treatments-xml/data/CF/6F/49/CF6F4925FFB0FF82A663FAC4E2A48FD1.xml

<document id="F92C4B40D0E68CF49BC6B2DED2CDABCD" ID-CLB-Dataset="5814" ID-DOI="10.2478/aemnp-2018-0027" ID-GBIF-Dataset="6677b4e9-a47f-405d-8d1f-579c4996cba5" ID-ISSN="1804-6487" ID-Zenodo-Dep="4504756" ID-ZooBank="72E7EE6C-593C-4ECD-A400-40E7851720C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612473151739" checkinUser="felipe" docAuthor="Yin, Zi-Wei" docDate="2018" docId="CF6F4925FFB0FF82A663FAC4E2A48FD1" docLanguage="en" docName="ActaEntMusNatPra.58.2.321-330.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2)" docSource="http://dx.doi.org/10.2478/aemnp-2018-0027" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Cuccodorodes Yin 2018, gen. nov." docType="treatment" docVersion="4" lastPageNumber="326" masterDocId="3356315DFFB1FF87A727FA35E05C8931" masterDocTitle="Cuccodorodes gen. nov., an apterous Batrisitae from the tropical Himalayas (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="330" masterPageNumber="321" pageNumber="322" updateTime="1698927370163" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="534F278C84C4B75325A7262E4752826E">Cuccodorodes gen. nov., an apterous Batrisitae from the tropical Himalayas (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="5D962CF122E23E13AD157275B1D04272">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="BF618AAB98999A4D15EDEC4A16DCEC49">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
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<treatment id="CF6F4925FFB0FF82A663FAC4E2A48FD1" ID-DOI="http://doi.org/10.5281/zenodo.5061035" ID-GBIF-Taxon="194787284" ID-Zenodo-Dep="5061035" LSID="urn:lsid:plazi:treatment:CF6F4925FFB0FF82A663FAC4E2A48FD1" httpUri="http://treatment.plazi.org/id/CF6F4925FFB0FF82A663FAC4E2A48FD1" lastPageId="5" lastPageNumber="326" pageId="1" pageNumber="322">
<subSubSection id="0FDCABB8FFB0FF86A663FAC4E2178838" box="[324,587,241,265]" pageId="1" pageNumber="322" type="nomenclature">
<paragraph id="4779F833FFB0FF86A663FAC4E2178838" blockId="1.[324,587,241,265]" box="[324,587,241,265]" pageId="1" pageNumber="322">
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<taxonomicName id="80C683B0FFB0FF86A663FAC4E1BC8838" ID-CoL="8XKP9" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[324,480,241,265]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="75B22421FFB0FF86A663FAC4E1BC8838" bold="true" box="[324,480,241,265]" italics="true" pageId="1" pageNumber="322">Cuccodorodes</emphasis>
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<taxonomicNameLabel id="6E81995AFFB0FF86A6C1FAC7E2178838" box="[486,587,242,265]" pageId="1" pageNumber="322" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="4779F833FFB0FF86A6BCFB25E1A88812" blockId="1.[411,500,272,291]" box="[411,500,272,291]" pageId="1" pageNumber="322">
(
<figureCitation id="DFFDE4B6FFB0FF86A686FB25E1B18812" box="[417,493,272,291]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="1.[151,182,2057,2076]" captionStartId-1="2.[151,182,2057,2076]" captionStartId-2="3.[151,182,1339,1358]" captionStartId-3="4.[151,182,1562,1581]" captionStartId-4="5.[151,182,1286,1305]" captionStartId-5="6.[151,182,2030,2049]" captionTargetBox-0="[185,1402,785,2041]" captionTargetBox-1="[167,1421,756,2017]" captionTargetBox-2="[151,1436,189,1320]" captionTargetBox-3="[239,1349,192,1543]" captionTargetBox-4="[153,1435,204,1259]" captionTargetBox-5="[155,1436,199,1888]" captionTargetId-0="figure-276@1.[180,1407,775,2045]" captionTargetId-1="figure-291@2.[151,1436,746,2021]" captionTargetId-2="figure-361@3.[151,1436,188,1321]" captionTargetId-3="figure-173@4.[239,1349,189,1546]" captionTargetId-4="figure-330@5.[151,1436,197,1262]" captionTargetId-5="figure-13@6.[151,1436,195,1893]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionText-0="Fig. 1. Dorsal habitus of Cuccodorodes darjeelingensis sp. nov. A – male; B – female." captionText-1="Fig. 2. Dorsal habitus of Cuccodorodes species. A – C. koshiensis sp. nov., male; B – C. weiperti sp. nov., male." captionText-2="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." captionText-3="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." captionText-4="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." captionText-5="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi-0="http://doi.org/10.5281/zenodo.4504758" figureDoi-1="http://doi.org/10.5281/zenodo.4504760" figureDoi-2="http://doi.org/10.5281/zenodo.4504762" figureDoi-3="http://doi.org/10.5281/zenodo.4504764" figureDoi-4="http://doi.org/10.5281/zenodo.4504766" figureDoi-5="http://doi.org/10.5281/zenodo.4504768" httpUri-0="https://zenodo.org/record/4504758/files/figure.png" httpUri-1="https://zenodo.org/record/4504760/files/figure.png" httpUri-2="https://zenodo.org/record/4504762/files/figure.png" httpUri-3="https://zenodo.org/record/4504764/files/figure.png" httpUri-4="https://zenodo.org/record/4504766/files/figure.png" httpUri-5="https://zenodo.org/record/4504768/files/figure.png" pageId="1" pageNumber="322">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="4779F833FFB0FF86A7B0FB0DE146885E" blockId="1.[151,760,312,719]" pageId="1" pageNumber="322">
<emphasis id="75B22421FFB0FF86A7B0FB0DE174887E" bold="true" box="[151,296,312,335]" pageId="1" pageNumber="322">
<typeStatus id="987D4691FFB0FF86A7B0FB0DE092887E" box="[151,206,312,335]" pageId="1" pageNumber="322">Type</typeStatus>
species.
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<taxonomicName id="80C683B0FFB0FF86A60BFB0DE2358861" ID-CoL="6BNR7" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[300,617,312,336]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
<emphasis id="75B22421FFB0FF86A60BFB0DE2358861" box="[300,617,312,336]" italics="true" pageId="1" pageNumber="322">Cuccodorodes darjeelingensis</emphasis>
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<taxonomicNameLabel id="6E81995AFFB0FF86A549FB0DE2E2887E" box="[622,702,312,335]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
(here designated).
</paragraph>
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<subSubSection id="0FDCABB8FFB0FF85A7B0FB4DE2478824" lastPageId="2" lastPageNumber="323" pageId="1" pageNumber="322" type="diagnosis">
<paragraph id="4779F833FFB0FF85A7B0FB4DE2478824" blockId="1.[151,760,312,719]" lastBlockId="2.[151,760,190,694]" lastPageId="2" lastPageNumber="323" pageId="1" pageNumber="322">
<emphasis id="75B22421FFB0FF86A7B0FB4DE1C388BE" bold="true" box="[151,415,376,399]" pageId="1" pageNumber="322">Comparative diagnosis.</emphasis>
At present 
<taxonomicName id="80C683B0FFB0FF86A532FB4DE2F088A1" authorityName="Yin" authorityYear="2018" box="[533,684,376,400]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="genus">
<emphasis id="75B22421FFB0FF86A532FB4DE2F088A1" box="[533,684,376,400]" italics="true" pageId="1" pageNumber="322">Cuccodorodes</emphasis>
</taxonomicName>
cannot be placed near any known genus of the Indomalayan 
<taxonomicName id="80C683B0FFB0FF86A59DFBADE09C88FE" authorityName="Yin" authorityYear="2018" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
. The relationship of the new genus to other members of the group is difficult to discern given the extensive amount of homoplasy in Batrisitae (
<bibRefCitation id="235785C2FFB0FF86A6F0FBCDE2B08B3E" author="LOBL I. &amp; KURBATOV S. A." box="[471,748,504,527]" firstAuthor="LOBL" journalOrPublisher="Revue Suisse de Zoologie" pageId="1" pageNumber="322" pagination="559 - 697" part="108" refId="ref4593" refString="LOBL I. &amp; KURBATOV S. A. 2001: The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae). Revue Suisse de Zoologie 108: 559 - 697." title="The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2001">LÖBL &amp; KURBATOV 2001</bibRefCitation>
). Members of 
<taxonomicName id="80C683B0FFB0FF86A603F82DE1E78B01" authorityName="Yin" authorityYear="2018" box="[292,443,536,560]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="genus">
<emphasis id="75B22421FFB0FF86A603F82DE1E78B01" box="[292,443,536,560]" italics="true" pageId="1" pageNumber="322">Cuccodorodes</emphasis>
</taxonomicName>
are probably confined to the Himalayan region due to the complete absence of functional wings. The concept of the new genus is based on a unique combination of many peculiar external characters, notwithstanding that these may be homoplasious in other batrisine genera: head rounded-triangular in shape, lacking distinct frontal rostrum, vertex with a distinct reversed U-shaped sulcus connecting nude foveae, temples rounded, eyes small; antennal clubs loosely assembled by apical three antennomeres. Pronotum roundly expanded laterally, narrowing at the apex and base, disc with three basolateral foveae, and five longitudinal sulci, lateral pair of sulci strongly sinuate, densely setose and broadening in basal half, overlapping small lateral antebasal foveae, transverse antebasal sulcus absent, lacking antebasal and discal tubercles, ventral surface of pronotal lateral expansion deeply sulcate. Elytra small, much narrower than abdomen, strongly constricted at the base, each elytron with single basal fovea, lacking discal striae; lacking metathoracic wings; abdomen large, all foveae densely setose; tergite 1 (IV) longest, much longer than tergite 2 (V), with thin, oblique inner and thick outer marginal carinae, tergite 1 (IV) with four, and sternite 2 (IV) with eight basal foveae which are connecting internally. Male has mesotibia spinose at the apex, sternite 2 (IV) decorated with arched median projection. Female has strongly sclerotized genital complex.
</paragraph>
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<paragraph id="4779F833FFB0FF86A7B0F23CE32A812D" blockId="1.[151,886,2056,2076]" box="[151,886,2056,2076]" pageId="1" pageNumber="322">
Fig. 1. Dorsal habitus of 
<taxonomicName id="80C683B0FFB0FF86A64BF23DE232812A" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[364,622,2056,2075]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
<emphasis id="75B22421FFB0FF86A64BF23DE232812A" box="[364,622,2056,2075]" italics="true" pageId="1" pageNumber="322">Cuccodorodes darjeelingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB0FF86A554F23CE2EA812D" box="[627,694,2057,2076]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
A – male; B – female.
</paragraph>
</caption>
<subSubSection id="0FDCABB8FFB3FF84A7B0FB2BE57D812D" lastPageId="3" lastPageNumber="324" pageId="2" pageNumber="323" type="description">
<paragraph id="4779F833FFB3FF85A7B0FB2BE2618844" blockId="2.[151,760,190,694]" pageId="2" pageNumber="323">
<emphasis id="75B22421FFB3FF85A7B0FB2BE1798804" bold="true" box="[151,293,286,309]" pageId="2" pageNumber="323">Description.</emphasis>
Body length 
<quantity id="803E55D6FFB3FF85A69CFB2BE2018804" box="[443,605,286,309]" metricMagnitude="-3" metricUnit="m" metricValue="3.0949999999999998" metricValueMax="3.26" metricValueMin="2.93" pageId="2" pageNumber="323" unit="mm" value="3.0949999999999998" valueMax="3.26" valueMin="2.93">2.93–3.26 mm</quantity>
; habitus (
<figureCitation id="DFFDE4B6FFB3FF85A5E0FB2BE09C8864" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[151,182,2057,2076]" captionStartId-1="2.[151,182,2057,2076]" captionTargetBox-0="[185,1402,785,2041]" captionTargetBox-1="[167,1421,756,2017]" captionTargetId-0="figure-276@1.[180,1407,775,2045]" captionTargetId-1="figure-291@2.[151,1436,746,2021]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Dorsal habitus of Cuccodorodes darjeelingensis sp. nov. A – male; B – female." captionText-1="Fig. 2. Dorsal habitus of Cuccodorodes species. A – C. koshiensis sp. nov., male; B – C. weiperti sp. nov., male." figureDoi-0="http://doi.org/10.5281/zenodo.4504758" figureDoi-1="http://doi.org/10.5281/zenodo.4504760" httpUri-0="https://zenodo.org/record/4504758/files/figure.png" httpUri-1="https://zenodo.org/record/4504760/files/figure.png" pageId="2" pageNumber="323">Figs 1–2</figureCitation>
) stout, antennae relatively short, not reaching half elytral length when angled posteriorly.
</paragraph>
<paragraph id="4779F833FFB3FF85A792FB4BE2E48B84" blockId="2.[151,760,190,694]" pageId="2" pageNumber="323">
Head (
<figureCitation id="DFFDE4B6FFB3FF85A626FB4BE1E388A4" box="[257,447,382,405]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="2" pageNumber="323">Figs 3A, 3C, 3E</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A6E8FB4BE18088A4" box="[463,476,382,405]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">4</figureCitation>
A–C) rounded-triangular; lacking distinct frontal rostrum, antennal tubercles strongly raised; vertexal foveae (
<figureCitation id="DFFDE4B6FFB3FF85A6BCFB8BE1AE88E4" box="[411,498,446,469]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A6D9FB8BE24D88E7" box="[510,529,446,470]" italics="true" pageId="2" pageNumber="323">vf</emphasis>
) nude and small, connected by complete reversed U or V-shaped sulcus (
<figureCitation id="DFFDE4B6FFB3FF85A5EBFBEBE0E68B24" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A7E0FBCBE0818B24" box="[199,221,510,533]" italics="true" pageId="2" pageNumber="323">vs</emphasis>
); with 11 antennomeres, clubs loosely formed by three apical antennomeres, antennomeres XI elongate and conical; ocular-mandibular carinae (
<figureCitation id="DFFDE4B6FFB3FF85A528F80BE23C8B64" box="[527,608,574,597]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4B</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A54EF80BE2C98B64" box="[617,661,574,597]" italics="true" pageId="2" pageNumber="323">omc</emphasis>
) distinct; eyes small; maxillary palpus with trapezoid palpomeres III, palpomeres IV narrowed at apex and base; punctiform gular foveae (
<figureCitation id="DFFDE4B6FFB3FF85A60DF8ABE1228B84" box="[298,382,670,693]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4C</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A6AEF8ABE1C28B87" box="[393,414,670,694]" italics="true" pageId="2" pageNumber="323">gf</emphasis>
) in shared round opening.
</paragraph>
<paragraph id="4779F833FFB3FF85A47DFA8BE5038B84" blockId="2.[828,1436,190,693]" pageId="2" pageNumber="323">
Pronotum (
<figureCitation id="DFFDE4B6FFB3FF85A4F0FA8BE4C889E4" box="[983,1172,190,213]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="2" pageNumber="323">Figs 3A, 3C, 3E</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A383FA8BE49A89E4" box="[1188,1222,190,213]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">4D</figureCitation>
) transverse, lateral margins roundly expanded near middle, narrowing at apex and base, anterior and posterior margins slightly curved; lateral antebasal foveae (
<figureCitation id="DFFDE4B6FFB3FF85A39CFB2BE5448804" box="[1211,1304,286,309]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">Fig. 5A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A200FB28E5188804" box="[1319,1348,285,309]" italics="true" pageId="2" pageNumber="323">laf</emphasis>
) small, covered by dense setae and not visible in dry specimens, median antebasal fovea (
<figureCitation id="DFFDE4B6FFB3FF85A36FFB6BE4C28844" box="[1096,1182,350,373]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4D</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A38DFB6BE48E8847" box="[1194,1234,350,374]" italics="true" pageId="2" pageNumber="323">maf</emphasis>
) tiny and obscure, lacking discal or antebasal spines/tubercles; disc convex; with three pairs of basolateral foveae (
<figureCitation id="DFFDE4B6FFB3FF85A3F8FBABE5668884" box="[1247,1338,414,437]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Figs 4D</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A260FBABE5368884" box="[1351,1386,414,437]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">5A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A250FBA8E5CF8884" box="[1399,1427,413,437]" italics="true" pageId="2" pageNumber="323">blf</emphasis>
) and one pair of basolateral pits (
<figureCitation id="DFFDE4B6FFB3FF85A3A9FB8BE4A688E4" box="[1166,1274,446,469]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">
Fig. 5A; 
<emphasis id="75B22421FFB3FF85A3CAFB8BE4A688E4" box="[1261,1274,446,469]" italics="true" pageId="2" pageNumber="323">p</emphasis>
</figureCitation>
); median (
<figureCitation id="DFFDE4B6FFB3FF85A24AFB8BE30388C4" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Figs 4D</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A44DFBEBE3D188C4" box="[874,909,478,501]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">5A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A4BDFBE8E3E388C4" box="[922,959,477,501]" italics="true" pageId="2" pageNumber="323">mls</emphasis>
), discal (
<figureCitation id="DFFDE4B6FFB3FF85A304FBEBE42188C4" box="[1059,1149,478,501]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Figs 4D</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A3AFFBEBE4F788C4" box="[1160,1195,478,501]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">5A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A390FBE8E48A88C4" box="[1207,1238,477,501]" italics="true" pageId="2" pageNumber="323">dls</emphasis>
), and lateral (
<figureCitation id="DFFDE4B6FFB3FF85A24AFBEBE3038B24" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Figs 4D</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB3FF85A44EFBCBE3D08B24" box="[873,908,510,533]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="2" pageNumber="323">5A</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A4B1FBC8E3F38B24" box="[918,943,509,533]" italics="true" pageId="2" pageNumber="323">lls</emphasis>
) longitudinal sulci present, lateral pair of sulci strongly sinuate, densely setose and broadening in basal half, lacking transverse antebasal sulcus; prosternite with distinct paranotal sulci (
<figureCitation id="DFFDE4B6FFB3FF85A319F86BE4CC8B44" box="[1086,1168,606,629]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4E</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A3BBF86BE4E88B44" box="[1180,1204,606,629]" italics="true" pageId="2" pageNumber="323">ps</emphasis>
); ventral side of lateral expansion deeply sulcate; opening of lateral procoxal foveae (
<figureCitation id="DFFDE4B6FFB3FF85A4B6F8ABE3B88B84" box="[913,996,670,693]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="2" pageNumber="323">Fig. 4E</figureCitation>
; 
<emphasis id="75B22421FFB3FF85A4C8F8A8E44B8B84" box="[1007,1047,669,693]" italics="true" pageId="2" pageNumber="323">lpcf</emphasis>
) at base of broad, setose sulci.
</paragraph>
<caption id="13B9A8BBFFB3FF85A7B0F23CE40D812D" ID-DOI="http://doi.org/10.5281/zenodo.4504760" ID-Zenodo-Dep="4504760" box="[151,1105,2056,2076]" httpUri="https://zenodo.org/record/4504760/files/figure.png" pageId="2" pageNumber="323" startId="2.[151,182,2057,2076]" targetBox="[167,1421,756,2017]" targetPageId="2">
<paragraph id="4779F833FFB3FF85A7B0F23CE40D812D" blockId="2.[151,1105,2056,2076]" box="[151,1105,2056,2076]" pageId="2" pageNumber="323">
Fig. 2. Dorsal habitus of 
<taxonomicName id="80C683B0FFB3FF85A64BF23DE1BA812A" authorityName="Yin" authorityYear="2018" box="[364,486,2056,2075]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="75B22421FFB3FF85A64BF23DE1BA812A" box="[364,486,2056,2075]" italics="true" pageId="2" pageNumber="323">Cuccodorodes</emphasis>
</taxonomicName>
species. A – 
<taxonomicName id="80C683B0FFB3FF85A571F23CE29B812A" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[598,711,2056,2076]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB3FF85A571F23CE29B812A" box="[598,711,2056,2076]" italics="true" pageId="2" pageNumber="323">C. koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB3FF85A5EBF23CE34D812D" box="[716,785,2057,2076]" pageId="2" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
, male; B – 
<taxonomicName id="80C683B0FFB3FF85A456F23CE38C812A" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[881,976,2057,2076]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="weiperti" status="sp. nov.">
<emphasis id="75B22421FFB3FF85A456F23CE38C812A" box="[881,976,2057,2076]" italics="true" pageId="2" pageNumber="323">C. weiperti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB3FF85A4F2F23CE446812D" box="[981,1050,2057,2076]" pageId="2" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
, male.
</paragraph>
</caption>
<caption id="13B9A8BBFFB2FF84A7B0FF0EE1BC8C59" ID-DOI="http://doi.org/10.5281/zenodo.4504762" ID-Zenodo-Dep="4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="3" pageNumber="324" startId="3.[151,182,1339,1358]" targetBox="[151,1436,189,1320]" targetPageId="3">
<paragraph id="4779F833FFB2FF84A7B0FF0EE1BC8C59" blockId="3.[151,1438,1338,1384]" pageId="3" pageNumber="324">
Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of 
<taxonomicName id="80C683B0FFB2FF84A475FF0FE4088C7C" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[850,1108,1338,1357]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
<emphasis id="75B22421FFB2FF84A475FF0FE4088C7C" box="[850,1108,1338,1357]" italics="true" pageId="3" pageNumber="324">Cuccodorodes darjeelingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB2FF84A37DFF0EE4C28C7F" box="[1114,1182,1339,1358]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
(A–B), 
<taxonomicName id="80C683B0FFB2FF84A3C4FF0EE5098C7C" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[1251,1365,1338,1358]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB2FF84A3C4FF0EE5098C7C" box="[1251,1365,1338,1358]" italics="true" pageId="3" pageNumber="324">C. koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB2FF84A27DFF0EE5C28C7F" box="[1370,1438,1339,1358]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
(C–D), and 
<taxonomicName id="80C683B0FFB2FF84A7DBFF60E1078C59" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[252,347,1365,1384]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="weiperti" status="sp. nov.">
<emphasis id="75B22421FFB2FF84A7DBFF60E1078C59" box="[252,347,1365,1384]" italics="true" pageId="3" pageNumber="324">C. weiperti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB2FF84A646FF60E1F88C59" box="[353,420,1365,1384]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
(E–F).
</paragraph>
</caption>
<paragraph id="4779F833FFB2FF84A792FF90E16C8F0D" blockId="3.[151,760,1445,2076]" pageId="3" pageNumber="324">
Elytra (
<figureCitation id="DFFDE4B6FFB2FF84A62DFF90E1328C8D" box="[266,366,1445,1468]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[151,182,2057,2076]" captionStartId-1="2.[151,182,2057,2076]" captionTargetBox-0="[185,1402,785,2041]" captionTargetBox-1="[167,1421,756,2017]" captionTargetId-0="figure-276@1.[180,1407,775,2045]" captionTargetId-1="figure-291@2.[151,1436,746,2021]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Dorsal habitus of Cuccodorodes darjeelingensis sp. nov. A – male; B – female." captionText-1="Fig. 2. Dorsal habitus of Cuccodorodes species. A – C. koshiensis sp. nov., male; B – C. weiperti sp. nov., male." figureDoi-0="http://doi.org/10.5281/zenodo.4504758" figureDoi-1="http://doi.org/10.5281/zenodo.4504760" httpUri-0="https://zenodo.org/record/4504758/files/figure.png" httpUri-1="https://zenodo.org/record/4504760/files/figure.png" pageId="3" pageNumber="324">Figs 1–2</figureCitation>
) strongly constricted at base; each elytron (
<figureCitation id="DFFDE4B6FFB2FF84A7D7FFF0E1638CED" box="[240,319,1477,1500]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="3" pageNumber="324">Fig. 4F</figureCitation>
) with one basal fovea (
<figureCitation id="DFFDE4B6FFB2FF84A515FFF0E2DF8CED" box="[562,643,1477,1500]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="3" pageNumber="324">Fig. 4F</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A5AAFFF1E2F28CED" box="[653,686,1476,1500]" italics="true" pageId="3" pageNumber="324">bef</emphasis>
), lacking subbasal fovea, lacking discal stria, with complete sutural stria, thin marginal carina present in posterior half of lateral side.
</paragraph>
<paragraph id="4779F833FFB2FF84A792FC70E1668FCD" blockId="3.[151,760,1445,2076]" pageId="3" pageNumber="324">
Mesoventrite with median foveae (
<figureCitation id="DFFDE4B6FFB2FF84A513FC70E2D78F6D" box="[564,651,1605,1628]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5B</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A5BFFC71E28C8F6D" box="[664,720,1604,1628]" italics="true" pageId="3" pageNumber="324">mmsf</emphasis>
) in transverse opening, with large lateral mesoventral foveae unforked internally (
<figureCitation id="DFFDE4B6FFB2FF84A657FCB0E19E8FAD" box="[368,450,1669,1692]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5B</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A6EBFCB1E1A48FAD" box="[460,504,1668,1692]" italics="true" pageId="3" pageNumber="324">lmsf</emphasis>
), with oval mesoventral foveae, metaventrite with lateral foveae (
<figureCitation id="DFFDE4B6FFB2FF84A57DFC90E2ED8F8D" box="[602,689,1701,1724]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5B</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A598FC91E2B48F8D" box="[703,744,1700,1724]" italics="true" pageId="3" pageNumber="324">lmtf</emphasis>
), posterior margin with narrow notch at middle; metacoxae well-separated.
</paragraph>
<paragraph id="4779F833FFB2FF84A792FD30E4D18FAD" blockId="3.[151,760,1445,2076]" lastBlockId="3.[828,1437,1445,2076]" pageId="3" pageNumber="324">
Abdomen with lateral margins of visible tergite 1 (IV) produced as thick, triangular ridge edged by thin inner and thick marginal carinae; tergites 2–3 (V–VI) with lateral margins similarly edged; tergite 4–5 (VII–VIII) lacking marginal carinae; tergite 1 (IV) longest, 2–3 (V–VI) distinctly shorter and subequal in length, 4 (VII) longer than 3 (VI); tergite 1 (IV) with one pair of mediobasal (
<figureCitation id="DFFDE4B6FFB2FF84A5EBFDF0E0E58ECD" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5C</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A7E3FDD1E0B08ECD" box="[196,236,2020,2044]" italics="true" pageId="3" pageNumber="324">mbf</emphasis>
) and one pair of basolateral foveae (
<figureCitation id="DFFDE4B6FFB2FF84A549FDD0E29D8ECD" box="[622,705,2021,2044]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5C</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A5EBFDD1E2B48ECD" box="[716,744,2020,2044]" italics="true" pageId="3" pageNumber="324">blf</emphasis>
), lacking discal carinae and basal impression, tergites 2–4 (V–VII) each with one pair of basolateral foveae. Visible sternite 2 (IV) with one pair of mediobasal (
<figureCitation id="DFFDE4B6FFB2FF84A22AFFF0E53C8CED" box="[1293,1376,1477,1500]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5C</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A24CFFF1E5CF8CED" box="[1387,1427,1476,1500]" italics="true" pageId="3" pageNumber="324">mbf</emphasis>
) and three pairs of basolateral foveae (
<figureCitation id="DFFDE4B6FFB2FF84A39AFFD0E5518CCD" box="[1213,1293,1509,1532]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5C</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A230FFD1E56F8CCD" box="[1303,1331,1508,1532]" italics="true" pageId="3" pageNumber="324">blf</emphasis>
) in broad, setose basolateral sulci, sternites 3–4 (V–VI) each with two pairs of basolateral foveae, and sternite 5 (VII) with one pair of basolateral foveae. All foveae of abdomen densely setose, those of tergite 1 (IV) and sternite 2 (IV) deep and connecting internally (
<figureCitation id="DFFDE4B6FFB2FF84A30BFCB0E4DD8FAD" box="[1068,1153,1669,1692]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="4779F833FFB2FF84A47DFC90E3FD8FED" blockId="3.[828,1437,1445,2076]" pageId="3" pageNumber="324">Legs relatively stout; tarsomeres II and III subequal in length.</paragraph>
<paragraph id="4779F833FFB2FF84A47DFCD0E57D812D" blockId="3.[828,1437,1445,2076]" pageId="3" pageNumber="324">
Male has relatively longer antennae than female (
<figureCitation id="DFFDE4B6FFB2FF84A256FCD0E3148E2D" captionStart="Fig" captionStartId="1.[151,182,2057,2076]" captionTargetBox="[185,1402,785,2041]" captionTargetId="figure-276@1.[180,1407,775,2045]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of Cuccodorodes darjeelingensis sp. nov. A – male; B – female." figureDoi="http://doi.org/10.5281/zenodo.4504758" httpUri="https://zenodo.org/record/4504758/files/figure.png" pageId="3" pageNumber="324">Fig. 1</figureCitation>
); mesotibia with one tubercle (
<figureCitation id="DFFDE4B6FFB2FF84A3ABFD30E4838E2D" box="[1164,1247,1797,1820]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="3" pageNumber="324">Fig. 4G</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A3CFFD30E4A18E2D" box="[1256,1277,1797,1820]" italics="true" pageId="3" pageNumber="324">at</emphasis>
) and two small spines (
<figureCitation id="DFFDE4B6FFB2FF84A4AAFD10E3BE8E0D" box="[909,994,1829,1852]" captionStart="Fig" captionStartId="4.[151,182,1562,1581]" captionTargetBox="[239,1349,192,1543]" captionTargetId="figure-173@4.[239,1349,189,1546]" captionTargetPageId="4" captionText="Fig. 4. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="3" pageNumber="324">Fig. 4G</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A4CAFD10E4598E0D" box="[1005,1029,1829,1852]" italics="true" pageId="3" pageNumber="324">as</emphasis>
) at apex; sternite 2 (IV) (
<figureCitation id="DFFDE4B6FFB2FF84A234FD10E3058E6D" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="3" pageNumber="324">Figs 3B, 3D, 3F</figureCitation>
) modified with arched median projection (
<figureCitation id="DFFDE4B6FFB2FF84A231FD70E3048E4D" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="3" pageNumber="324">Figs 3B, 3D, 3F</figureCitation>
, 
<figureCitation id="DFFDE4B6FFB2FF84A443FD50E3D98E4D" box="[868,901,1893,1916]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">5C</figureCitation>
; 
<emphasis id="75B22421FFB2FF84A4B5FD50E3EF8E4D" box="[914,947,1893,1916]" italics="true" pageId="3" pageNumber="324">mp</emphasis>
); sternite 7 (IX) semi-membranous (
<figureCitation id="DFFDE4B6FFB2FF84A219FD50E3CC8EAD" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="3" pageNumber="324">Figs 6A, 6D, 6G</figureCitation>
); aedeagus (
<figureCitation id="DFFDE4B6FFB2FF84A33FFDB0E4018EAD" box="[1048,1117,1925,1948]" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="3" pageNumber="324">Figs 6</figureCitation>
B–C, 6E–F, 6H–I) with parameres fused to median lobe to form elongate ventral lobe, with large basal capsule and foramen, with well-developed basoventral projection. Female genital complex composed of strongly sclerotized structure (
<figureCitation id="DFFDE4B6FFB2FF84A3BBF230E482812D" box="[1180,1246,2053,2076]" captionStart="Fig" captionStartId="5.[151,182,1286,1305]" captionTargetBox="[153,1435,204,1259]" captionTargetId="figure-330@5.[151,1436,197,1262]" captionTargetPageId="5" captionText="Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – meso- and metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="3" pageNumber="324">Figs 5</figureCitation>
D–E).
</paragraph>
</subSubSection>
<caption id="13B9A8BBFFB5FF83A7B0FC2FE1068FA6" ID-DOI="http://doi.org/10.5281/zenodo.4504764" ID-Zenodo-Dep="4504764" httpUri="https://zenodo.org/record/4504764/files/figure.png" pageId="4" pageNumber="325" startId="4.[151,182,1562,1581]" targetBox="[239,1349,192,1543]" targetPageId="4">
<paragraph id="4779F833FFB5FF83A7B0FC2FE1068FA6" blockId="4.[151,1437,1561,1687]" pageId="4" pageNumber="325">
Fig. 4. Morphology of 
<taxonomicName id="80C683B0FFB5FF83A671FC2CE18C8F1D" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[342,464,1561,1580]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="75B22421FFB5FF83A671FC2CE18C8F1D" box="[342,464,1561,1580]" italics="true" pageId="4" pageNumber="325">Cuccodorodes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB5FF83A6F2FC2FE2788F1C" box="[469,548,1562,1581]" pageId="4" pageNumber="325" rank="genus">gen. nov.</taxonomicNameLabel>
, based on 
<taxonomicName id="80C683B0FFB5FF83A55FFC2FE2B48F1D" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[632,744,1561,1581]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB5FF83A55FFC2FE2B48F1D" box="[632,744,1561,1581]" italics="true" pageId="4" pageNumber="325">C. koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB5FF83A5CBFC2FE3728F1C" box="[748,814,1562,1581]" pageId="4" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
A–C – head in dorsal (A), lateral (B), and ventral (C) view; D – pronotum; E – prosternite; F – left elytron; G – apex of male mesotibia.Abbreviations: as – apical spine; at – apical tooth; bef – basal elytral fovea; blf – basolateral foveae; dls – discal longitudinal sulcus; gf – gular foveae; lls – lateral longitudinal sulcus; lpcf – lateral procoxal foveae; maf – median antebasal fovea; mls – median longitudinal sulcus; omc – ocular-mandibular carina; ps – paranotal sulcus; vf – vertex foveae; vs – vertexal sulcus. Scale bars: 0.2 mm in A–F; 0.05 mm in G.
</paragraph>
</caption>
<subSubSection id="0FDCABB8FFB5FF83A7B0FCF1E2078E2A" pageId="4" pageNumber="325" type="etymology">
<paragraph id="4779F833FFB5FF83A7B0FCF1E2078E2A" blockId="4.[151,760,1732,1819]" pageId="4" pageNumber="325">
<emphasis id="75B22421FFB5FF83A7B0FCF1E14A8FEA" bold="true" box="[151,278,1732,1755]" pageId="4" pageNumber="325">Etymology.</emphasis>
The new genus is named in honor of Dr. Giulio Cuccodoro, who has offered support and help during my visits to 
<collectingRegion id="850236D1FFB5FF83A7D5FD31E11B8E2A" box="[242,327,1796,1819]" country="Switzerland" name="Geneve" pageId="4" pageNumber="325">Geneva</collectingRegion>
. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="0FDCABB8FFB5FF83A639FD6FE2EA8111" pageId="4" pageNumber="325" type="discussion">
<paragraph id="4779F833FFB5FF83A639FD6FE22D8E40" blockId="4.[286,625,1881,1905]" box="[286,625,1881,1905]" pageId="4" pageNumber="325">
<heading id="1C314F5FFFB5FF83A639FD6FE22D8E40" box="[286,625,1881,1905]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="325" reason="3">
<emphasis id="75B22421FFB5FF83A639FD6FE22D8E40" bold="true" box="[286,625,1881,1905]" pageId="4" pageNumber="325">
Key to males of 
<taxonomicName id="80C683B0FFB5FF83A6F2FD6CE22D8E40" authorityName="Yin" authorityYear="2018" box="[469,625,1881,1905]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="75B22421FFB5FF83A6F2FD6CE22D8E40" bold="true" box="[469,625,1881,1905]" italics="true" pageId="4" pageNumber="325">Cuccodorodes</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="4779F833FFB5FF83A7B0FDBCE2EA8111" blockId="4.[151,760,1929,2080]" pageId="4" pageNumber="325">
<emphasis id="75B22421FFB5FF83A7B0FDBCE0808E91" bold="true" box="[151,220,1929,1952]" pageId="4" pageNumber="325">Notes.</emphasis>
The three new species described below are exceptionally similar in external morphology, and probably restricted to a small area in eastern 
<collectingCountry id="3FD1B8A3FFB5FF83A6BCFDFCE1868ED1" box="[411,474,1993,2016]" name="Nepal" pageId="4" pageNumber="325">Nepal</collectingCountry>
and northern 
<collectingCountry id="3FD1B8A3FFB5FF83A54EFDFCE2C38ED1" box="[617,671,1993,2016]" name="India" pageId="4" pageNumber="325">India</collectingCountry>
(
<figureCitation id="DFFDE4B6FFB5FF83A58BFDFCE2B68ED1" box="[684,746,1993,2016]" captionStart="Fig" captionStartId="7.[1049,1082,757,776]" captionTargetBox="[152,1032,189,801]" captionTargetId="figure-635@7.[152,1035,189,801]" captionTargetPageId="7" captionText="Fig.7. Map showing the distribution of Cuccodorodes species." figureDoi="http://doi.org/10.5281/zenodo.4504770" httpUri="https://zenodo.org/record/4504770/files/figure.png" pageId="4" pageNumber="325">Fig. 7</figureCitation>
). Reliable species identification of additional material from this region requires an examination of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="0FDCABB8FFB5FF82A41BFCF1E2A48FD1" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325" type="key">
<keyStep id="FC32E096FFB5FF83A41BFCF1E5C08EEA" pageId="4" pageNumber="325">
<paragraph id="4779F833FFB5FF83A41BFCF1E5C08E6A" blockId="4.[828,1437,1732,2075]" pageId="4" pageNumber="325">
<keyLead id="FC375B06FFB5FF83A41BFCF1E5C08E6A" pageId="4" pageNumber="325">
1 Vertexal sulcus evenly convergent anteriorly, more or less V-shaped (
<figureCitation id="DFFDE4B6FFB5FF83A301FCD1E4208FCA" box="[1062,1148,1764,1787]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="4" pageNumber="325">Fig. 3C</figureCitation>
); sternite 2 (IV) with area surrounded by median projection broadly triangular (
<figureCitation id="DFFDE4B6FFB5FF83A44EFD11E3E38E0A" box="[873,959,1828,1851]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="4" pageNumber="325">Fig. 3D</figureCitation>
). (
<collectingCountry id="3FD1B8A3FFB5FF83A4FDFD11E4438E0A" box="[986,1055,1828,1851]" name="Nepal" pageId="4" pageNumber="325">Nepal</collectingCountry>
: 
<collectingRegion id="850236D1FFB5FF83A30EFD11E4308E0A" box="[1065,1132,1828,1851]" country="Nepal" name="Kosi" pageId="4" pageNumber="325">Koshi</collectingRegion>
; 
<figureCitation id="DFFDE4B6FFB5FF83A351FD11E4EB8E0A" box="[1142,1207,1828,1851]" captionStart="Fig" captionStartId="7.[1049,1082,757,776]" captionTargetBox="[152,1032,189,801]" captionTargetId="figure-635@7.[152,1035,189,801]" captionTargetPageId="7" captionText="Fig.7. Map showing the distribution of Cuccodorodes species." figureDoi="http://doi.org/10.5281/zenodo.4504770" httpUri="https://zenodo.org/record/4504770/files/figure.png" pageId="4" pageNumber="325">Fig. 7</figureCitation>
). ............................... ................................................. 
<emphasis id="75B22421FFB5FF83A388FD76E5C08E6A" bold="true" box="[1199,1436,1859,1883]" pageId="4" pageNumber="325">
<taxonomicName id="80C683B0FFB5FF83A388FD76E51C8E6A" ID-CoL="328TB" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[1199,1344,1859,1883]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB5FF83A388FD76E49D8E6A" bold="true" box="[1199,1217,1859,1883]" italics="true" pageId="4" pageNumber="325">C</emphasis>
. 
<emphasis id="75B22421FFB5FF83A3E9FD76E51C8E6A" bold="true" box="[1230,1344,1859,1883]" italics="true" pageId="4" pageNumber="325">koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB5FF83A260FD71E5C08E6A" box="[1351,1436,1860,1883]" pageId="4" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="4779F833FFB5FF83A41BFD51E5C08EEA" blockId="4.[828,1437,1732,2075]" pageId="4" pageNumber="325">
<keyLead id="FC375B06FFB5FF83A41BFD51E5C08EEA" pageId="4" pageNumber="325">
– Vertexal sulcus truncate anteriorly, more or less U-shaped (
<figureCitation id="DFFDE4B6FFB5FF83A4FCFDB1E4308EAA" box="[987,1132,1924,1947]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="4" pageNumber="325">Figs 3A, 3E</figureCitation>
); sternite 2 (IV) with area surrounded by median projection narrowly triangular (
<figureCitation id="DFFDE4B6FFB5FF83A44EFDF1E3B78EEA" box="[873,1003,1988,2011]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="4" pageNumber="325">Figs 3B, 3F</figureCitation>
). .......................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC32E096FFB5FF82A41BFDD1E2A48FD1" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325">
<paragraph id="4779F833FFB5FF82A41BFDD1E2A48F71" blockId="4.[828,1437,1732,2075]" lastBlockId="5.[151,760,1481,1760]" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325">
<keyLead id="FC375B06FFB5FF82A41BFDD1E2A48F71" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325">
2 Median area of sternite3 (V) distinctly microsculptured (
<figureCitation id="DFFDE4B6FFB5FF83A44EF231E39C812A" box="[873,960,2052,2075]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="4" pageNumber="325">Fig. 3B</figureCitation>
); membranous basal half of sternite 7 (IX) obliquely and lengthily elongate (
<figureCitation id="DFFDE4B6FFB4FF82A50DFFFCE2DD8CD1" box="[554,641,1481,1504]" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="5" pageNumber="326">Fig. 6A</figureCitation>
); aedeagus in ventral view lacking narrowly elongate sclerite at left side (
<figureCitation id="DFFDE4B6FFB4FF82A606FC3CE1298F11" box="[289,373,1545,1568]" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="5" pageNumber="326">Fig. 6C</figureCitation>
). (
<collectingCountry id="3FD1B8A3FFB4FF82A6B7FC3CE1908F11" box="[400,460,1545,1568]" name="India" pageId="5" pageNumber="326">India</collectingCountry>
: Darjeeling; 
<figureCitation id="DFFDE4B6FFB4FF82A573FC3CE2C98F11" box="[596,661,1545,1568]" captionStart="Fig" captionStartId="7.[1049,1082,757,776]" captionTargetBox="[152,1032,189,801]" captionTargetId="figure-635@7.[152,1035,189,801]" captionTargetPageId="7" captionText="Fig.7. Map showing the distribution of Cuccodorodes species." figureDoi="http://doi.org/10.5281/zenodo.4504770" httpUri="https://zenodo.org/record/4504770/files/figure.png" pageId="5" pageNumber="326">Fig. 7</figureCitation>
). ........... ........................................ 
<emphasis id="75B22421FFB4FF82A6F0FC1DE2A48F71" bold="true" box="[471,760,1576,1600]" pageId="5" pageNumber="326">
<taxonomicName id="80C683B0FFB4FF82A6F0FC1DE2C08F71" ID-CoL="6BNR7" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[471,668,1576,1600]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
<emphasis id="75B22421FFB4FF82A6F0FC1DE1B58F71" bold="true" box="[471,489,1576,1600]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="75B22421FFB4FF82A6D1FC1DE2C08F71" bold="true" box="[502,668,1576,1600]" italics="true" pageId="5" pageNumber="326">darjeelingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A585FC1CE2A48F71" box="[674,760,1577,1600]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<caption id="13B9A8BBFFB4FF82A7B0FF33E4E68CB5" ID-DOI="http://doi.org/10.5281/zenodo.4504766" ID-Zenodo-Dep="4504766" httpUri="https://zenodo.org/record/4504766/files/figure.png" pageId="5" pageNumber="326" startId="5.[151,182,1286,1305]" targetBox="[153,1435,204,1259]" targetPageId="5">
<paragraph id="4779F833FFB4FF82A7B0FF33E4E68CB5" blockId="5.[151,1437,1285,1412]" pageId="5" pageNumber="326">
Fig. 5. Morphology of 
<taxonomicName id="80C683B0FFB4FF82A672FF30E1938C29" ID-CoL="8XKP9" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[341,463,1285,1304]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="75B22421FFB4FF82A672FF30E1938C29" box="[341,463,1285,1304]" italics="true" pageId="5" pageNumber="326">Cuccodorodes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A6F3FF33E27F8C28" box="[468,547,1286,1305]" pageId="5" pageNumber="326" rank="genus">gen. nov.</taxonomicNameLabel>
, based on 
<taxonomicName id="80C683B0FFB4FF82A550FF33E2BA8C29" ID-CoL="328TB" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[631,742,1285,1305]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB4FF82A550FF33E2BA8C29" box="[631,742,1285,1305]" italics="true" pageId="5" pageNumber="326">C. koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A5CDFF33E3708C28" box="[746,812,1286,1305]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
unless specified otherwise.A – prothorax, in dorso-lateral view; B – mesoand metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – 
<taxonomicName id="80C683B0FFB4FF82A225FF15E5C08C02" ID-CoL="6BNR7" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[1282,1436,1312,1331]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="darjeelingensis" status="sp. nov.">
<emphasis id="75B22421FFB4FF82A225FF15E5C08C02" box="[1282,1436,1312,1331]" italics="true" pageId="5" pageNumber="326">C. darjeelingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A7B0FF0EE0818C7F" box="[151,221,1339,1358]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
; E – 
<taxonomicName id="80C683B0FFB4FF82A620FF0EE1248C7F" ID-CoL="328TB" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[263,376,1339,1358]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="koshiensis" status="sp. nov.">
<emphasis id="75B22421FFB4FF82A620FF0EE1248C7F" box="[263,376,1339,1358]" italics="true" pageId="5" pageNumber="326">C. koshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A659FF0EE1988C7F" box="[382,452,1339,1358]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="4779F833FFB4FF82A7B0FC7CE2A48FD1" blockId="5.[151,760,1481,1760]" pageId="5" pageNumber="326">
<keyLead id="FC375B06FFB4FF82A7B0FC7CE2A48FD1" pageId="5" pageNumber="326">
– Median area of sternite 3 (V) lacking microsculpture (
<figureCitation id="DFFDE4B6FFB4FF82A7E2FC5CE1458FB1" box="[197,281,1641,1664]" captionStart="Fig" captionStartId="3.[151,182,1339,1358]" captionTargetBox="[151,1436,189,1320]" captionTargetId="figure-361@3.[151,1436,188,1321]" captionTargetPageId="3" captionText="Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F)." figureDoi="http://doi.org/10.5281/zenodo.4504762" httpUri="https://zenodo.org/record/4504762/files/figure.png" pageId="5" pageNumber="326">Fig. 3F</figureCitation>
); membranous basal half of sternite 7 (IX) roundly expanded (
<figureCitation id="DFFDE4B6FFB4FF82A6ADFCBCE1838F91" box="[394,479,1673,1696]" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="5" pageNumber="326">Fig. 6G</figureCitation>
); aedeagus in ventral view with narrowly elongate sclerite at left side (
<figureCitation id="DFFDE4B6FFB4FF82A5B9FC9CE2B78FF1" box="[670,747,1705,1728]" captionStart="Fig" captionStartId="6.[151,182,2030,2049]" captionTargetBox="[155,1436,199,1888]" captionTargetId="figure-13@6.[151,1436,195,1893]" captionTargetPageId="6" captionText="Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I." figureDoi="http://doi.org/10.5281/zenodo.4504768" httpUri="https://zenodo.org/record/4504768/files/figure.png" pageId="5" pageNumber="326">Fig. 6I</figureCitation>
). (
<collectingCountry id="3FD1B8A3FFB4FF82A7E3FCFCE1558FD1" box="[196,265,1737,1760]" name="Nepal" pageId="5" pageNumber="326">Nepal</collectingCountry>
: 
<collectingRegion id="850236D1FFB4FF82A633FCFCE1008FD1" box="[276,348,1737,1760]" country="Nepal" name="Mechi" pageId="5" pageNumber="326">Mechi</collectingRegion>
; 
<figureCitation id="DFFDE4B6FFB4FF82A641FCFCE1FB8FD1" box="[358,423,1737,1760]" captionStart="Fig" captionStartId="7.[1049,1082,757,776]" captionTargetBox="[152,1032,189,801]" captionTargetId="figure-635@7.[152,1035,189,801]" captionTargetPageId="7" captionText="Fig.7. Map showing the distribution of Cuccodorodes species." figureDoi="http://doi.org/10.5281/zenodo.4504770" httpUri="https://zenodo.org/record/4504770/files/figure.png" pageId="5" pageNumber="326">Fig. 7</figureCitation>
). .............. 
<emphasis id="75B22421FFB4FF82A502FCFDE2A48FD1" bold="true" box="[549,760,1736,1760]" pageId="5" pageNumber="326">
<taxonomicName id="80C683B0FFB4FF82A502FCFDE2C78FD1" ID-CoL="328TC" authority="Yin, 2018" authorityName="Yin" authorityYear="2018" box="[549,667,1736,1760]" class="Insecta" family="Staphylinidae" genus="Cuccodorodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="weiperti" status="sp. nov.">
<emphasis id="75B22421FFB4FF82A502FCFDE26B8FD1" bold="true" box="[549,567,1736,1760]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="75B22421FFB4FF82A563FCFDE2C78FD1" bold="true" box="[580,667,1736,1760]" italics="true" pageId="5" pageNumber="326">weiperti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6E81995AFFB4FF82A585FCFCE2A48FD1" box="[674,760,1737,1760]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>