction="north" minutes="8" orientation="latitude" pageId="76" pageNumber="753" precision="1" seconds="28.68" value="0.14130001">0°8′28.68″N</geoCoordinate>
,
<geoCoordinate id="EE36F6BEFFA3F655FDAEFD647EA5FD73" box="[590,755,691,714]" degrees="117" direction="east" minutes="25" orientation="longitude" pageId="76" pageNumber="753" precision="1" seconds="51.60" value="117.431">117°25′51.60″E</geoCoordinate>
.
<collectorName id="26F7F5AFFFA3F655FF70FD027CAAFD52" box="[144,252,722,744]" pageId="76" pageNumber="753">Tortonian</collectorName>
age, 9.4–9.8
<collectorName id="26F7F5AFFFA3F655FE70FD037EA4FD52" box="[400,754,722,744]" pageId="76" pageNumber="753">Ma. Collector N. Santodomingo</collectorName>
,
<date id="FFBCB6B9FFA3F655FF70FD217D4AFCBC" box="[144,284,753,775]" pageId="76" pageNumber="753" value="2011-07-02">
<collectingDate id="EFF84F51FFA3F655FF70FD217D4AFCBC" box="[144,284,753,775]" pageId="76" pageNumber="753" value="2011-07-02">2 July 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FCF37D47FC82" blockId="76.[144,759,803,1650]" box="[144,273,803,824]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF70FCF37D47FC82" box="[144,273,803,824]" italics="true" pageId="76" pageNumber="753">Description</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70FC917EB1FB97" blockId="76.[144,759,803,1650]" pageId="76" pageNumber="753">
<taxonomicName id="4C02EBFAFFA3F655FF70FC917CAAFCEC" authorityName="Ehrenberg" authorityYear="1834" box="[144,252,833,854]" genus="Corallum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFA3F655FF70FC917CAAFCEC" box="[144,252,833,854]" italics="true" pageId="76" pageNumber="753">Corallum</emphasis>
</taxonomicName>
. Cylindrical branches, extending randomly and sparsely from main branches, mostly broken, tapering, branch length 8.73–19.77–
<quantity id="4CFA3D9CFFA3F655FDCEFCAE7EF0FC2E" box="[558,678,894,916]" metricMagnitude="-2" metricUnit="m" metricValue="2.542" pageId="76" pageNumber="753" unit="mm" value="25.42">25.42 mm</quantity>
, angle 29.62–37.00–45.12°, basal branch diameter 7.65–8.52–
<quantity id="4CFA3D9CFFA3F655FF70FC6C7CA5FC6B" box="[144,243,956,977]" metricMagnitude="-3" metricUnit="m" metricValue="9.83" pageId="76" pageNumber="753" unit="mm" value="9.83">9.83 mm</quantity>
, middle branch diameter 4.97–6.67–
<quantity id="4CFA3D9CFFA3F655FD70FC6B7EA4FC6B" box="[656,754,955,977]" metricMagnitude="-3" metricUnit="m" metricValue="8.54" pageId="76" pageNumber="753" unit="mm" value="8.54">8.54 mm</quantity>
, branch tip diameter 3.72–4.44–
<quantity id="4CFA3D9CFFA3F655FDEBFC0A7E27FC4A" box="[523,625,986,1008]" metricMagnitude="-3" metricUnit="m" metricValue="5.48" pageId="76" pageNumber="753" unit="mm" value="5.48">5.48 mm</quantity>
, some portions of branches lacking radial corallites. Overall length of
<typeStatus id="54B92EDBFFA3F655FF4EFBC77D59FB97" box="[174,271,1047,1069]" pageId="76" pageNumber="753" type="holotype">holotype</typeStatus>
<quantity id="4CFA3D9CFFA3F655FEF8FBC77D2BFB97" box="[280,381,1047,1069]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="76" pageNumber="753" unit="mm" value="15.9">15.9 mm</quantity>
, mid branch diameter
<quantity id="4CFA3D9CFFA3F655FD6BFBC77EB5FB97" box="[651,739,1047,1069]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="76" pageNumber="753" unit="mm" value="5.1">5.1 mm</quantity>
.
</paragraph>
<paragraph id="8BBD9079FFA3F655FF48FBE67DBCF9AC" blockId="76.[144,759,803,1650]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF48FBE67D4EFBF1" box="[168,280,1078,1099]" italics="true" pageId="76" pageNumber="753">Corallites.</emphasis>
Axial corallites visible at some branch tips, 0.93–1.05–
<quantity id="4CFA3D9CFFA3F655FEEBFB847D3BFBD0" box="[267,365,1108,1130]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="76" pageNumber="753" unit="mm" value="1.19">1.19 mm</quantity>
exsert, outer diameter 1.12–1.32–
<quantity id="4CFA3D9CFFA3F655FF70FBA37CA6FB33" box="[144,240,1139,1161]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="76" pageNumber="753" unit="mm" value="1.5">1.50 mm</quantity>
, inner diameter 0.52–0.69–
<quantity id="4CFA3D9CFFA3F655FDF9FBA37E2EFB33" box="[537,632,1139,1161]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="76" pageNumber="753" unit="mm" value="0.79">0.79 mm</quantity>
, wall thickness 0.15–0.16–
<quantity id="4CFA3D9CFFA3F655FEA6FB427DFDFB1D" box="[326,427,1170,1192]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="76" pageNumber="753" unit="mm" value="0.16">0.16 mm</quantity>
, primary septa two-thirds R with smooth and straight axial margins, secondary septa up to one-quarter R, S1&gt;&gt;S2; radial corallites dimorphic, tubular appressed towards the branch tips, round to oval calices, profile length 1.18–1.56–
<quantity id="4CFA3D9CFFA3F655FDADFADC7EF9FA98" box="[589,687,1292,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="76" pageNumber="753" unit="mm" value="1.78">1.78 mm</quantity>
, angle 17.94–29.80–41.82°, outer diameter 0.71–0.80–
<quantity id="4CFA3D9CFFA3F655FD72FAFB7EA4FAFA" box="[658,754,1323,1344]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="76" pageNumber="753" unit="mm" value="0.93">0.93 mm</quantity>
, inner diameter 0.34–0.47–
<quantity id="4CFA3D9CFFA3F655FE20FA997E72FAE5" box="[448,548,1353,1375]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="76" pageNumber="753" unit="mm" value="0.56">0.56 mm</quantity>
; mostly immersed radials, not touching, sparsely distributed, inner diameter 0.36–0.42–
<quantity id="4CFA3D9CFFA3F655FE9DFA567DBBFA26" box="[381,493,1414,1436]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="76" pageNumber="753" unit="mm" value="0.5">0.50 mm</quantity>
, angle 35.25–40.69– 46.12°, distance between centres 3.20–4.45–6.00 mm, primary septa up to one-quarter R, secondary septa mostly absent or visible as points. Corallite sequence arrangement 1–[1–3]–[3–6]–?.
</paragraph>
<paragraph id="8BBD9079FFA3F655FF48F9F07E3AF9C8" blockId="76.[144,759,803,1650]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF48F9F07D62F98F" box="[168,308,1568,1589]" italics="true" pageId="76" pageNumber="753">Coenosteum.</emphasis>
Elaborated spinules evenly and densely distributed both on and between radial corallites. Coenosteum amount 2.54–3.83–
<quantity id="4CFA3D9CFFA3F655FDE2F98C7E3EF9C8" box="[514,616,1628,1650]" metricMagnitude="-3" metricUnit="m" metricValue="5.89" pageId="76" pageNumber="753" unit="mm" value="5.89">5.89 mm</quantity>
.
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70F95F7D58F91E" blockId="76.[144,759,1679,1791]" box="[144,270,1679,1700]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF70F95F7D58F91E" box="[144,270,1679,1700]" italics="true" pageId="76" pageNumber="753">Occurrence</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70F97D7CBEF945" blockId="76.[144,759,1679,1791]" pageId="76" pageNumber="753">
<materialsCitation id="3B6A9A24FFA3F655FF70F97D7CB2F945" collectionCode="TF" pageId="76" pageNumber="753" specimenCode="TF502" specimenCount="1">
Late Miocene. Only known as a fossil from the
<typeStatus id="54B92EDBFFA3F655FD25F97D7EA0F978" box="[709,758,1709,1730]" pageId="76" pageNumber="753">type</typeStatus>
locality, outcrop
<specimenCode id="DBA43802FFA3F655FEA4F91B7DD9F95B" box="[324,399,1739,1761]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="76" pageNumber="753">TF502</specimenCode>
, Bontang, of Tortonian age, 9.4– 9.8 Ma
</materialsCitation>
.
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70F8CC7D3CF88B" blockId="76.[144,759,1820,1903]" box="[144,362,1820,1841]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FF70F8CC7D3CF88B" box="[144,362,1820,1841]" italics="true" pageId="76" pageNumber="753">Palaeoenvironment</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FF70F8EA7F9CFEEC" blockId="76.[144,759,1820,1903]" lastBlockId="76.[806,1421,198,342]" pageId="76" pageNumber="753">
<taxonomicName id="4C02EBFAFFA3F655FF70F8EA7D05F8EA" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" box="[144,339,1850,1872]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">
<emphasis id="B9764C6BFFA3F655FF70F8EA7D05F8EA" box="[144,339,1850,1872]" italics="true" pageId="76" pageNumber="753">Acropora renemai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245F110FFA3F655FEBBF8EB7DFBF8EA" box="[347,429,1851,1872]" pageId="76" pageNumber="753" rank="species">sp. nov.</taxonomicNameLabel>
was found within a grey silty to sand-rich matrix together with a large phacelloid colony of
<taxonomicName id="4C02EBFAFFA3F655FC76FF167877FF61" authorityName="Dana" authorityYear="1846" box="[918,1057,198,219]" class="Anthozoa" family="Merulinidae" genus="Caulastraea" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFA3F655FC76FF167877FF61" box="[918,1057,198,219]" italics="true" pageId="76" pageNumber="753">Caulastraea</emphasis>
</taxonomicName>
. Other small massive corals of the genera
<taxonomicName id="4C02EBFAFFA3F655FC41FF357877FF40" authorityName="de Blainville" authorityYear="1830" box="[929,1057,229,250]" class="Anthozoa" family="Merulinidae" genus="Dipsastraea" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFA3F655FC41FF357877FF40" box="[929,1057,229,250]" italics="true" pageId="76" pageNumber="753">Dipsastraea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C02EBFAFFA3F655FBCFFF35782BFF40" authorityName="Link" authorityYear="1807" box="[1071,1149,229,250]" class="Anthozoa" family="Merulinidae" genus="Favites" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFA3F655FBCFFF35782BFF40" box="[1071,1149,229,250]" italics="true" pageId="76" pageNumber="753">Favites</emphasis>
</taxonomicName>
and
<taxonomicName id="4C02EBFAFFA3F655FB53FF34794DFF43" box="[1203,1307,228,249]" class="Anthozoa" family="Merulinidae" genus="Platygyra" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFA3F655FB53FF34794DFF43" box="[1203,1307,228,249]" italics="true" pageId="76" pageNumber="753">Platygyra</emphasis>
</taxonomicName>
also occur in the same facies. This environment is interpreted as a shallow-water environment with calm waters, probably a lagoon.
</paragraph>
<paragraph id="8BBD9079FFA3F655FCC6FE547FDDFE23" blockId="76.[806,1422,388,1144]" box="[806,907,388,409]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FCC6FE547FDDFE23" box="[806,907,388,409]" italics="true" pageId="76" pageNumber="753">Remarks</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FCC6FE737FBFFBC2" blockId="76.[806,1422,388,1144]" pageId="76" pageNumber="753">
Branches recovered show only primary branching order, and thus a corymbose or caespito-corymbose form is inferred for this species. We have neither photographic evidence nor field notes to confirm that all recovered fragments belong to the same colony, even though they are from the same sample. For that reason, only one fragment was designated as the
<typeStatus id="54B92EDBFFA3F655FB5CFD8A794FFDCA" box="[1212,1305,602,624]" pageId="76" pageNumber="753" type="holotype">holotype</typeStatus>
(NHMUK PI AZ7439), and the rest of the specimens as
<typeStatus id="54B92EDBFFA3F655FAC0FDA979D8FD34" box="[1312,1422,633,654]" pageId="76" pageNumber="753" type="paratype">paratypes</typeStatus>
(NHMUK PI AZ9117). Among extant species
<taxonomicName id="4C02EBFAFFA3F655FAC7FD487FD7FD76" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">
<emphasis id="B9764C6BFFA3F655FAC7FD487FD7FD76" italics="true" pageId="76" pageNumber="753">Acropora renemai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245F110FFA3F655FC69FD667F88FD71" box="[905,990,694,715]" pageId="76" pageNumber="753" rank="species">sp. nov.</taxonomicNameLabel>
closely resembles
<taxonomicName id="4C02EBFAFFA3F655FB50FD667FA0FD51" authority="(Ehrenberg, 1874)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1874" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="squarrosa">
<emphasis id="B9764C6BFFA3F655FB50FD6679D8FD76" box="[1200,1422,694,716]" italics="true" pageId="76" pageNumber="753">Acropora squarrosa</emphasis>
(Ehrenberg, 1874)
</taxonomicName>
from the
<collectingRegion id="49C65E9BFFA3F655FB8AFD05789DFD50" box="[1130,1227,725,747]" country="Egypt" name="Al Bahr al Ahmar" pageId="76" pageNumber="753">Red Sea</collectingRegion>
, and therefore it is placed into the
<taxonomicName id="4C02EBFAFFA3F655FC05FD23787AFCB2" box="[997,1068,755,776]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="loripes">
<emphasis id="B9764C6BFFA3F655FC05FD23787AFCB2" box="[997,1068,755,776]" italics="true" pageId="76" pageNumber="753">loripes</emphasis>
</taxonomicName>
species group.
<taxonomicName id="4C02EBFAFFA3F655FB2FFD2379DBFCB3" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" box="[1231,1421,755,777]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">
<emphasis id="B9764C6BFFA3F655FB2FFD2379DBFCB3" box="[1231,1421,755,777]" italics="true" pageId="76" pageNumber="753">Acropora renemai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245F110FFA3F655FCC6FCC27F2BFC9D" box="[806,893,786,807]" pageId="76" pageNumber="753" rank="species">sp. nov.</taxonomicNameLabel>
also resembles
<taxonomicName id="4C02EBFAFFA3F655FBD7FCC279DBFC9D" authority="Veron, 2000" authorityName="Veron" authorityYear="2000" box="[1079,1421,786,807]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="maryae">
<emphasis id="B9764C6BFFA3F655FBD7FCC278ADFC9D" box="[1079,1275,786,807]" italics="true" pageId="76" pageNumber="753">Acropora maryae</emphasis>
<bibRefCitation id="EF93ED88FFA3F655FAE4FCC279DBFC9D" author="Veron JEN" box="[1284,1421,786,807]" pageId="76" pageNumber="753" refId="ref53488" refString="Veron JEN. 2000. Corals of the world. Townsville: Australian Institute of Marine Science." type="book" year="2000">Veron, 2000</bibRefCitation>
</taxonomicName>
from the
<collectingRegion id="49C65E9BFFA3F655FC74FCE07FA5FCFF" box="[916,1011,816,838]" country="Egypt" name="Al Bahr al Ahmar" pageId="76" pageNumber="753">Red Sea</collectingRegion>
synonymized with
<taxonomicName id="4C02EBFAFFA3F655FB38FCE1793EFCFC" baseAuthorityName="Ehrenberg" baseAuthorityYear="1874" box="[1240,1384,817,838]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="squarrosa">
<emphasis id="B9764C6BFFA3F655FB38FCE1793EFCFC" box="[1240,1384,817,838]" italics="true" pageId="76" pageNumber="753">A. squarrosa</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF93ED88FFA3F655FCC6FC9F7874FCDF" author="Wallace CC &amp; Done BJ &amp; Muir PR" box="[806,1058,847,869]" pageId="76" pageNumber="753" pagination="1 - 255" refId="ref54139" refString="Wallace CC, Done BJ, Muir PR. 2012. Revision and catalogue of worldwide staghorn corals Acropora and Isopora (Scleractinia: Acroporidae) in the Museum of Tropical Queensland. Memoirs of the Queensland Museum 57: 1 - 255." type="journal article" year="2012">
Wallace
<emphasis id="B9764C6BFFA3F655FC6EFC807F97FCDE" box="[910,961,847,869]" italics="true" pageId="76" pageNumber="753">et al</emphasis>
. (2012)
</bibRefCitation>
. According to
<bibRefCitation id="EF93ED88FFA3F655FB34FC9F79DEFCDF" author="Wallace CC" box="[1236,1416,847,869]" pageId="76" pageNumber="753" refId="ref53823" refString="Wallace CC. 1999. Staghorn corals of the world: a revision of the coral Genus Acropora (Scleractinia; Astrocoeniina; Acroporidae) worldwide, with emphasis on morphology, phylogeny and biogeography. Collingwood: CSIRO Publishing." type="book" year="1999">Wallace (1999)</bibRefCitation>
,
<taxonomicName id="4C02EBFAFFA3F655FCC6FCBE7FE0FC39" baseAuthorityName="Ehrenberg" baseAuthorityYear="1874" box="[806,950,878,899]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="squarrosa">
<emphasis id="B9764C6BFFA3F655FCC6FCBE7FE0FC39" box="[806,950,878,899]" italics="true" pageId="76" pageNumber="753">A. squarrosa</emphasis>
</taxonomicName>
is restricted to the
<collectingRegion id="49C65E9BFFA3F655FB7EFCBD78AAFC39" box="[1182,1276,877,899]" country="Egypt" name="Al Bahr al Ahmar" pageId="76" pageNumber="753">Red Sea</collectingRegion>
today. Comparisons with the
<typeStatus id="54B92EDBFFA3F655FBEDFC5C782EFC18" box="[1037,1144,908,930]" pageId="76" pageNumber="753" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C02EBFAFFA3F655FB46FC5C79DAFC18" authority="NHMUK" authorityName="NHMUK" box="[1190,1420,908,930]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="loripes">
<emphasis id="B9764C6BFFA3F655FB46FC5C7940FC1B" box="[1190,1302,908,929]" italics="true" pageId="76" pageNumber="753">A. loripes</emphasis>
NHMUK
</taxonomicName>
1892.6.8.219 from the Great Barrier Reef show that corallites in
<taxonomicName id="4C02EBFAFFA3F655FC56FC1A7866FC65" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" box="[950,1072,970,991]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">
<emphasis id="B9764C6BFFA3F655FC56FC1A7866FC65" box="[950,1072,970,991]" italics="true" pageId="76" pageNumber="753">A. renemai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245F110FFA3F655FBD9FC1A78D8FC65" box="[1081,1166,970,991]" pageId="76" pageNumber="753" rank="species">sp. nov.</taxonomicNameLabel>
are smaller and more scattered, and branches seem to be more delicate. Elaborated spinules of
<taxonomicName id="4C02EBFAFFA3F655FC12FBD7783BFBA6" authority="Santodomingo &amp; Wallace &amp; Johnson, 2015" authorityName="Santodomingo &amp; Wallace &amp; Johnson" authorityYear="2015" box="[1010,1133,1031,1052]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="renemai" status="sp. nov.">
<emphasis id="B9764C6BFFA3F655FC12FBD7783BFBA6" box="[1010,1133,1031,1052]" italics="true" pageId="76" pageNumber="753">A. renemai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245F110FFA3F655FB97FBD77898FBA6" box="[1143,1230,1031,1052]" pageId="76" pageNumber="753" rank="species">sp. nov.</taxonomicNameLabel>
are also similar to
<taxonomicName id="4C02EBFAFFA3F655FCA9FBF57FEEFB80" box="[841,952,1061,1082]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="loripes">
<emphasis id="B9764C6BFFA3F655FCA9FBF57FEEFB80" box="[841,952,1061,1082]" italics="true" pageId="76" pageNumber="753">A. loripes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C02EBFAFFA3F655FC1DFBF578C4FB81" baseAuthorityName="Ehrenberg" baseAuthorityYear="1874" box="[1021,1170,1061,1083]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="squarrosa">
<emphasis id="B9764C6BFFA3F655FC1DFBF578C4FB81" box="[1021,1170,1061,1083]" italics="true" pageId="76" pageNumber="753">A. squarrosa</emphasis>
</taxonomicName>
. This species represents the earliest occurrence of the
<taxonomicName id="4C02EBFAFFA3F655FB33FB947949FBE3" box="[1235,1311,1092,1113]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="76" pageNumber="753" phylum="Cnidaria" rank="species" species="loripes">
<emphasis id="B9764C6BFFA3F655FB33FB947949FBE3" box="[1235,1311,1092,1113]" italics="true" pageId="76" pageNumber="753">loripes</emphasis>
</taxonomicName>
group in the fossil record.
</paragraph>
<paragraph id="8BBD9079FFA3F655FCC6FB767FCBFB01" blockId="76.[806,1421,1190,1334]" box="[806,925,1190,1211]" pageId="76" pageNumber="753">
<emphasis id="B9764C6BFFA3F655FCC6FB767FCBFB01" box="[806,925,1190,1211]" italics="true" pageId="76" pageNumber="753">Etymology</emphasis>
</paragraph>
<paragraph id="8BBD9079FFA3F655FCC6FB157802FA8C" blockId="76.[806,1421,1190,1334]" pageId="76" pageNumber="753">
This species is named after Willem Renema for his contribution to the palaeontological and biological studies of the geology, stratigraphy and ecology of larger benthic foraminifera of
<collectingCountry id="F315D0E9FFA3F655FC3DFAF07806FA8C" box="[989,1104,1312,1334]" name="Indonesia" pageId="76" pageNumber="753">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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