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<mods:title id="456197991D758A445A148E9B66F58DA8">Taxonomy of ' Euconnus complex'. Part XVII. Status of subgenera defined by male antennal characters: Androconnus Franz and Cladoconnus Reitter (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C316652D8B5078078BC85DFEFCBFFBE5" box="[151,852,1014,1040]" pageId="1" pageNumber="370" type="nomenclature">
<paragraph id="8BB336A68B5078078BC85DFEFCBFFBE5" blockId="1.[151,852,1014,1040]" box="[151,852,1014,1040]" pageId="1" pageNumber="370">
<heading id="D0FB81CA8B5078078BC85DFEFCBFFBE5" bold="true" box="[151,852,1014,1040]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
<emphasis id="B978EAB48B5078078BC85DFEFCBFFBE5" bold="true" box="[151,852,1014,1040]" pageId="1" pageNumber="370">
1. Morphological structures of 
<taxonomicName id="4C0C4D258B50780789465DFEFCBFFBE5" ID-CoL="8XRFM" authority="" box="[537,852,1014,1040]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="subGenus" subGenus="Androconnus">
<emphasis id="B978EAB48B50780789465DFEFD7EFBE5" bold="true" box="[537,661,1014,1040]" italics="true" pageId="1" pageNumber="370">Euconnus</emphasis>
(
<emphasis id="B978EAB48B50780789FA5DFEFCA2FBE5" bold="true" box="[677,841,1014,1040]" italics="true" pageId="1" pageNumber="370">Androconnus</emphasis>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316652D8B5078078BC85A35FE40FB6B" pageId="1" pageNumber="370" type="discussion">
<paragraph id="8BB336A68B5078078BC85A35FE40FB6B" blockId="1.[151,1437,1085,2010]" pageId="1" pageNumber="370">
The general body shape of 
<taxonomicName id="4C0C4D258B5078078A835A36FD20FBA3" authorityName="Franz" authorityYear="1986" box="[476,715,1085,1110]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="bechyneae">
<emphasis id="B978EAB48B5078078A835A36FD20FBA3" box="[476,715,1085,1110]" italics="true" pageId="1" pageNumber="370">Euconnus bechyneae</emphasis>
</taxonomicName>
(
<figureCitation id="13372A238B50780789805A35FCC8FBA3" box="[735,803,1085,1110]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1305,1328]" captionTargetBox="[161,1423,352,1279]" captionTargetId="figure@2.[151,1436,337,1290]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus. Euconnus (Androconnus) bechyneae Franz, male (1); Euconnus (Cladoconnus) motschulskyi (Sturm), male (2); Euconnus (Cladoconnus) denticornis (Müller &amp; Kunze), female (3)." httpUri="https://zenodo.org/record/1242547/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
) does not deviate from that in many other species of 
<taxonomicName id="4C0C4D258B5078078BC85A6AFEEEFB8C" authorityName="Thomson" authorityYear="1859" box="[151,261,1122,1145]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5078078BC85A6AFEEEFB8C" box="[151,261,1122,1145]" italics="true" pageId="1" pageNumber="370">Euconnus</emphasis>
</taxonomicName>
; it is slender, moderately convex, with deep constrictions between the head and pronotum and between the pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="C316652D8B5078058B985AA1FC87F973" lastPageId="3" lastPageNumber="372" pageId="1" pageNumber="370" type="description">
<paragraph id="8BB336A68B5078078B985AA1FCECFA6F" blockId="1.[151,1437,1085,2010]" pageId="1" pageNumber="370">
The head capsule (
<figureCitation id="13372A238B5078078AF75AA1FE1CFB37" box="[424,503,1193,1218]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Figs 4</figureCitation>
̄5) is divided into the posterior 'neck' region and anterior, exposed part by an occipital constriction subequal to half width of the head. The anterior part of the head is slightly elongate; the vertex demarcated from the 'neck' region by a transverse impression and very weakly bulging posterodorsad; the frons subtrapezoidal, transverse; antennal insertions moderately broadly separated; composite eyes located anterolaterally; tempora much longer than eyes, rounded and distinctly convergent caudad. The tempora and genae/ postgenae are sparsely covered with long, thick bristles; additionally, there are two groups of 2¯3 modified, flattened and distally broadened setae on the postgenae.
</paragraph>
<paragraph id="8BB336A68B5078078B985BADFAACF984" blockId="1.[151,1437,1085,2010]" pageId="1" pageNumber="370">
Mouthparts (
<figureCitation id="13372A238B5078078A075BADFE48FA4B" box="[344,419,1445,1470]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Figs 4</figureCitation>
̄5) similar to those in most species of 
<taxonomicName id="4C0C4D258B50780788045BAEFC22FA48" authorityName="Thomson" authorityYear="1859" box="[859,969,1446,1469]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B50780788045BAEFC22FA48" box="[859,969,1446,1469]" italics="true" pageId="1" pageNumber="370">Euconnus</emphasis>
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; mandibles subtriangular, evenly curved, lacking mesal teeth; mentum (
<figureCitation id="13372A238B5078078AB85BC1FDC0FA17" box="[487,555,1481,1506]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B50780789685BC2FDB2FA14" box="[567,601,1482,1505]" italics="true" pageId="1" pageNumber="370">mn</emphasis>
) subhexagonal with narrowly separated and unremarkable labial palps and a small elongate ligula bearing a pair of short setae between the palps; submentum (
<figureCitation id="13372A238B5078078FCF5BE5FB31F9F3" box="[1168,1242,1517,1542]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B5078078FB35BE6FAF2F9F0" box="[1260,1305,1518,1541]" italics="true" pageId="1" pageNumber="370">smn</emphasis>
) short and transverse; hypostomal ridges (
<figureCitation id="13372A238B5078078AA45819FDA9F9DF" box="[507,578,1553,1578]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B507807890F5819FD82F9DF" box="[592,617,1553,1578]" italics="true" pageId="1" pageNumber="370">hr</emphasis>
) long and recurved, but posteriorly not reaching posterior tentorial pits (
<figureCitation id="13372A238B5078078BC0583DFF0FF9BB" box="[159,228,1589,1614]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B5078078BAD583EFEFDF9B8" box="[242,278,1590,1613]" italics="true" pageId="1" pageNumber="370">ptp</emphasis>
), which are located in front of a transverse impression demarcating the 'neck' region ventrally, the pits are elongate, slot-like. Maxillae and maxillary palpomeres typical as in 
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<emphasis id="B978EAB48B50780788E65852FBCCF984" box="[953,1063,1626,1649]" italics="true" pageId="1" pageNumber="370">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B5078078F705852FB87F984" box="[1071,1132,1626,1649]" pageId="1" pageNumber="370" sensu="stricto">s. str.</taxonomicNameLabel>
; labrum transverse.
</paragraph>
<paragraph id="8BB336A68B5078078B985875FF04F864" blockId="1.[151,1437,1085,2010]" pageId="1" pageNumber="370">
The antennae (
<figureCitation id="13372A238B5078078A325875FE5FF963" box="[365,436,1661,1686]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1305,1328]" captionTargetBox="[161,1423,352,1279]" captionTargetId="figure@2.[151,1436,337,1290]" captionTargetPageId="2" captionText="FIGURES 1–3. Dorsal habitus. Euconnus (Androconnus) bechyneae Franz, male (1); Euconnus (Cladoconnus) motschulskyi (Sturm), male (2); Euconnus (Cladoconnus) denticornis (Müller &amp; Kunze), female (3)." httpUri="https://zenodo.org/record/1242547/files/figure.png" pageId="1" pageNumber="370">Figs 1</figureCitation>
, 
<figureCitation id="13372A238B5078078A9D5875FE38F963" box="[450,467,1661,1686]" captionStart="FIGURES6–8" captionStartId="5.[151,264,1581,1604]" captionTargetBox="[151,1436,375,1560]" captionTargetId="figure@5.[151,1436,375,1560]" captionTargetPageId="5" captionText="FIGURES6–8. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male.Distal portion of left antenna in dorsal (6) and mesal (7) views; antennomeres X–XI in mesal view (8). Abbreviations:anIX–XI,antennomere IX–XI." httpUri="https://zenodo.org/record/1242551/files/figure.png" pageId="1" pageNumber="370">6</figureCitation>
¯8) are slender and strikingly long in relation to the body; in both sexes antennomeres VIĪXI form an indistinctly delimited, pentamerous club, which in males is broader than in females, and has three terminal antennomeres modified. In the 
<typeStatus id="54B788048B507807893958CEFD7DF92B" box="[614,662,1734,1758]" pageId="1" pageNumber="370">type</typeStatus>
species of 
<taxonomicName id="4C0C4D258B507807884358CDFC5BF92B" authorityName="Franz" authorityYear="1986" box="[796,944,1733,1758]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B507807884358CDFC5BF92B" box="[796,944,1733,1758]" italics="true" pageId="1" pageNumber="370">Androconnus</emphasis>
</taxonomicName>
, antennomeres IX and X have their mesal portions flattened or impressed, and the anteromesal margin of each forms a subtriangular projection. In the antennomere X (
<figureCitation id="13372A238B5078078A3B5905FE5BF8D3" box="[356,432,1805,1830]" captionStart="FIGURES6–8" captionStartId="5.[151,264,1581,1604]" captionTargetBox="[151,1436,375,1560]" captionTargetId="figure@5.[151,1436,375,1560]" captionTargetPageId="5" captionText="FIGURES6–8. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male.Distal portion of left antenna in dorsal (6) and mesal (7) views; antennomeres X–XI in mesal view (8). Abbreviations:anIX–XI,antennomere IX–XI." httpUri="https://zenodo.org/record/1242551/files/figure.png" pageId="1" pageNumber="370">Fig. 8</figureCitation>
), there is a glandular opening at base of the projection. The antennomere XI is asymmetrically impressed dorsally and dorsomesally. Apical portions of antennomeres VIĪX are covered with irregular granulation, and impressed portions of antennomere XI have scale-like microsculpture and are nearly asetose.
</paragraph>
<paragraph id="8BB336A68B5078048B985995FE25FEE8" blockId="1.[151,1437,1085,2010]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
The prothorax (
<figureCitation id="13372A238B5078078A255995FE2DF843" box="[378,454,1949,1974]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="1" pageNumber="370">Figs 4</figureCitation>
̄5) of 
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<emphasis id="B978EAB48B507807894B5995FD43F843" box="[532,680,1949,1974]" italics="true" pageId="1" pageNumber="370">Androconnus</emphasis>
</taxonomicName>
is bell-shaped, slightly elongate, broadest slightly behind middle, with rounded sides and more narrowing anteriorly than posteriorly, so that the anterior pronotal margin is shorter than posterior margin; anterior and posterior corners well-defined but obtuse-angled and blunt. The pronotal base bears two pairs of very small and shallow antebasal pits (
<figureCitation id="13372A238B53780488685EB4FC6BFF21" box="[823,896,188,213]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
; 
<emphasis id="B978EAB48B53780488D05EB4FC52FF20" box="[911,953,188,213]" italics="true" pageId="2" pageNumber="371">abp</emphasis>
), narrow but distinct transverse antebasal groove (
<figureCitation id="13372A238B5378048BAA5ED7FED7FF0D" box="[245,316,223,248]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
; 
<emphasis id="B978EAB48B5378048A165ED7FE98FF0D" box="[329,371,223,248]" italics="true" pageId="2" pageNumber="371">abg</emphasis>
), and short sublateral carinae (
<figureCitation id="13372A238B537804898C5ED7FCF2FF0D" box="[723,793,223,248]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
; 
<emphasis id="B978EAB48B53780488795ED7FCAEFF0D" box="[806,837,223,248]" italics="true" pageId="2" pageNumber="371">slc</emphasis>
). In addition to thin setae, pronotum bears thick and sparse bristles on each side.
</paragraph>
<caption id="DF73662E8B5378048BC85B11FBECFABB" httpUri="https://zenodo.org/record/1242547/files/figure.png" pageId="2" pageNumber="371" startId="2.[151,264,1305,1328]" targetBox="[161,1423,352,1279]" targetPageId="2">
<paragraph id="8BB336A68B5378048BC85B11FBECFABB" blockId="2.[151,1436,1305,1358]" pageId="2" pageNumber="371">
<emphasis id="B978EAB48B5378048BC85B11FED0FADB" bold="true" box="[151,315,1305,1328]" pageId="2" pageNumber="371">FIGURES 1–3.</emphasis>
Dorsal habitus. 
<taxonomicName id="4C0C4D258B5378048ABA5B12FC75FADA" authority="Franz" authorityName="Franz" authorityYear="1986" box="[485,926,1305,1327]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="bechyneae" subGenus="Androconnus">
<emphasis id="B978EAB48B5378048ABA5B12FDA2FADA" box="[485,585,1306,1327]" italics="true" pageId="2" pageNumber="371">Euconnus</emphasis>
(
<emphasis id="B978EAB48B53780489055B11FD0BFADA" box="[602,736,1305,1327]" italics="true" pageId="2" pageNumber="371">Androconnus</emphasis>
) 
<emphasis id="B978EAB48B53780489AF5B11FCB0FADA" box="[752,859,1305,1327]" italics="true" pageId="2" pageNumber="371">bechyneae</emphasis>
Franz
</taxonomicName>
, male (1); 
<taxonomicName id="4C0C4D258B5378048F505B12FF0CFABB" authority="(Sturm)" authorityName="Sturm" baseAuthorityName="Sturm" baseAuthorityYear="1838" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="motschulskyi" subGenus="Cladoconnus">
<emphasis id="B978EAB48B5378048F505B12FB98FADA" box="[1039,1139,1306,1327]" italics="true" pageId="2" pageNumber="371">Euconnus</emphasis>
(
<emphasis id="B978EAB48B5378048FDC5B11FAE2FADA" box="[1155,1289,1305,1327]" italics="true" pageId="2" pageNumber="371">Cladoconnus</emphasis>
) 
<emphasis id="B978EAB48B5378048E465B11FA77FADA" box="[1305,1436,1305,1327]" italics="true" pageId="2" pageNumber="371">motschulskyi</emphasis>
(Sturm)
</taxonomicName>
, male (2); 
<taxonomicName id="4C0C4D258B5378048A0B5B31FC65FABB" authority="(Muller &amp; Kunze)" authorityName="Muller &amp; Kunze" baseAuthorityName="Muller &amp; Kunze" baseAuthorityYear="1822" box="[340,910,1336,1358]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="denticornis" subGenus="Cladoconnus">
<emphasis id="B978EAB48B5378048A0B5B31FE53FABB" box="[340,440,1337,1358]" italics="true" pageId="2" pageNumber="371">Euconnus</emphasis>
(
<emphasis id="B978EAB48B5378048A985B30FDA7FABB" box="[455,588,1336,1358]" italics="true" pageId="2" pageNumber="371">Cladoconnus</emphasis>
) 
<emphasis id="B978EAB48B53780489045B30FD26FABB" box="[603,717,1336,1358]" italics="true" pageId="2" pageNumber="371">denticornis</emphasis>
(Müller &amp; Kunze)
</taxonomicName>
, female (3).
</paragraph>
</caption>
<paragraph id="8BB336A68B5378048B985B73FE37FA0A" blockId="2.[151,1437,1403,2004]" pageId="2" pageNumber="371">
Ventral prothoracic structures are typical of 
<taxonomicName id="4C0C4D258B53780489E85B74FCCEFA66" authorityName="Jałoszyński" authorityYear="2018" box="[695,805,1404,1427]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B53780489E85B74FCCEFA66" box="[695,805,1404,1427]" italics="true" pageId="2" pageNumber="371">Euconnus</emphasis>
</taxonomicName>
: the basisternal part of prosternum (
<figureCitation id="13372A238B5378048FE05B73FAEFFA61" box="[1215,1284,1403,1428]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B5378048E4D5B73FAD9FA61" box="[1298,1330,1403,1428]" italics="true" pageId="2" pageNumber="371">bst</emphasis>
) is much shorter than the coxal part; the intercoxal region lacks process or carina; notosternal sutures (
<figureCitation id="13372A238B5378048FB85B96FADBFA42" box="[1255,1328,1438,1463]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B5378048E1E5BA8FA8EFA42" box="[1345,1381,1440,1463]" italics="true" pageId="2" pageNumber="371">nss</emphasis>
) are complete; hypomeral ridges (
<figureCitation id="13372A238B5378048AB85BCBFDC5FA29" box="[487,558,1475,1500]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
; 
<emphasis id="B978EAB48B53780489625BCBFD89FA29" box="[573,610,1475,1500]" italics="true" pageId="2" pageNumber="371">hyr</emphasis>
) complete and the inner (adcoxal) part of each hypomeron is distinctly microsculptured and asetose.
</paragraph>
<paragraph id="8BB336A68B5378048B985803FBBAF97A" blockId="2.[151,1437,1403,2004]" pageId="2" pageNumber="371">
The mesoventrite (
<figureCitation id="13372A238B5378048AC65803FE37F9D6" box="[409,476,1547,1572]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
) has a broad and well-defined anterior ridge, distinct anterior impressions functioning as procoxal rests (
<figureCitation id="13372A238B5378048A005826FE48F9B2" box="[351,419,1582,1607]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
; 
<emphasis id="B978EAB48B5378048AF05838FE3FF9B2" box="[431,468,1584,1607]" italics="true" pageId="2" pageNumber="371">pcr</emphasis>
), which are sharply demarcated anteriorly and separated at middle, but not demarcated posteriorly, and filled with sparse, short setae. The mesoventral intercoxal process (
<figureCitation id="13372A238B5378048F32585BFB5DF99E" box="[1133,1206,1619,1644]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
; 
<emphasis id="B978EAB48B5378048F99585CFAEBF99E" box="[1222,1280,1620,1643]" italics="true" pageId="2" pageNumber="371">msvp</emphasis>
) is keel-like, anteriorly connected to the anterior ridge and posteriorly fused with the metaventrite.
</paragraph>
<paragraph id="8BB336A68B5378048B985893FC63F946" blockId="2.[151,1437,1403,2004]" box="[199,904,1691,1716]" pageId="2" pageNumber="371">
The mesoscutellum is not exposed in intact specimens (
<figureCitation id="13372A238B53780488695893FC90F946" box="[822,891,1691,1716]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BB336A68B5378048B9858B6FA93F909" blockId="2.[151,1437,1403,2004]" pageId="2" pageNumber="371">
The metaventrite (
<figureCitation id="13372A238B5378048AC858B6FE37F922" box="[407,476,1726,1751]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
) is slightly transverse; the metaventral intercoxal process (
<figureCitation id="13372A238B5378048F2458B6FB2AF922" box="[1147,1217,1726,1751]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
; 
<emphasis id="B978EAB48B5378048F9058C8FAEEF922" box="[1231,1285,1728,1751]" italics="true" pageId="2" pageNumber="371">mtvp</emphasis>
) is relatively narrow but distinctly separating metacoxae, its posterior margin is concave, and posterolateral corners rounded.
</paragraph>
<paragraph id="8BB336A68B5378048B98590EFB1BF8EA" blockId="2.[151,1437,1403,2004]" box="[199,1264,1798,1823]" pageId="2" pageNumber="371">
Each elytron bears two asetose, small and shallow but distinct basal elytral foveae (
<figureCitation id="13372A238B5378048F32590EFB59F8EA" box="[1133,1202,1798,1823]" captionStart="FIGURES 4–5" captionStartId="3.[151,264,1383,1406]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@3.[151,1436,193,1362]" captionTargetPageId="3" captionText="FIGURES 4–5. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
; 
<emphasis id="B978EAB48B5378048FE2590EFB34F8EA" box="[1213,1247,1798,1823]" italics="true" pageId="2" pageNumber="371">bef</emphasis>
).
</paragraph>
<paragraph id="8BB336A68B5378048B985923FDECF821" blockId="2.[151,1437,1403,2004]" pageId="2" pageNumber="371">
The aedeagus in the 
<typeStatus id="54B788048B5378048AE95924FE0DF8B1" box="[438,486,1836,1860]" pageId="2" pageNumber="371">type</typeStatus>
species of 
<taxonomicName id="4C0C4D258B53780489355923FD15F8B1" authorityName="Franz" authorityYear="1986" box="[618,766,1835,1860]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B53780489355923FD15F8B1" box="[618,766,1835,1860]" italics="true" pageId="2" pageNumber="371">Androconnus</emphasis>
</taxonomicName>
(
<figureCitation id="13372A238B53780488505923FC82F8B6" box="[783,873,1835,1860]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1414,1437]" captionTargetBox="[151,1436,265,1393]" captionTargetId="figure@6.[151,1436,265,1393]" captionTargetPageId="6" captionText="FIGURES 9–12. Morphological structures of Euconnus (Androconnus) bechyneae Franz, male. Pterothorax and anterior portion ofabdomen inventral view (9); aedeagusinlateroventral (10), ventral (11) and lateral(12)views.Abbreviations:msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." httpUri="https://zenodo.org/record/1242553/files/figure.png" pageId="2" pageNumber="371">Figs 10</figureCitation>
̄12) is elongate, moderately strongly sclerotized, with free parameres bearing apical and subapical setae and two pairs of apical (dorsal and ventral) projections. Aedeagi illustrated by 
<bibRefCitation id="EF9D4B578B5378048AC5597CFDD1F879" author="Franz, H." box="[410,570,1907,1932]" journalOrPublisher="Koleopterologische Rundschau" pageId="2" pageNumber="371" pagination="3 – 42" part="58" refId="ref4336" refString="Franz, H. (1986 a) Monographie der Scydmaeniden Venezuelas I. Teil (Genera Scydmaenus Latreille, Neuraphanax Reitter, Protoconnus Franz, Homoconnus Schaufuss und Euconnus Rhomboconnus subg. nov. sowie Androconnus subg. nov.). Koleopterologische Rundschau, 58, 3 - 42." title="Monographie der Scydmaeniden Venezuelas I. Teil (Genera Scydmaenus Latreille, Neuraphanax Reitter, Protoconnus Franz, Homoconnus Schaufuss und Euconnus Rhomboconnus subg. nov. sowie Androconnus subg. nov.)" type="journal article" year="1986">Franz (1986a)</bibRefCitation>
for other species of 
<taxonomicName id="4C0C4D258B5378048877597BFC57F879" authorityName="Franz" authorityYear="1986" box="[808,956,1907,1932]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5378048877597BFC57F879" box="[808,956,1907,1932]" italics="true" pageId="2" pageNumber="371">Androconnus</emphasis>
</taxonomicName>
are variable in form, some have similarly paired distal projections as those in 
<taxonomicName id="4C0C4D258B53780489705990FD2DF85A" authorityName="Franz" authorityYear="1986" box="[559,710,1942,1967]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="bechyneae">
<emphasis id="B978EAB48B53780489705990FD2DF85A" box="[559,710,1942,1967]" italics="true" pageId="2" pageNumber="371">E. bechyneae</emphasis>
</taxonomicName>
, and some have subtriangular, undivided apices; endophalli can be symmetrical or asymmetrical.
</paragraph>
<caption id="DF73662E8B5278058BC85B6FFE6AFA0D" httpUri="https://zenodo.org/record/1242549/files/figure.png" pageId="3" pageNumber="372" startId="3.[151,264,1383,1406]" targetBox="[151,1436,193,1362]" targetPageId="3">
<paragraph id="8BB336A68B5278058BC85B6FFE6AFA0D" blockId="3.[151,1436,1383,1528]" pageId="3" pageNumber="372">
<emphasis id="B978EAB48B5278058BC85B6FFED6FA89" bold="true" box="[151,317,1383,1406]" pageId="3" pageNumber="372">FIGURES 4–5.</emphasis>
Morphological structures of 
<taxonomicName id="4C0C4D258B527805892F5B61FBC5FA88" authority="Franz" authorityName="Franz" authorityYear="1986" box="[624,1070,1383,1406]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="bechyneae" subGenus="Androconnus">
<emphasis id="B978EAB48B527805892F5B61FD3FFA8B" box="[624,724,1385,1406]" italics="true" pageId="3" pageNumber="372">Euconnus</emphasis>
(
<emphasis id="B978EAB48B52780589B95B6FFC87FA88" box="[742,876,1383,1405]" italics="true" pageId="3" pageNumber="372">Androconnus</emphasis>
) 
<emphasis id="B978EAB48B52780588215B6FFC02FA88" box="[894,1001,1383,1405]" italics="true" pageId="3" pageNumber="372">bechyneae</emphasis>
Franz
</taxonomicName>
, male. Head, prothorax and elytral base in dorsal view (4); head, prothorax and anterior portion of pterothorax in ventral view (5). Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; nss, notosternal suture; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex.
</paragraph>
</caption>
<paragraph id="8BB336A68B5278058B98582CFC87F973" blockId="3.[151,1436,1572,1705]" pageId="3" pageNumber="372">
<emphasis id="B978EAB48B5278058B98582CFEB4F9C8" bold="true" box="[199,351,1572,1597]" pageId="3" pageNumber="372">Conclusions.</emphasis>
<taxonomicName id="4C0C4D258B5278058A39582CFE11F9C8" authorityName="Franz" authorityYear="1986" box="[358,506,1572,1597]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058A39582CFE11F9C8" box="[358,506,1572,1597]" italics="true" pageId="3" pageNumber="372">Androconnus</emphasis>
</taxonomicName>
has all the diagnostic characters of 
<taxonomicName id="4C0C4D258B52780588CC582EFBEAF9C8" authorityName="Jałoszyński" authorityYear="2018" box="[915,1025,1574,1597]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B52780588CC582EFBEAF9C8" box="[915,1025,1574,1597]" italics="true" pageId="3" pageNumber="372">Euconnus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EF9D4B578B5278058F60582CFAFEF9C8" author="Jaloszynski" box="[1087,1301,1572,1597]" pageId="3" pageNumber="372" refId="ref4534" refString="Jaloszynski, P. (2012) Taxonomy of ' Euconnus complex'. Part I. Morphology of Euconnus s. str. and revision of Euconnomorphus Franz and Venezolanoconnus Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3555, 55 - 82." type="journal article" year="2012">Jałoszyński (2012)</bibRefCitation>
, and can be maintained as a subgenus. Although it is similar to several other subgenera of 
<taxonomicName id="4C0C4D258B5278058F425842FB60F994" authorityName="Jałoszyński" authorityYear="2018" box="[1053,1163,1610,1633]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058F425842FB60F994" box="[1053,1163,1610,1633]" italics="true" pageId="3" pageNumber="372">Euconnus</emphasis>
</taxonomicName>
(see remarks below), it differs in a combination of several characters from each of them.
</paragraph>
</subSubSection>
<subSubSection id="C316652D8B5278038B985898FE0EFEB5" lastPageId="5" lastPageNumber="374" pageId="3" pageNumber="372" type="diagnosis">
<paragraph id="8BB336A68B5278058B985898FA85F95C" blockId="3.[151,1436,1572,1705]" box="[199,1390,1680,1705]" pageId="3" pageNumber="372">
<emphasis id="B978EAB48B5278058B985898FEDBF95C" bold="true" box="[199,304,1680,1705]" pageId="3" pageNumber="372">Remark.</emphasis>
<taxonomicName id="4C0C4D258B5278058A685898FE20F95C" authorityName="Franz" authorityYear="1986" box="[311,459,1680,1705]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058A685898FE20F95C" box="[311,459,1680,1705]" italics="true" pageId="3" pageNumber="372">Androconnus</emphasis>
</taxonomicName>
differs from hitherto redefined subgenera of 
<taxonomicName id="4C0C4D258B527805889B589AFBD9F95C" authorityName="Jałoszyński" authorityYear="2018" box="[964,1074,1682,1705]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B527805889B589AFBD9F95C" box="[964,1074,1682,1705]" italics="true" pageId="3" pageNumber="372">Euconnus</emphasis>
</taxonomicName>
in the following characters:
</paragraph>
<paragraph id="8BB336A68B5278058BC858D1FDD2F818" blockId="3.[151,1436,1752,2030]" pageId="3" pageNumber="372">
• 
<emphasis id="B978EAB48B5278058B9858D0FD9CF904" bold="true" box="[199,631,1752,1777]" pageId="3" pageNumber="372">
from 
<taxonomicName id="4C0C4D258B5278058A5258D0FD9CF904" authority="Franz, 1971" authorityName="Franz" authorityYear="1971" box="[269,631,1752,1777]" class="Insecta" genus="Austroconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058A5258D0FE32F904" bold="true" box="[269,473,1752,1777]" italics="true" pageId="3" pageNumber="372">Austroconophron</emphasis>
<bibRefCitation id="EF9D4B578B5278058ABA58D0FD9CF904" author="Franz, H." box="[485,631,1752,1777]" journalOrPublisher="Koleopterologische Rundschau" pageId="3" pageNumber="372" pagination="43 – 157" part="49" refId="ref4307" refString="Franz, H. (1971) Untersuchungen uber die Scydmaenidenfauna Neukaledoniens (Col., Scyd.). Koleopterologische Rundschau, 49, 43 - 157." title="Untersuchungen über die Scydmaenidenfauna Neukaledoniens (Col., Scyd.)" type="journal article" year="1971">
Franz, 
<quantity id="4CF49B438B527805896058D0FD9CF904" box="[575,631,1752,1777]" metricMagnitude="1" metricUnit="m" metricValue="5.00634" pageId="3" pageNumber="372" unit="in" value="1971.0">1971</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous antennae (gradually thickened in 
<taxonomicName id="4C0C4D258B5278058F9658D0FA60F904" authorityName="Franz" authorityYear="1971" box="[1225,1419,1752,1777]" class="Insecta" genus="Austroconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058F9658D0FA60F904" box="[1225,1419,1752,1777]" italics="true" pageId="3" pageNumber="372">Austroconophron</emphasis>
</taxonomicName>
); pronotum with antebasal pits, groove and sublateral carinae (lacking any of those structures in 
<taxonomicName id="4C0C4D258B5278058FE358F5FA95F8E3" authorityName="Franz" authorityYear="1971" box="[1212,1406,1789,1814]" class="Insecta" genus="Austroconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058FE358F5FA95F8E3" box="[1212,1406,1789,1814]" italics="true" pageId="3" pageNumber="372">Austroconophron</emphasis>
</taxonomicName>
); • 
<emphasis id="B978EAB48B5278058B985928FDA4F8CC" bold="true" box="[199,591,1824,1849]" pageId="3" pageNumber="372">
from 
<taxonomicName id="4C0C4D258B5278058A545928FDA4F8CC" authority="Reitter, 1909" authorityName="Reitter" authorityYear="1909" box="[267,591,1824,1849]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058A545928FE40F8CC" bold="true" box="[267,427,1824,1849]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
<bibRefCitation id="EF9D4B578B5278058AEA5928FDA4F8CC" author="Reitter" box="[437,591,1824,1849]" pageId="3" pageNumber="372" refId="ref5582" refString="Reitter, E. (1909) Fauna Germanica. Die Kafer des Deutschen Reiches. Vol. 2. K. G. Lutz, Stuttgart, 392 pp." type="book" year="1909">
Reitter, 
<quantity id="4CF49B438B52780589485928FDA4F8CC" box="[535,591,1824,1849]" metricMagnitude="1" metricUnit="m" metricValue="4.84886" pageId="3" pageNumber="372" unit="in" value="1909.0">1909</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous antennal club (indistinctly tetramerous in 
<taxonomicName id="4C0C4D258B5278058FAC5928FA61F8CC" authorityName="Reitter" authorityYear="1909" box="[1267,1418,1824,1849]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058FAC5928FA61F8CC" box="[1267,1418,1824,1849]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
</taxonomicName>
); short mandibles lacking mesal teeth (strikingly long and slender mandibles, each with one mesal tooth in 
<taxonomicName id="4C0C4D258B5278058BC85960FEC5F874" authorityName="Reitter" authorityYear="1909" box="[151,302,1896,1921]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058BC85960FEC5F874" box="[151,302,1896,1921]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
</taxonomicName>
); hypostomal ridges not connected posteriorly (connected or nearly connected in 
<taxonomicName id="4C0C4D258B5278058FAC5960FA61F874" authorityName="Reitter" authorityYear="1909" box="[1267,1418,1896,1921]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058FAC5960FA61F874" box="[1267,1418,1896,1921]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
</taxonomicName>
); pronotum with an even number of antebasal pits (with an uneven number of pits in 
<taxonomicName id="4C0C4D258B5278058F365985FAEBF853" authorityName="Reitter" authorityYear="1909" box="[1129,1280,1933,1958]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058F365985FAEBF853" box="[1129,1280,1933,1958]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
</taxonomicName>
, which has a median pit); anterolateral corners of prosternum weakly projecting anteriorly (strongly projecting, forming large subtriangular lobes in 
<taxonomicName id="4C0C4D258B5278058ACF59DDFDCCF81B" authorityName="Reitter" authorityYear="1909" box="[400,551,2005,2030]" class="Insecta" family="Staphylinidae" genus="Euconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5278058ACF59DDFDCCF81B" box="[400,551,2005,2030]" italics="true" pageId="3" pageNumber="372">Euconophron</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BB336A68B5578028BC85E90FF11FEE9" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985E9FFE2CFF45" bold="true" box="[199,455,151,176]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A565E90FE94FF5A" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[265,383,152,175]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978EAB48B5578028A565E90FE94FF5A" bold="true" box="[265,383,152,175]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B5578028AD85E9FFE2CFF45" box="[391,455,151,176]" pageId="4" pageNumber="373" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
in pentamerous antennae (gradually thickened or indistinctly tetramerous in 
<taxonomicName id="4C0C4D258B5578028E725E91FA70FF45" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[1325,1435,153,176]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978EAB48B5578028E725E91FA70FF45" box="[1325,1435,153,176]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B5578028BC85EB5FF3FFF21" box="[151,212,188,212]" pageId="4" pageNumber="373" sensu="stricto">s. str.</taxonomicNameLabel>
); hypostomal ridges not connected posteriorly (connected or nearly connected in 
<taxonomicName id="4C0C4D258B5578028F3D5EB5FB3BFF21" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[1122,1232,189,212]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978EAB48B5578028F3D5EB5FB3BFF21" box="[1122,1232,189,212]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B5578028F885EB5FAFFFF21" box="[1239,1300,188,212]" pageId="4" pageNumber="373" sensu="stricto">s. str.</taxonomicNameLabel>
); mandibles lacking mesal teeth (with one mesal tooth in 
<taxonomicName id="4C0C4D258B55780289F65EE9FCFCFF0D" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[681,791,225,248]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978EAB48B55780289F65EE9FCFCFF0D" box="[681,791,225,248]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B557802887D5EE8FC88FF0D" box="[802,867,224,248]" pageId="4" pageNumber="373" sensu="stricto">s. str.</taxonomicNameLabel>
); antennae in males with modified antennomeres VIII–XI;
</paragraph>
<paragraph id="8BB336A68B5578028BC85F20FB5BFCCD" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985F2FFD88FEB5" bold="true" box="[199,611,295,320]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A4F5F2FFD88FEB5" authority="Franz, 1986 b" authorityName="Franz" authorityYear="1986" box="[272,611,295,320]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A4F5F2FFE5BFEB5" bold="true" box="[272,432,295,320]" italics="true" pageId="4" pageNumber="373">Filonapochus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AE05F2FFD88FEB5" author="Franz, H." box="[447,611,295,320]" journalOrPublisher="Mathematisch-Naturwissenschaftliche Klasse" pageId="4" pageNumber="373" pagination="1 – 393" part="125" refId="ref4388" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." title="Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Österreichische Akademie der Wissenschaften" type="journal article" year="1986">Franz, 1986b</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous antennal club (antennae very slender and gradually thickened in 
<taxonomicName id="4C0C4D258B5578028A785F44FE2AFE90" authorityName="Franz" authorityYear="1986" box="[295,449,332,357]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A785F44FE2AFE90" box="[295,449,332,357]" italics="true" pageId="4" pageNumber="373">Filonapochus</emphasis>
</taxonomicName>
); elongate head (head short, round in 
<taxonomicName id="4C0C4D258B55780288345F44FBEEFE90" authorityName="Franz" authorityYear="1986" box="[875,1029,332,357]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288345F44FBEEFE90" box="[875,1029,332,357]" italics="true" pageId="4" pageNumber="373">Filonapochus</emphasis>
</taxonomicName>
); head lacking ventrolateral cavities near occipital constriction; pronotum broadest only slightly behind middle (at base in 
<taxonomicName id="4C0C4D258B5578028F345F67FAEEFE7D" authorityName="Franz" authorityYear="1986" box="[1131,1285,367,392]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F345F67FAEEFE7D" box="[1131,1285,367,392]" italics="true" pageId="4" pageNumber="373">Filonapochus</emphasis>
</taxonomicName>
) and bearing antebasal pits, transverse groove and sublateral carinae (lacking any of those structures in 
<taxonomicName id="4C0C4D258B5578028FD75F9CFAC9FE58" authorityName="Franz" authorityYear="1986" box="[1160,1314,404,429]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028FD75F9CFAC9FE58" box="[1160,1314,404,429]" italics="true" pageId="4" pageNumber="373">Filonapochus</emphasis>
</taxonomicName>
); • 
<emphasis id="B978EAB48B5578028B985FBFFD71FE25" bold="true" box="[199,666,439,464]" pageId="4" pageNumber="373">
from 
<emphasis id="B978EAB48B5578028A525FBFFE47FE25" bold="true" box="[269,428,439,464]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AE85FBFFD71FE25" author="Jaloszynski" box="[439,666,439,464]" pageId="4" pageNumber="373" refId="ref4871" refString="Jaloszynski, P. (2016 b) Taxonomy of ' Euconnus complex'. Part VII. Glabriconnus, a new subgenus of Euconnus from Madagascar (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4084 (4), 572 - 586. https: // doi. org / 10.11646 / zootaxa. 4084.4.7" type="journal article" year="2016" yearSuffix="b">Jałoszyński, 2016b</bibRefCitation>
</emphasis>
in an elongate head (in 
<emphasis id="B978EAB48B557802889C5FBFFBB6FE25" box="[963,1117,439,464]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
head short, subpentagonal, ' 
<taxonomicName id="4C0C4D258B5578028BC35FD4FEF9FE00" authorityName="Broun" authorityYear="1893" box="[156,274,476,501]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC35FD4FEF9FE00" box="[156,274,476,501]" italics="true" pageId="4" pageNumber="373">Sciacharis</emphasis>
</taxonomicName>
-like'); pentamerous antennal club (antennae gradually thickened in 
<emphasis id="B978EAB48B5578028F455FD4FB5FFE00" box="[1050,1204,476,501]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
); mandibles lacking mesal teeth (one mesal tooth in 
<emphasis id="B978EAB48B55780289715FF7FD23FDED" box="[558,712,511,536]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
); pronotum broadest behind middle (in front of middle in 
<emphasis id="B978EAB48B5578028BC85C2CFEDAFDC8" box="[151,305,548,573]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
); sides of pronotum lacking longitudinal hypomeral grooves (present in 
<emphasis id="B978EAB48B5578028FD75C2CFAC9FDC8" box="[1160,1314,548,573]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
); pronotal base with two pairs of pits, transverse groove and sublateral carinae (only with one pair of pits in 
<emphasis id="B978EAB48B5578028F845C4FFA9EFD95" box="[1243,1397,583,608]" italics="true" pageId="4" pageNumber="373">Glabriconnus</emphasis>
); • 
<emphasis id="B978EAB48B5578028B985C64FDD5FD70" bold="true" box="[199,574,620,645]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A575C64FDD5FD70" authority="Franz, 1963" authorityName="Franz" authorityYear="1963" box="[264,574,620,645]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A575C64FE40FD76" bold="true" box="[264,427,620,643]" italics="true" pageId="4" pageNumber="373">Heteroconnus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AEE5C64FDD5FD70" author="Franz, H." box="[433,574,620,645]" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde" pageId="4" pageNumber="373" pagination="1 – 93" part="120" refId="ref4263" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." title="Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten)" type="journal article" year="1963">
Franz, 
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</emphasis>
in hypostomal ridges not reaching posterior tentorial pits; antennae with club (gradually thickened in 
<taxonomicName id="4C0C4D258B5578028AFF5C99FDD7FD5D" authorityName="Franz" authorityYear="1963" box="[416,572,657,680]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028AFF5C99FDD7FD5D" box="[416,572,657,680]" italics="true" pageId="4" pageNumber="373">Heteroconnus</emphasis>
</taxonomicName>
); pronotum with rounded sides (constricted behind middle in 
<taxonomicName id="4C0C4D258B5578028FB05C99FA60FD5D" authorityName="Franz" authorityYear="1963" box="[1263,1419,657,680]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028FB05C99FA60FD5D" box="[1263,1419,657,680]" italics="true" pageId="4" pageNumber="373">Heteroconnus</emphasis>
</taxonomicName>
); an unmodified pronotum in males (with a median impression, carinae and tufts of setae in 
<taxonomicName id="4C0C4D258B5578028FD45CBDFACCFD39" authorityName="Franz" authorityYear="1963" box="[1163,1319,693,716]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028FD45CBDFACCFD39" box="[1163,1319,693,716]" italics="true" pageId="4" pageNumber="373">Heteroconnus</emphasis>
</taxonomicName>
); • 
<emphasis id="B978EAB48B5578028B985CDFFDD1FD05" bold="true" box="[199,570,727,752]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A575CDFFDD1FD05" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[264,570,727,752]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A575CDFFE97FD05" bold="true" box="[264,380,727,752]" italics="true" pageId="4" pageNumber="373">Napochus</emphasis>
<bibRefCitation id="EF9D4B578B5578028ADB5CDFFDD1FD05" author="Thomson" box="[388,570,727,752]" pageId="4" pageNumber="373" refId="ref5651" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. 1. Berlingska Boktryckeriet, Lund, 290 pp. https: // doi. org / 10.5962 / bhl. title. 138677" type="book" year="1859">
Thomson, 
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</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous (tetramerous in 
<taxonomicName id="4C0C4D258B55780288FB5CDFFBFFFD05" authorityName="Thomson" authorityYear="1859" box="[932,1044,727,752]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288FB5CDFFBFFFD05" box="[932,1044,727,752]" italics="true" pageId="4" pageNumber="373">Napochus</emphasis>
</taxonomicName>
) antennal club; pronotum broadest only slightly behind middle and strongly rounded at sides (subconical in 
<taxonomicName id="4C0C4D258B55780288935CF4FBD7FCE0" authorityName="Thomson" authorityYear="1859" box="[972,1084,764,789]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288935CF4FBD7FCE0" box="[972,1084,764,789]" italics="true" pageId="4" pageNumber="373">Napochus</emphasis>
</taxonomicName>
); prosternum lacking any trace of prosternal process (typically an indistinct carina present in 
<taxonomicName id="4C0C4D258B55780288155D17FC51FCCD" authorityName="Thomson" authorityYear="1859" box="[842,954,799,824]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288155D17FC51FCCD" box="[842,954,799,824]" italics="true" pageId="4" pageNumber="373">Napochus</emphasis>
</taxonomicName>
, sometimes reduced);
</paragraph>
<paragraph id="8BB336A68B5578028BC85D4DFB73FC19" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985D4CFDB4FCA8" bold="true" box="[199,607,836,861]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A4C5D4CFDB4FCA8" authority="Franz, 1986 b" authorityName="Franz" authorityYear="1986" box="[275,607,836,861]" class="Insecta" family="Staphylinidae" genus="Nodoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A4C5D4CFE4EFCA8" bold="true" box="[275,421,836,861]" italics="true" pageId="4" pageNumber="373">Nodoconnus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AE85D4CFDB4FCA8" author="Franz, H." box="[439,607,836,861]" journalOrPublisher="Mathematisch-Naturwissenschaftliche Klasse" pageId="4" pageNumber="373" pagination="1 – 393" part="125" refId="ref4388" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." title="Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Österreichische Akademie der Wissenschaften" type="journal article" year="1986">Franz, 1986b</bibRefCitation>
</taxonomicName>
</emphasis>
in elongate head (subpentagonal, ' 
<taxonomicName id="4C0C4D258B5578028F485D4CFB66FCA8" authorityName="Broun" authorityYear="1893" box="[1047,1165,836,861]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F485D4CFB66FCA8" box="[1047,1165,836,861]" italics="true" pageId="4" pageNumber="373">Sciacharis</emphasis>
</taxonomicName>
-like' in 
<taxonomicName id="4C0C4D258B5578028FA05D4CFA60FCA8" authorityName="Franz" authorityYear="1986" box="[1279,1419,836,861]" class="Insecta" family="Staphylinidae" genus="Nodoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028FA05D4CFA60FCA8" box="[1279,1419,836,861]" italics="true" pageId="4" pageNumber="373">Nodoconnus</emphasis>
</taxonomicName>
); hypostomal ridges extending posteromesad and not connected at middle (ridges connected and forming one transverse carina parallel to the posterior mental margin in 
<taxonomicName id="4C0C4D258B557802881D5D84FC25FC50" authorityName="Franz" authorityYear="1986" box="[834,974,908,933]" class="Insecta" family="Staphylinidae" genus="Nodoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B557802881D5D84FC25FC50" box="[834,974,908,933]" italics="true" pageId="4" pageNumber="373">Nodoconnus</emphasis>
</taxonomicName>
); pronotum broadest behind middle and rounded at sides (broadest in front of middle and with sinuate sides in 
<taxonomicName id="4C0C4D258B55780288A35DA7FB63FC3D" authorityName="Franz" authorityYear="1986" box="[1020,1160,943,968]" class="Insecta" family="Staphylinidae" genus="Nodoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288A35DA7FB63FC3D" box="[1020,1160,943,968]" italics="true" pageId="4" pageNumber="373">Nodoconnus</emphasis>
</taxonomicName>
); pronotal base with a transverse groove and sublateral carinae (only with one pair of diffuse pits in 
<taxonomicName id="4C0C4D258B55780288A45DDCFB6CFC18" authorityName="Franz" authorityYear="1986" box="[1019,1159,980,1005]" class="Insecta" family="Staphylinidae" genus="Nodoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288A45DDCFB6CFC18" box="[1019,1159,980,1005]" italics="true" pageId="4" pageNumber="373">Nodoconnus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BB336A68B5578028BC85DF0FD9EFBAD" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985DFFFD93FBE5" bold="true" box="[199,632,1015,1040]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A495DFFFD93FBE5" authority="Franz, 1963" authorityName="Franz" authorityYear="1963" box="[278,632,1015,1040]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A495DFFFE2CFBE5" bold="true" box="[278,455,1015,1040]" italics="true" pageId="4" pageNumber="373">Paratetramelus</emphasis>
<bibRefCitation id="EF9D4B578B5578028A825DFFFD93FBE5" author="Franz, H." box="[477,632,1015,1040]" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde" pageId="4" pageNumber="373" pagination="1 – 93" part="120" refId="ref4263" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." title="Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten)" type="journal article" year="1963">
Franz, 
<quantity id="4CF49B438B557802891F5DFFFD93FBE5" box="[576,632,1015,1040]" metricMagnitude="1" metricUnit="m" metricValue="4.98602" pageId="4" pageNumber="373" unit="in" value="1963.0">1963</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in elongate head (
<taxonomicName id="4C0C4D258B55780288205DFFFC1EFBE5" authorityName="Broun" authorityYear="1893" box="[895,1013,1015,1040]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288205DFFFC1EFBE5" box="[895,1013,1015,1040]" italics="true" pageId="4" pageNumber="373">Sciacharis</emphasis>
</taxonomicName>
-like, subpentagonal and short in 
<taxonomicName id="4C0C4D258B5578028BC85A14FEAEFBC0" authorityName="Franz" authorityYear="1963" box="[151,325,1052,1077]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC85A14FEAEFBC0" box="[151,325,1052,1077]" italics="true" pageId="4" pageNumber="373">Paratetramelus</emphasis>
</taxonomicName>
); antennae with club (gradually thickened in 
<taxonomicName id="4C0C4D258B55780288075A14FBEDFBC0" authorityName="Franz" authorityYear="1963" box="[856,1030,1052,1077]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288075A14FBEDFBC0" box="[856,1030,1052,1077]" italics="true" pageId="4" pageNumber="373">Paratetramelus</emphasis>
</taxonomicName>
); hypostomal ridges not connected posteriorly (connected in 
<taxonomicName id="4C0C4D258B5578028AE95A37FD8FFBAD" authorityName="Franz" authorityYear="1963" box="[438,612,1087,1112]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028AE95A37FD8FFBAD" box="[438,612,1087,1112]" italics="true" pageId="4" pageNumber="373">Paratetramelus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BB336A68B5578028BC85A6DFB48FB89" blockId="4.[151,1436,151,1581]" box="[151,1187,1124,1149]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985A6CFDCAFB88" bold="true" box="[199,545,1124,1149]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A575A6CFDCAFB88" authority="Casey, 1897" authorityName="Casey" authorityYear="1897" box="[264,545,1124,1149]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A575A6CFE65FB88" bold="true" box="[264,398,1124,1149]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
<bibRefCitation id="EF9D4B578B5578028ACA5A6CFDCAFB88" author="Casey, T. L." box="[405,545,1124,1149]" journalOrPublisher="Annals of the New York Academy of Science" pageId="4" pageNumber="373" pagination="285 – 684" part="9" publicationUrl="https://doi.org/10.1111/j.1749-6632.1896.tb55435.x" refId="ref4215" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Science, 9, 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" title="Coleopterological notices, VII" type="journal article" year="1897">
Casey, 
<quantity id="4CF49B438B5578028AB65A6CFDCAFB88" box="[489,545,1124,1149]" metricMagnitude="1" metricUnit="m" metricValue="4.81838" pageId="4" pageNumber="373" unit="in" value="1897.0">1897</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous antennal club (trimerous in 
<taxonomicName id="4C0C4D258B5578028F4F5A6CFB7AFB88" authorityName="Casey" authorityYear="1897" box="[1040,1169,1124,1149]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F4F5A6CFB7AFB88" box="[1040,1169,1124,1149]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BB336A68B5578028BC85A80FEC8FAF9" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985A8FFDF1FB55" bold="true" box="[199,538,1159,1184]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A575A8FFDF1FB55" authority="Casey, 1897" authorityName="Casey" authorityYear="1897" box="[264,538,1159,1184]" class="Insecta" family="Staphylinidae" genus="Pycnophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A575A8FFE63FB55" bold="true" box="[264,392,1159,1184]" italics="true" pageId="4" pageNumber="373">Pycnophus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AD15A8FFDF1FB55" author="Casey, T. L." box="[398,538,1159,1184]" journalOrPublisher="Annals of the New York Academy of Science" pageId="4" pageNumber="373" pagination="285 – 684" part="9" publicationUrl="https://doi.org/10.1111/j.1749-6632.1896.tb55435.x" refId="ref4215" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Science, 9, 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" title="Coleopterological notices, VII" type="journal article" year="1897">
Casey, 
<quantity id="4CF49B438B5578028ABD5A8FFDF1FB55" box="[482,538,1159,1184]" metricMagnitude="1" metricUnit="m" metricValue="4.81838" pageId="4" pageNumber="373" unit="in" value="1897.0">1897</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in pentamerous antennal club (tetramerous in 
<taxonomicName id="4C0C4D258B5578028F7F5A8FFB70FB55" authorityName="Casey" authorityYear="1897" box="[1056,1179,1159,1184]" class="Insecta" family="Staphylinidae" genus="Pycnophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F7F5A8FFB70FB55" box="[1056,1179,1159,1184]" italics="true" pageId="4" pageNumber="373">Pycnophus</emphasis>
</taxonomicName>
); elongate head (large, subcircular, often broader than long in 
<taxonomicName id="4C0C4D258B55780289135AA4FD2CFB30" authorityName="Casey" authorityYear="1897" box="[588,711,1196,1221]" class="Insecta" family="Staphylinidae" genus="Pycnophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780289135AA4FD2CFB30" box="[588,711,1196,1221]" italics="true" pageId="4" pageNumber="373">Pycnophus</emphasis>
</taxonomicName>
); pronotum broadest slightly behind middle (typically at base in 
<taxonomicName id="4C0C4D258B5578028BC85AC7FEF9FB1D" authorityName="Casey" authorityYear="1897" box="[151,274,1231,1256]" class="Insecta" family="Staphylinidae" genus="Pycnophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC85AC7FEF9FB1D" box="[151,274,1231,1256]" italics="true" pageId="4" pageNumber="373">Pycnophus</emphasis>
</taxonomicName>
) and bearing two pairs of pits and a transverse groove (one pair of pits and typically lacking groove in 
<taxonomicName id="4C0C4D258B5578028BC85AFCFEF9FAF8" authorityName="Casey" authorityYear="1897" box="[151,274,1268,1293]" class="Insecta" family="Staphylinidae" genus="Pycnophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC85AFCFEF9FAF8" box="[151,274,1268,1293]" italics="true" pageId="4" pageNumber="373">Pycnophus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BB336A68B5578028BC85B10FC2EFA68" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985B1FFD86FAC5" bold="true" box="[199,621,1303,1328]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A535B1FFD86FAC5" authority="Franz, 1986 a" authorityName="Franz" authorityYear="1986" box="[268,621,1303,1328]" class="Insecta" family="Staphylinidae" genus="Rhomboconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A535B1FFE29FAC5" bold="true" box="[268,450,1303,1328]" italics="true" pageId="4" pageNumber="373">Rhomboconnus</emphasis>
<bibRefCitation id="EF9D4B578B5578028A925B1FFD86FAC5" author="Franz, H." box="[461,621,1303,1328]" journalOrPublisher="Koleopterologische Rundschau" pageId="4" pageNumber="373" pagination="3 – 42" part="58" refId="ref4336" refString="Franz, H. (1986 a) Monographie der Scydmaeniden Venezuelas I. Teil (Genera Scydmaenus Latreille, Neuraphanax Reitter, Protoconnus Franz, Homoconnus Schaufuss und Euconnus Rhomboconnus subg. nov. sowie Androconnus subg. nov.). Koleopterologische Rundschau, 58, 3 - 42." title="Monographie der Scydmaeniden Venezuelas I. Teil (Genera Scydmaenus Latreille, Neuraphanax Reitter, Protoconnus Franz, Homoconnus Schaufuss und Euconnus Rhomboconnus subg. nov. sowie Androconnus subg. nov.)" type="journal article" year="1986">Franz, 1986a</bibRefCitation>
</taxonomicName>
</emphasis>
in ventral surface of head lacking a well-defined longitudinal groove connecting posterior tentorial pits and apices of hypostomal ridges; hypostomal ridges not connected posteriorly; tempora gradually, and not stepwise narrowing caudad and lacking a well-defined row of short and very dense bristles; pronotum with distinct sublateral carinae and transverse groove;
</paragraph>
<paragraph id="8BB336A68B5578028BC85BA0FEC8F9D9" blockId="4.[151,1436,151,1581]" pageId="4" pageNumber="373">
• 
<emphasis id="B978EAB48B5578028B985BAFFD9DFA35" bold="true" box="[199,630,1447,1472]" pageId="4" pageNumber="373">
from 
<taxonomicName id="4C0C4D258B5578028A535BAFFD9DFA35" authority="Motschulsky, 1870" authorityName="Motschulsky" authorityYear="1870" box="[268,630,1447,1472]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028A535BAFFE62FA35" bold="true" box="[268,393,1447,1472]" italics="true" pageId="4" pageNumber="373">Tetramelus</emphasis>
<bibRefCitation id="EF9D4B578B5578028AC95BAFFD9DFA35" author="Motschulsky" box="[406,630,1447,1472]" pageId="4" pageNumber="373" refId="ref5470" refString="Motschulsky, V. (1870) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. 7 - ieme Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42, 252 - 272." type="journal article" year="1870">
Motschulsky, 
<quantity id="4CF49B438B55780289615BAFFD9DFA35" box="[574,630,1447,1472]" metricMagnitude="1" metricUnit="m" metricValue="4.7498" pageId="4" pageNumber="373" unit="in" value="1870.0">1870</quantity>
</bibRefCitation>
</taxonomicName>
</emphasis>
in a short clypeus (strikingly long in 
<taxonomicName id="4C0C4D258B5578028F6A5BAFFB5AFA35" authorityName="Motschulsky" authorityYear="1870" box="[1077,1201,1447,1472]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F6A5BAFFB5AFA35" box="[1077,1201,1447,1472]" italics="true" pageId="4" pageNumber="373">Tetramelus</emphasis>
</taxonomicName>
); mandibles lacking mesal teeth (one mesal tooth typically present in 
<taxonomicName id="4C0C4D258B55780289BB5BC4FC8BFA10" authorityName="Motschulsky" authorityYear="1870" box="[740,864,1484,1509]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780289BB5BC4FC8BFA10" box="[740,864,1484,1509]" italics="true" pageId="4" pageNumber="373">Tetramelus</emphasis>
</taxonomicName>
); pronotum with sides rounded in posterior half (typically constricted near posterior third in 
<taxonomicName id="4C0C4D258B55780289E65BE7FCDEF9FD" authorityName="Motschulsky" authorityYear="1870" box="[697,821,1519,1544]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780289E65BE7FCDEF9FD" box="[697,821,1519,1544]" italics="true" pageId="4" pageNumber="373">Tetramelus</emphasis>
</taxonomicName>
); pronotal base lacking median carina (present in 
<taxonomicName id="4C0C4D258B5578028BC8581CFEF8F9D8" authorityName="Motschulsky" authorityYear="1870" box="[151,275,1556,1581]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC8581CFEF8F9D8" box="[151,275,1556,1581]" italics="true" pageId="4" pageNumber="373">Tetramelus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB336A68B5578028B985855FD41F885" blockId="4.[151,1437,1628,2013]" pageId="4" pageNumber="373">
From each of these subgenera, except one, 
<taxonomicName id="4C0C4D258B55780289FA5854FCD2F980" authorityName="Franz" authorityYear="1986" box="[677,825,1628,1653]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780289FA5854FCD2F980" box="[677,825,1628,1653]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
</taxonomicName>
differs not only in the modified antennae of males, but also in some other characters. The only exception seems to be 
<taxonomicName id="4C0C4D258B55780288CA5877FBFDF96D" authorityName="Casey" authorityYear="1897" box="[917,1046,1663,1688]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288CA5877FBFDF96D" box="[917,1046,1663,1688]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
</taxonomicName>
, and in future 
<taxonomicName id="4C0C4D258B5578028F8D5877FA8DF96D" authorityName="Franz" authorityYear="1986" box="[1234,1382,1663,1688]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F8D5877FA8DF96D" box="[1234,1382,1663,1688]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0C4D258B5578028BC858ACFEF3F948" authorityName="Casey" authorityYear="1897" box="[151,280,1700,1725]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028BC858ACFEF3F948" box="[151,280,1700,1725]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
</taxonomicName>
may be placed as synonyms. However, the worldwide subgenus 
<taxonomicName id="4C0C4D258B5578028F5F58ACFB6AF948" authorityName="Casey" authorityYear="1897" box="[1024,1153,1700,1725]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F5F58ACFB6AF948" box="[1024,1153,1700,1725]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
</taxonomicName>
requires further study to understand its limits and character variability, whereas 
<taxonomicName id="4C0C4D258B557802885158CFFC49F915" authorityName="Franz" authorityYear="1986" box="[782,930,1735,1760]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B557802885158CFFC49F915" box="[782,930,1735,1760]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
</taxonomicName>
seems to represent a group of species welldefined not only morphologically, but also geographically (restricted to the northern part of Neotropical region) and may constitute a monophyletic unit. For this reason, I prefer to maintain 
<taxonomicName id="4C0C4D258B55780288A45907FB64F8DD" authorityName="Franz" authorityYear="1986" box="[1019,1167,1807,1832]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B55780288A45907FB64F8DD" box="[1019,1167,1807,1832]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
</taxonomicName>
as a separate subgenus, pending further study. 
<taxonomicName id="4C0C4D258B5578028AC1593CFDD9F8B8" authorityName="Franz" authorityYear="1986" box="[414,562,1844,1869]" class="Insecta" family="Staphylinidae" genus="Androconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028AC1593CFDD9F8B8" box="[414,562,1844,1869]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0C4D258B5578028930593CFD1BF8B8" authorityName="Casey" authorityYear="1897" box="[623,752,1844,1869]" class="Insecta" family="Staphylinidae" genus="Psomophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028930593CFD1BF8B8" box="[623,752,1844,1869]" italics="true" pageId="4" pageNumber="373">Psomophus</emphasis>
</taxonomicName>
are most similar to 
<taxonomicName id="4C0C4D258B5578028880593DFBA6F8B9" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[991,1101,1845,1868]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B978EAB48B5578028880593DFBA6F8B9" box="[991,1101,1845,1868]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57CF8B5578028F07593DFB7DF8B9" box="[1112,1174,1845,1868]" pageId="4" pageNumber="373" sensu="stricto">s. str.</taxonomicNameLabel>
and 
<taxonomicName id="4C0C4D258B5578028F8D593CFA7FF8B8" authorityName="Franz" authorityYear="1971" box="[1234,1428,1844,1869]" class="Insecta" genus="Austroconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B5578028F8D593CFA7FF8B8" box="[1234,1428,1844,1869]" italics="true" pageId="4" pageNumber="373">Austroconophron</emphasis>
</taxonomicName>
; these taxa require a more comprehensive study.
</paragraph>
<paragraph id="8BB336A68B5578038B985974FE0EFEB5" blockId="4.[151,1437,1628,2013]" lastBlockId="5.[151,1437,151,320]" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373">
<emphasis id="B978EAB48B5578028B985974FCFCF860" bold="true" box="[199,791,1916,1941]" pageId="4" pageNumber="373">
Emended diagnosis of 
<taxonomicName id="4C0C4D258B5578028A805974FCFAF860" authority="" box="[479,785,1916,1941]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="subGenus" subGenus="Androconnus">
<emphasis id="B978EAB48B5578028A805974FDBEF866" bold="true" box="[479,597,1916,1939]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
(
<emphasis id="B978EAB48B55780289355974FCECF860" bold="true" box="[618,775,1916,1941]" italics="true" pageId="4" pageNumber="373">Androconnus</emphasis>
)
</taxonomicName>
.
</emphasis>
<taxonomicName id="4C0C4D258B557802887D5975FC7BF861" authorityName="Jałoszyński" authorityYear="2018" box="[802,912,1917,1940]" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB48B557802887D5975FC7BF861" box="[802,912,1917,1940]" italics="true" pageId="4" pageNumber="373">Euconnus</emphasis>
</taxonomicName>
with strikingly long antennae; antennal club indistinctly pentamerous; antennomeres IX–XI modified in males, asymmetrical, with impressions and/or mesal tooth-like projections; head elongate, with vertex indistinctly bulging posterodorsad; tempora longer than eyes and covered with bristles; hypostomal ridges long but not reaching posterior tentorial pits and not connected posteriorly; mandibles short and lacking mesal teeth; pronotum broadest slightly behind middle, with rounded sides; pronotal base with paired pits, sublateral carinae and transverse groove; interprocoxal area lacking any trace of carina or process; sides of pronotum with bristles; basal elytral foveae small but distinct; metaventral intercoxal process distinctly emarginate.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>