252 lines
37 KiB
XML
252 lines
37 KiB
XML
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<mods:title id="BC45DA9F4BF790BC6BDAA0558DB6B43F">Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae</mods:title>
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<mods:name id="A6532A4760A8B730A0E345F68A21894E" type="personal">
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<mods:namePart id="43D8F65F276DB68177266943F7AAE1FC">Andrew Hamilton, K. G.</mods:namePart>
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<mods:title id="48A4C174496F629FD2343C026DF7B7CB">Zootaxa</mods:title>
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<mods:date id="F15E866CC7E06EA426893C0BC0D969A6">2016</mods:date>
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<mods:number id="9AD097FDB51E30501464A5DA63063AD3">4169</mods:number>
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<treatment id="03A5742DFFC3AE1DFF76AC15CF63FD79" ID-DOI="http://doi.org/10.5281/zenodo.5613355" ID-GBIF-Taxon="124552105" ID-Zenodo-Dep="5613355" LSID="urn:lsid:plazi:treatment:03A5742DFFC3AE1DFF76AC15CF63FD79" httpUri="http://treatment.plazi.org/id/03A5742DFFC3AE1DFF76AC15CF63FD79" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
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<subSubSection id="C31696B0FFC3AE1CFF76AC15CF2FFA8B" box="[151,607,1344,1370]" pageId="22" pageNumber="223" type="nomenclature">
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<paragraph id="8BB3C53BFFC3AE1CFF76AC15CF2FFA8B" blockId="22.[151,607,1344,1370]" box="[151,607,1344,1370]" pageId="22" pageNumber="223">
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<heading id="D0FB7257FFC3AE1CFF76AC15CF2FFA8B" bold="true" box="[151,607,1344,1370]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
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<emphasis id="B9781929FFC3AE1CFF76AC15CF2FFA8B" bold="true" box="[151,607,1344,1370]" pageId="22" pageNumber="223">
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<taxonomicName id="4C0CBEB8FFC3AE1CFF76AC15CCADFA8B" ID-CoL="63WHS" authority="Lallemand" authorityName="Lallemand" box="[151,477,1344,1370]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B9781929FFC3AE1CFF76AC15CC3DFA8B" bold="true" box="[151,333,1344,1370]" italics="true" pageId="22" pageNumber="223">Tomaspisinella</emphasis>
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Lallemand
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</taxonomicName>
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, redefined
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31696B0FFC3AE1CFF76ACD2C841FA6A" pageId="22" pageNumber="223" type="reference_group">
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<paragraph id="8BB3C53BFFC3AE1CFF76ACD2C9DEFA4C" blockId="22.[151,1329,1415,1467]" box="[151,1198,1415,1437]" pageId="22" pageNumber="223">
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<treatmentCitationGroup id="AB1CE215FFC3AE1CFF76ACD2C9DEFA4C" box="[151,1198,1415,1437]" pageId="22" pageNumber="223">
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<taxonomicName id="4C0CBEB8FFC3AE1CFF76ACD2CF66FA4C" ID-CoL="63WHS" authority="Lallemand, 1927: 117" authorityName="Lallemand" authorityPageNumber="117" authorityYear="1927" box="[151,534,1415,1437]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B9781929FFC3AE1CFF76ACD2CC40FA4C" box="[151,304,1415,1437]" italics="true" pageId="22" pageNumber="223">Tomaspisinella</emphasis>
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<treatmentCitation id="0AADE32AFFC3AE1CFED4ACD2CF66FA4C" author="Lallemand" box="[309,534,1415,1437]" page="117" pageId="22" pageNumber="223" year="1927">
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<bibRefCitation id="EF9DB8CAFFC3AE1CFED4ACD2CC94FA4C" author="Lallemand" box="[309,484,1415,1437]" pageId="22" pageNumber="223" refString="Lallemand, V. (1927) Descriptions de Cercopides nouveaux provenant de la collection du British Museum. Transactions of the Entomological Society of London, 1927, 99 - 118. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1927. tb 00063. x" type="journal article" year="1927">Lallemand, 1927</bibRefCitation>
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: 117
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</treatmentCitation>
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</taxonomicName>
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. Type-species by original designation:
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<taxonomicName id="4C0CBEB8FFC3AE1CFC40ACDDC9DAFA4C" ID-CoL="7CFC5" authority="Lallemand, 1927" authorityName="Lallemand" authorityYear="1927" box="[929,1194,1415,1437]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="parva">
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<emphasis id="B9781929FFC3AE1CFC40ACDDCE83FA4C" box="[929,1011,1416,1437]" italics="true" pageId="22" pageNumber="223">T. parva</emphasis>
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<bibRefCitation id="EF9DB8CAFFC3AE1CFC1AACD2C9DAFA4C" author="Lallemand" box="[1019,1194,1415,1437]" pageId="22" pageNumber="223" refString="Lallemand, V. (1927) Descriptions de Cercopides nouveaux provenant de la collection du British Museum. Transactions of the Entomological Society of London, 1927, 99 - 118. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1927. tb 00063. x" type="journal article" year="1927">Lallemand, 1927</bibRefCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BB3C53BFFC3AE1CFF76ACF3C841FA6A" blockId="22.[151,1329,1415,1467]" box="[151,1329,1445,1467]" pageId="22" pageNumber="223">
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<treatmentCitationGroup id="AB1CE215FFC3AE1CFF76ACF3C841FA6A" box="[151,1329,1445,1467]" pageId="22" pageNumber="223">
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<taxonomicName id="4C0CBEB8FFC3AE1CFF76ACF3CF72FA6A" ID-CoL="4VFY" authority="Lallemand, 1949: 31" authorityName="Lallemand" authorityPageNumber="31" authorityYear="1949" box="[151,514,1445,1467]" class="Insecta" family="Cercopidae" genus="Hemitomaspis" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B9781929FFC3AE1CFF76ACF3CC58FA6A" box="[151,296,1446,1467]" italics="true" pageId="22" pageNumber="223">Hemitomaspis</emphasis>
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<treatmentCitation id="0AADE32AFFC3AE1CFECFACF0CF72FA6A" author="Lallemand" box="[302,514,1445,1467]" page="31" pageId="22" pageNumber="223" year="1949">
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<bibRefCitation id="EF9DB8CAFFC3AE1CFECFACF0CCAEFA6A" author="Lallemand" box="[302,478,1445,1467]" pageId="22" pageNumber="223" refString="Lallemand, V. (1949) Revision des Cercopinae (Hemiptera Homoptera). Memoires de l'Institut royal des Sciences naturelle de Belgique, 32, 1 - 193." type="journal article" year="1949">Lallemand, 1949</bibRefCitation>
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: 31
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</treatmentCitation>
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</taxonomicName>
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,
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<emphasis id="B9781929FFC3AE1CFDECACF0CF15FA6A" bold="true" box="[525,613,1445,1467]" pageId="22" pageNumber="223">syn.nov.</emphasis>
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Type-species by original designation:
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<taxonomicName id="4C0CBEB8FFC3AE1CFC0AACF3C85DFA6A" ID-CoL="57CDZ" authority="Jacobi, 1908" authorityName="Jacobi" authorityYear="1908" box="[1003,1325,1445,1467]" class="Insecta" family="Cercopidae" genus="Tomaspis" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="caligata">
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<emphasis id="B9781929FFC3AE1CFC0AACF3C9D3FA6A" box="[1003,1187,1445,1467]" italics="true" pageId="22" pageNumber="223">Tomaspis caligata</emphasis>
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<bibRefCitation id="EF9DB8CAFFC3AE1CFB4BACF0C85DFA6A" author="Jacobi" box="[1194,1325,1445,1467]" pageId="22" pageNumber="223" refString="Jacobi, A. (1908) Neue Cercopiden des Andengebietes. Sitzungsberichte der Gesellschaft naturforschender Freunde, 8, 200 - 215. http: // dx. doi. org / 10.5962 / bhl. part. 12854" type="journal article" year="1908">Jacobi, 1908</bibRefCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31696B0FFC3AE1CFF76ACBDCE13F9D0" box="[151,867,1512,1537]" pageId="22" pageNumber="223" type="distribution">
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||
<paragraph id="8BB3C53BFFC3AE1CFF76ACBDCE13F9D0" blockId="22.[151,1437,1512,2005]" box="[151,867,1512,1537]" pageId="22" pageNumber="223">
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<emphasis id="B9781929FFC3AE1CFF76ACBDCC40F9D0" bold="true" box="[151,304,1512,1537]" pageId="22" pageNumber="223">Distribution.</emphasis>
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Neotropical, from
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<collectingCountry id="F31B85ABFFC3AE1CFDE6ACBDCFF0F9D0" box="[519,640,1512,1537]" name="Costa Rica" pageId="22" pageNumber="223">
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<collectingRegion id="49C80BD9FFC3AE1CFDE6ACBDCF37F9D1" box="[519,583,1512,1536]" country="New Zealand" name="West Coast" pageId="22" pageNumber="223">Costa</collectingRegion>
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Rica
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</collectingCountry>
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south to
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<collectingCountry id="F31B85ABFFC3AE1CFD09ACBDCE2FF9D0" box="[744,863,1512,1537]" name="Venezuela" pageId="22" pageNumber="223">Venezuela</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31696B0FFC3AE1CFF26AF59CC36F964" pageId="22" pageNumber="223" type="diagnosis">
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<paragraph id="8BB3C53BFFC3AE1CFF26AF59CC36F964" blockId="22.[151,1437,1512,2005]" pageId="22" pageNumber="223">
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<emphasis id="B9781929FFC3AE1CFF26AF59CC31F9F4" bold="true" box="[199,321,1548,1573]" pageId="22" pageNumber="223">Diagnosis.</emphasis>
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Superficially similar to
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<taxonomicName id="4C0CBEB8FFC3AE1CFDB4AF58CFC3F9F5" box="[597,691,1549,1572]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9781929FFC3AE1CFDB4AF58CFC3F9F5" box="[597,691,1549,1572]" italics="true" pageId="22" pageNumber="223">Menytes</emphasis>
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</taxonomicName>
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(
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<figureCitation id="1337D9BEFFC3AE1CFD22AF59CE63F9F4" box="[707,787,1548,1573]" captionStart="FIGURES 14 – 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 – 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="22" pageNumber="223">Fig. 14</figureCitation>
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A) but without sulcate face; color usually black to brown (except
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<taxonomicName id="4C0CBEB8FFC3AE1CFF15AF64CC4EF999" box="[244,318,1585,1608]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="jocosa" status="sp. nov.">
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<emphasis id="B9781929FFC3AE1CFF15AF64CC4EF999" box="[244,318,1585,1608]" italics="true" pageId="22" pageNumber="223">jocosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24BA452FFC3AE1CFEA9AF65CCD5F998" box="[328,421,1584,1609]" pageId="22" pageNumber="223" rank="species">
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<emphasis id="B9781929FFC3AE1CFEA9AF65CCD5F998" bold="true" box="[328,421,1584,1609]" pageId="22" pageNumber="223">sp. nov.</emphasis>
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</taxonomicNameLabel>
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), body shorter, broader across the head, with convex tegmina (
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<figureCitation id="1337D9BEFFC3AE1CFB6DAF65C991F998" box="[1164,1249,1584,1609]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="22" pageNumber="223">Fig. 35</figureCitation>
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) and distinctive antenna (
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<figureCitation id="1337D9BEFFC3AE1CFF1EAF01CC24F9BC" box="[255,340,1620,1645]" captionStart="FIGURES 17 – 19" captionStartId="34.[151,264,1878,1900]" captionTargetBox="[180,1422,193,1855]" captionTargetId="figure@34.[180,1427,193,1856]" captionTargetPageId="34" captionText="FIGURES 17 – 19. Antennal characters of apomorphic new-world Cercopinae: Olcotomaspis (Phymatostethini, 17), Zuata (Neaenini, 18) and Tomaspisella (Neaenini, 19): 17 A, Olcotomaspis versicolor (Lallemand); 17 B, same, from another angle, showing paired basiconic sensillae on inner side; 17 C, O. laterinotata (Fowler); 17 D, O. (= Hyalotomaspis) clarissa; 18 A, Zuata infecta sp. nov.; 18 B, same, detail (note peglike seta surrounded by large socket and tiny secondary seta behind it); 19 A, Tomaspisella parva Lallemand, male; 19 B, same, female; 19 C, same, detail of pits." httpUri="https://zenodo.org/record/262592/files/figure.png" pageId="22" pageNumber="223">Figs 19</figureCitation>
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A–C) with postpedicel retracted into pedicel, as in Orthoraphini (Aphrophorinae) and basiconic sensillum elongate and setiform, as in
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<taxonomicName id="4C0CBEB8FFC3AE1CFDBFAF2CCFCBF941" box="[606,699,1657,1680]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9781929FFC3AE1CFDBFAF2CCFCBF941" box="[606,699,1657,1680]" italics="true" pageId="22" pageNumber="223">Menytes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0CBEB8FFC3AE1CFD1AAF2DCE82F940" authority="Neaenini" authorityName="Neaenini" box="[763,1010,1656,1681]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B9781929FFC3AE1CFD1AAF2DCE1FF940" box="[763,879,1656,1681]" italics="true" pageId="22" pageNumber="223">Simorhina</emphasis>
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(Neaenini)
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</taxonomicName>
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and
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<taxonomicName id="4C0CBEB8FFC3AE1CFBD0AF2DCC33F964" authority="Amyot & Serville (Ischnorhinini)" authorityName="Amyot & Serville (Ischnorhinini" class="Insecta" family="Cercopidae" genus="Sphenorhina" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="B9781929FFC3AE1CFBD0AF2DC9CFF940" box="[1073,1215,1656,1681]" italics="true" pageId="22" pageNumber="223">Sphenorhina</emphasis>
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Amyot & Serville (Ischnorhinini)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31696B0FFC3AE1DFF26AF95CF63FD79" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="description">
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||
<paragraph id="8BB3C53BFFC3AE1DFF26AF95C963FE59" blockId="22.[151,1437,1512,2005]" lastBlockId="23.[151,1437,151,680]" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
|
||
<emphasis id="B9781929FFC3AE1CFF26AF95CC29F908" bold="true" box="[199,345,1728,1753]" pageId="22" pageNumber="223">Description.</emphasis>
|
||
Head distinctly to slightly narrower than pronotum (
|
||
<bibRefCitation id="EF9DB8CAFFC3AE1CFC47AF95C828F909" author="Carvalho" box="[934,1368,1728,1753]" pageId="22" pageNumber="223" refString="Carvalho, G. S. & Webb, M. D. (2005) Cercopid Spittle Bugs of the New World (Hemiptera, Auchenorrhyncha, Cercopidae). Pensoft, Sofia-Moscow, 271 pp." type="book" year="2005">Carvalho & Webb 2005, figs 280–281</bibRefCitation>
|
||
); eyes varying from globose to transverse; crown short, weakly concave; antennal ledge arched above coronal margin; frons convexly inflated. Pronotum with anterior margin steeply declivous (
|
||
<figureCitation id="1337D9BEFFC3AE1CFBE1AE5DC936F8F0" box="[1024,1094,1800,1825]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1857,1879]" captionTargetBox="[218,1367,473,1822]" captionTargetId="figure@5.[218,1368,473,1835]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Examples of Microsarganini (1 – 3), Neaenini (4 – 5) and genera related to Phymatostetha (6 – 8) in dorsolateral aspect (A), dorsal aspect (B), lateral or ventrolateral aspect of head and thorax (C) and hind wing characters including detail of costal margin (D): 1, Microsargane sp.; 2, Microclimax luteosignatus; 3, Microlaqueus isolatus; 4, Helioptera gilvum; 5, Tomaspisinella lucifer sp. nov.; 6, Olcotomaspis laterinotata (Fowler), comb. nov.; 7, Hyalotomaspis clarissa (Jacobi); 8, Tomaspisina frontalis (Walker)." httpUri="https://zenodo.org/record/262586/files/figure.png" pageId="22" pageNumber="223">Fig. 5</figureCitation>
|
||
C), anterior border weakly to distinctly bowed; lateral margins as long as eyes to much shorter than half length of eyes, surface variable from medially carinate and transversely rugulose (
|
||
<figureCitation id="1337D9BEFFC3AE1CFD6CAE05CFAEF8B8" box="[653,734,1872,1897]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="22" pageNumber="223">Fig. 26</figureCitation>
|
||
) to smooth with scattered pits (
|
||
<figureCitation id="1337D9BEFFC3AE1CFBDEAE05C9E1F8B9" box="[1087,1169,1872,1897]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="22" pageNumber="223">Fig. 27</figureCitation>
|
||
) or densely setose (
|
||
<figureCitation id="1337D9BEFFC3AE1CFA8EAE05CDC5F85C" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="22" pageNumber="223">Fig. 35</figureCitation>
|
||
,
|
||
<emphasis id="B9781929FFC3AE1CFF20AE20CC62F85D" box="[193,274,1909,1932]" italics="true" pageId="22" pageNumber="223">Merinx</emphasis>
|
||
subg. nov.), the nominate subgenus having fine setae (
|
||
<figureCitation id="1337D9BEFFC3AE1CFC98AE21CECCF85C" box="[889,956,1908,1933]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1857,1879]" captionTargetBox="[218,1367,473,1822]" captionTargetId="figure@5.[218,1368,473,1835]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Examples of Microsarganini (1 – 3), Neaenini (4 – 5) and genera related to Phymatostetha (6 – 8) in dorsolateral aspect (A), dorsal aspect (B), lateral or ventrolateral aspect of head and thorax (C) and hind wing characters including detail of costal margin (D): 1, Microsargane sp.; 2, Microclimax luteosignatus; 3, Microlaqueus isolatus; 4, Helioptera gilvum; 5, Tomaspisinella lucifer sp. nov.; 6, Olcotomaspis laterinotata (Fowler), comb. nov.; 7, Hyalotomaspis clarissa (Jacobi); 8, Tomaspisina frontalis (Walker)." httpUri="https://zenodo.org/record/262586/files/figure.png" pageId="22" pageNumber="223">Fig. 5</figureCitation>
|
||
C). Tegmina convex, shiny to hirsute; hind wing with 3 apical cells (Cu is unbranched; appendix large, even width around tip of wing. Fore femora distinctly longer than hind femora. Hind tibiae with 2 spines on outer edge, both long in smallest species, but basal one tiny in largest species; hind tibial pectin with 8–9 spines; hind basitarsal pecten of 6–7 black-tipped spines, that of second tarsomere with 8–9 such spines (hind legs missing from
|
||
<typeStatus id="54B77B99FFC2AE1DFCBFA9E8CEFEFF04" box="[862,910,189,213]" pageId="23" pageNumber="224">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFC53A9E8C97CFF05" box="[946,1036,189,212]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="parva">
|
||
<emphasis id="B9781929FFC2AE1DFC53A9E8C97CFF05" box="[946,1036,189,212]" italics="true" pageId="23" pageNumber="224">T. parva</emphasis>
|
||
</taxonomicName>
|
||
); arolia variable from much shorter than claws (in the type-species) as in Ischnorhinini, to longer than the claws (in
|
||
<emphasis id="B9781929FFC2AE1DFBC1A9B4C901FF29" box="[1056,1137,225,248]" italics="true" pageId="23" pageNumber="224">Merinx</emphasis>
|
||
subg. nov.). Male pygofer usually elongate and deeply notched dorsally on each side of anal tube (
|
||
<figureCitation id="1337D9BEFFC2AE1DFC35A851C925FECC" box="[980,1109,260,285]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="23" pageNumber="224">Figs 28–35</figureCitation>
|
||
A) as in
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFB5AA850C868FECD" box="[1211,1304,261,284]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9781929FFC2AE1DFB5AA850C868FECD" box="[1211,1304,261,284]" italics="true" pageId="23" pageNumber="224">Menytes</emphasis>
|
||
</taxonomicName>
|
||
; subgenital plates absent or broad, fused to pygofer, nearly vertical, upper angles with decurved, pointed tips, sometimes with long, median processes (
|
||
<figureCitation id="1337D9BEFFC2AE1DFE4EA819CF73FEB5" box="[431,515,332,357]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="23" pageNumber="224">Fig. 34</figureCitation>
|
||
D); styles unarmed or absent; phallobase short, sometimes fused to theca to form aedeagus; shaft unarmed or with paired, retrorse processes (
|
||
<figureCitation id="1337D9BEFFC2AE1DFCD5A83ACEFBFE59" box="[820,907,367,392]" captionStart="FIGURES 25 – 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 – 35. Neaenini, Menytes (25) and Tomaspisinella (26 – 35): 25 A – B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 – 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="23" pageNumber="224">Figs 28</figureCitation>
|
||
C, 34–35C).
|
||
</paragraph>
|
||
<paragraph id="8BB3C53BFFC2AE1DFF26A8C1CF63FD79" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9781929FFC2AE1DFF26A8C1CCE3FE7C" bold="true" box="[199,403,404,429]" pageId="23" pageNumber="224">Included species.</emphasis>
|
||
The original description included
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFCFEA8C1C9E0FE7C" authority="Jacobi" authorityName="Jacobi" box="[799,1168,404,429]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="caligata">
|
||
<emphasis id="B9781929FFC2AE1DFCFEA8C1C95CFE7C" box="[799,1068,404,429]" italics="true" pageId="23" pageNumber="224">Tomaspisinella caligata</emphasis>
|
||
(Jacobi)
|
||
</taxonomicName>
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFB79A8C0C808FE7C" authority="Lallemand" authorityName="Lallemand" box="[1176,1400,404,429]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="parva">
|
||
<emphasis id="B9781929FFC2AE1DFB79A8C0C987FE7D" box="[1176,1271,405,428]" italics="true" pageId="23" pageNumber="224">T. parva</emphasis>
|
||
Lallemand
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFA66A8C0CC1BFE01" authority="Jacobi" authorityName="Jacobi" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="minuscula">
|
||
<emphasis id="B9781929FFC2AE1DFA66A8C0CC7AFE01" italics="true" pageId="23" pageNumber="224">T. minuscula</emphasis>
|
||
(Jacobi)
|
||
</taxonomicName>
|
||
and 2 species now assigned to
|
||
<emphasis id="B9781929FFC2AE1DFD27A8ECCE7FFE01" box="[710,783,441,464]" italics="true" pageId="23" pageNumber="224">
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFD27A8ECCE7BFE01" box="[710,779,441,464]" class="Insecta" family="Cercopidae" genus="Zuata" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">Zuata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EF9DB8CAFFC2AE1DFCF4A8E2C941FE01" author="Carvalho" box="[789,1073,439,464]" pageId="23" pageNumber="224" refString="Carvalho, G. S. & Webb, M. D. (2005) Cercopid Spittle Bugs of the New World (Hemiptera, Auchenorrhyncha, Cercopidae). Pensoft, Sofia-Moscow, 271 pp." type="book" year="2005">Carvalho & Webb (2005)</bibRefCitation>
|
||
added
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFB60A8ECC8E8FE01" authority="Fowler" authorityName="Fowler" box="[1153,1432,439,464]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="apicifasciata">
|
||
<emphasis id="B9781929FFC2AE1DFB60A8ECC85DFE01" box="[1153,1325,440,464]" italics="true" pageId="23" pageNumber="224">T. apicifasciata</emphasis>
|
||
(Fowler)
|
||
</taxonomicName>
|
||
, known only from the female type, and these identities (except that of
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFC11A889C913FE24" box="[1008,1123,476,501]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="minuscula">
|
||
<emphasis id="B9781929FFC2AE1DFC11A889C913FE24" box="[1008,1123,476,501]" italics="true" pageId="23" pageNumber="224">minuscula</emphasis>
|
||
</taxonomicName>
|
||
) have been confirmed by examination of the antennae.
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFE08AB54CFD2FDC9" box="[489,674,513,536]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="punctatissima">
|
||
<emphasis id="B9781929FFC2AE1DFE08AB54CFD2FDC9" box="[489,674,513,536]" italics="true" pageId="23" pageNumber="224">T. punctatissima</emphasis>
|
||
</taxonomicName>
|
||
(Stål:
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFD11A8AACE15FDC9" box="[752,869,511,536]" class="Insecta" family="Aphrophoridae" genus="Lepyronia" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">Lepyronia</taxonomicName>
|
||
)
|
||
<emphasis id="B9781929FFC2AE1DFC94A8AACE9EFDC9" bold="true" box="[885,1006,511,536]" pageId="23" pageNumber="224">comb.nov.</emphasis>
|
||
and
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFBC9AB54C865FDC9" authority="Fowler" authorityName="Fowler" box="[1064,1301,511,536]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="ignobilis" status="comb. nov.">
|
||
<emphasis id="B9781929FFC2AE1DFBC9AB54C9D8FDC9" box="[1064,1192,511,536]" italics="true" pageId="23" pageNumber="224">T. ignobilis</emphasis>
|
||
(Fowler)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24BA452FFC2AE1DFAC2A8AAC8ECFDC9" box="[1315,1436,511,536]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9781929FFC2AE1DFAC2A8AAC8ECFDC9" bold="true" box="[1315,1436,511,536]" pageId="23" pageNumber="224">comb.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were formerly placed in
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFE4CAB70CFBDFDEC" authority="Lallemand" authorityName="Lallemand" box="[429,717,548,573]" class="Insecta" family="Cercopidae" genus="Hemitomaspis" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9781929FFC2AE1DFE4CAB70CF3DFDED" box="[429,589,549,572]" italics="true" pageId="23" pageNumber="224">Hemitomaspis</emphasis>
|
||
Lallemand
|
||
</taxonomicName>
|
||
and are here transferred to
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFBE4AB71C9DDFDEC" box="[1029,1197,548,573]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9781929FFC2AE1DFBE4AB71C9DDFDEC" box="[1029,1197,548,573]" italics="true" pageId="23" pageNumber="224">Tomaspisinella</emphasis>
|
||
</taxonomicName>
|
||
. These together with 6 new species increase the genus to 12 species. The typical subgenus appears to be paraphyletic with respect to 3 other apomorphic subgenera, with
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFDC2AB38CFF9FD54" box="[547,649,620,645]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="lucifer" status="sp. nov.">
|
||
<emphasis id="B9781929FFC2AE1DFDC2AB38CFF9FD54" box="[547,649,620,645]" italics="true" pageId="23" pageNumber="224">T. lucifer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24BA452FFC2AE1DFD73AB39CF9DFD54" box="[658,749,620,645]" pageId="23" pageNumber="224" rank="species">
|
||
<emphasis id="B9781929FFC2AE1DFD73AB39CF9DFD54" bold="true" box="[658,749,620,645]" pageId="23" pageNumber="224">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
intermediate in pronotal characters and
|
||
<taxonomicName id="4C0CBEB8FFC2AE1DFB5BAB38C848FD54" box="[1210,1336,620,645]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="diabolos" status="sp. nov.">
|
||
<emphasis id="B9781929FFC2AE1DFB5BAB38C848FD54" box="[1210,1336,620,645]" italics="true" pageId="23" pageNumber="224">T. diabolos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24BA452FFC2AE1DFAA0AB39C8ECFD54" box="[1345,1436,620,645]" pageId="23" pageNumber="224" rank="species">
|
||
<emphasis id="B9781929FFC2AE1DFAA0AB39C8ECFD54" bold="true" box="[1345,1436,620,645]" pageId="23" pageNumber="224">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
intermediate in genital characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |