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<document id="B4D905F1EF125BBB9EE67071AFC508AB" ID-CLB-Dataset="2598" ID-DOI="10.5852/ejt.2022.788.1625" ID-GBIF-Dataset="d6e6119f-4100-4cfc-9cc3-0d6951ee2c28" ID-ISSN="2118-9773" ID-Zenodo-Dep="5846636" ID-ZooBank="46C3CFCA-0CED-4432-AFD8-F4CFC1E0E1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642092502222" checkinUser="felipe" docAuthor="Castillo, Stephanie, Rédei, Dávid &amp; Weirauch, Christiane" docDate="2022" docId="015D8E70FFBFFFB1FDC30524FADBBFE7" docLanguage="en" docName="ejt-788_1-95.pdf" docOrigin="European Journal of Taxonomy 788 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.788.1625" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Voconia ifana Castillo &amp; Rédei &amp; Weirauch 2022, comb. nov." docType="treatment" docVersion="10" lastPageNumber="54" masterDocId="FD64F608FF8BFF84FFE7031FFFACBA3D" masterDocTitle="Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs" masterLastPageNumber="95" masterPageNumber="1" pageNumber="53" updateTime="1699302197121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3BEB3706428231CC0C9088ACCAF34C64">Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs</mods:title>
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<mods:affiliation id="8720EC6F1C8AE23F921C5FC5D073A97F">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<mods:namePart id="FD0FEC3B16B0683352B311F051642AEA">Rédei, Dávid</mods:namePart>
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<mods:affiliation id="9A71ED1F6F72289A0168E98E1A79398F">National Chung Hsing University, 145 Xingda Rd., 40227 Taichung, Taiwan.</mods:affiliation>
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<mods:affiliation id="0FDDB5F9027B71787DF6173445FC53DD">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<mods:date id="FA0A25A2C4D4D7204E1B06B61CC710BF">2022</mods:date>
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<subSubSection id="C1EE6CEDFFBFFFB0FDC30524FBBCBC6B" box="[548,1040,1595,1622]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="894B3F66FFBFFFB0FDC30524FBBCBC6B" blockId="52.[548,1040,1595,1657]" box="[548,1040,1595,1622]" pageId="52" pageNumber="53">
<heading id="D203880AFFBFFFB0FDC30524FBBCBC6B" box="[548,1040,1595,1622]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="53" reason="2">
<taxonomicName id="4EF444E5FFBFFFB0FDC30524FC2BBC6B" authority="(Villiers, 1963)" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" baseAuthorityName="Villiers" baseAuthorityYear="1963" box="[548,903,1595,1622]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="ifana" status="comb. nov.">
<emphasis id="BB80E374FFBFFFB0FDC30524FD61BC68" bold="true" box="[548,717,1595,1621]" italics="true" pageId="52" pageNumber="53">Voconia ifana</emphasis>
(Villiers, 1963)
</taxonomicName>
<taxonomicNameLabel id="A0B35E0FFFBFFFB0FC690524FBBCBC6B" box="[910,1040,1595,1622]" pageId="52" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
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<figureCitation id="11CF23E3FFBFFFB0FD170540FCEFBC44" box="[752,835,1631,1657]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="52" pageNumber="53">Fig. 17</figureCitation>
</paragraph>
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<paragraph id="894B3F66FFBFFFB0FF5A05BFFD52BC87" blockId="52.[189,766,1696,1722]" box="[189,766,1696,1722]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="A9E41848FFBFFFB0FF5A05BFFD52BC87" box="[189,766,1696,1722]" pageId="52" pageNumber="53">
<treatmentCitation id="08551977FFBFFFB0FF5A05BFFD0DBC87" author="Villiers A." box="[189,673,1696,1722]" page="989" pageId="52" pageNumber="53" year="1963">
<taxonomicName id="4EF444E5FFBFFFB0FF5A05BFFD0DBC87" ID-CoL="4NSZB" authority="Villiers, 1963 a: 989" authorityName="Villiers" authorityPageNumber="989" authorityYear="1963" box="[189,673,1696,1722]" class="Insecta" family="Reduviidae" genus="Pseudocethera" kingdom="Animalia" order="Hemiptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="ifana">
<emphasis id="BB80E374FFBFFFB0FF5A05BFFE03BC87" box="[189,431,1696,1722]" italics="true" pageId="52" pageNumber="53">Pseudocethera ifana</emphasis>
<bibRefCitation id="ED654297FFBFFFB0FE5105BFFD0DBC87" author="Villiers A." box="[438,673,1696,1722]" pageId="52" pageNumber="53" pagination="969 - 994" refId="ref52220" refString="Villiers A. 1963 a. Contribution a l'etude de la faune de la basse Casamance. Hemipteres Reduviides et Henicocephalides. Bulletin de l'Instiut Francais d'Afrique Noire, Ser. A 25 (3): 969 - 994." type="journal article" year="1963">Villiers, 1963a: 989</bibRefCitation>
</taxonomicName>
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, fig. 11.
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</paragraph>
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<subSubSection id="C1EE6CEDFFBFFFB0FF5A05FEFEA5BDE7" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="894B3F66FFBFFFB0FF5A05FEFE36BCC6" blockId="52.[189,410,1761,1787]" box="[189,410,1761,1787]" pageId="52" pageNumber="53">
<heading id="D203880AFFBFFFB0FF5A05FEFE36BCC6" bold="true" box="[189,410,1761,1787]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="BB80E374FFBFFFB0FF5A05FEFE36BCC6" bold="true" box="[189,410,1761,1787]" pageId="52" pageNumber="53">Revised diagnosis</emphasis>
</heading>
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<paragraph id="894B3F66FFBFFFB0FF5A040FFEA5BDE7" blockId="52.[189,1399,1808,2010]" pageId="52" pageNumber="53">
Distinguished from other species of
<taxonomicName id="4EF444E5FFBFFFB0FD92040EFD78BD17" authorityName="Stal" authorityYear="1866" box="[629,724,1809,1834]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="BB80E374FFBFFFB0FD92040EFD78BD17" box="[629,724,1809,1834]" italics="true" pageId="52" pageNumber="53">Voconia</emphasis>
</taxonomicName>
by the following combination of characters: short body length (about
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); coarsely granulose head and pronotum; maxillary plates are not adjacent to and far surpass clypeal apex; maxillary plates in lateral view are wide, about twice the width of scape; antennifer with long lateral spine; eyes do not reach ventral head margin in lateral view. It is recognized from micropterous individuals of
<taxonomicName id="4EF444E5FFBFFFB0FDB40481FD52BD8A" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" baseAuthorityName="Villiers" baseAuthorityYear="1964" box="[595,766,1949,1975]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="BB80E374FFBFFFB0FDB40481FD52BD8A" box="[595,766,1949,1975]" italics="true" pageId="52" pageNumber="53">V. schoutedeni</emphasis>
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and
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<emphasis id="BB80E374FFBFFFB0FCDA0481FC18BD8A" box="[829,948,1949,1975]" italics="true" pageId="52" pageNumber="53">V. monodi</emphasis>
</taxonomicName>
by the toothed apex of the maxillary plates.
</paragraph>
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<emphasis id="BB80E374FFBEFFB1FF5A0216FEC0BB1F" bold="true" box="[189,364,264,291]" pageId="53" pageNumber="54">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF0D0228FEB3BB86" blockId="53.[189,1399,311,443]" pageId="53" pageNumber="54">
<materialsCitation id="399C353BFFBEFFB1FF0D0228FEB3BB86" ID-GBIF-Occurrence="3436573355" collectingDate="1961-11-21" collectionCode="MNHN" country="Senegal" location="Tobor Forest" municipality="Foret classee de Tobor" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="BB80E374FFBEFFB1FF0D0228FEF0BB6C" bold="true" box="[234,348,311,337]" pageId="53" pageNumber="54">
<typeStatus id="564F81C4FFBEFFB1FF0D0228FEF0BB6C" box="[234,348,311,337]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F1E37FF6FFBEFFB1FF5A0245FEEEBB49" box="[189,322,346,372]" name="Senegal" pageId="53" pageNumber="54">SENEGAL</collectingCountry>
<specimenCount id="9FF2F4EFFFBEFFB1FE830244FED1BB48" box="[356,381,347,373]" pageId="53" pageNumber="54" type="male"></specimenCount>
; “
<collectingMunicipality id="692FA51CFFBEFFB1FE700245FD1FBB49" box="[407,691,346,372]" pageId="53" pageNumber="54">Forêt classée de Tobor</collectingMunicipality>
” [
<location id="8C2B69BDFFBEFFB1FD280245FCD8BB48" LSID="urn:lsid:plazi:treatment:015D8E70FFBFFFB1FDC30524FADBBFE7:8C2B69BDFFBEFFB1FD280245FCD8BB48" box="[719,884,346,373]" country="Senegal" municipality="Foret classee de Tobor" name="Tobor Forest" pageId="53" pageNumber="54">Tobor Forest</location>
];
<date id="FD4A19A6FFBEFFB1FC6C0245FB9ABB49" box="[907,1078,346,373]" pageId="53" pageNumber="54" value="1961-11-21">
<collectingDate id="ED0EE04EFFBEFFB1FC6C0245FB9ABB49" box="[907,1078,346,373]" pageId="53" pageNumber="54" value="1961-11-21">21 Nov. 1961</collectingDate>
</date>
; “Mission IFAN” [Institut Fondamental dAfrique Noire exped.]; “sous un morceau de bois mort” [under a piece of dead wood];
<collectionCode id="EFE5A7A3FFBEFFB1FF5902BEFEB0BB86" box="[190,284,417,443]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="53" pageNumber="54">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFBEFFB1FF5A02FDFE95BE50" pageId="53" pageNumber="54" type="description">
<paragraph id="894B3F66FFBEFFB1FF5A02FDFEE2BBC1" blockId="53.[189,334,482,508]" box="[189,334,482,508]" pageId="53" pageNumber="54">
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<emphasis id="BB80E374FFBEFFB1FF5A02FDFEE2BBC1" bold="true" box="[189,334,482,508]" pageId="53" pageNumber="54">Description</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF0D010EFE86B816" blockId="53.[189,706,529,591]" box="[234,298,529,555]" pageId="53" pageNumber="54">
<emphasis id="BB80E374FFBEFFB1FF0D010EFE86B816" bold="true" box="[234,298,529,555]" pageId="53" pageNumber="54">Male</emphasis>
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A012BFD6EB873" blockId="53.[189,706,529,591]" box="[189,706,564,591]" pageId="53" pageNumber="54">
BODY LENGTH. About
<quantity id="4E0C9283FFBEFFB1FE51012BFDBFB873" box="[438,531,564,591]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="53" pageNumber="54" unit="mm" value="6.5">6.5 mm</quantity>
; micropterous.
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A016AFC00B8AD" blockId="53.[189,940,629,656]" box="[189,940,629,656]" pageId="53" pageNumber="54">
COLORATION. As described for
<taxonomicName id="4EF444E5FFBEFFB1FDF90169FD3FB8B2" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" baseAuthorityName="Villiers" baseAuthorityYear="1963" box="[542,659,629,655]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="BB80E374FFBEFFB1FDF90169FD3FB8B2" box="[542,659,629,655]" italics="true" pageId="53" pageNumber="54">V. monodi</emphasis>
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, except legs testaceous.
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A01A8FC00B8C9" blockId="53.[187,1398,695,756]" pageId="53" pageNumber="54">INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with setigerous tubercles; interocular region with two pairs of large setigerous tubercles paramedially.</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A0004FDD0BE34" blockId="53.[189,1401,795,1033]" pageId="53" pageNumber="54">STRUCTURE. Head: scape not reaching head apex; antennifer with long lateral spine; postocular region in dorsal view as long as eye, lateral margins subrectangular; maxillary plates in lateral view about twice the width of scape and apex toothed; eye not reaching ventral head margin in lateral view; ventrolateral swelling of buccula with lateral protrusion that surpasses buccular margin; labial segment I in lateral view straight, surpassing posteroventral eye margin. Thorax: scutellar spine reduced, subhorizontal. Abdomen: anterior margin of terga carinulate; tergum II with paired prominent longitudinal carinae reaching posterior margin of segment.</paragraph>
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<emphasis id="BB80E374FFBEFFB1FF0D072FFEE9BE77" bold="true" box="[234,325,1072,1098]" pageId="53" pageNumber="54">Female</emphasis>
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A074CFE95BE50" blockId="53.[189,325,1072,1133]" box="[189,313,1107,1133]" pageId="53" pageNumber="54">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFBEFFB1FF5A078BFD76BEE0" pageId="53" pageNumber="54" type="distribution">
<paragraph id="894B3F66FFBEFFB1FF5A078BFE68BE92" blockId="53.[189,452,1172,1199]" box="[189,452,1172,1199]" pageId="53" pageNumber="54">
<emphasis id="BB80E374FFBEFFB1FF5A078BFEFABE93" bold="true" box="[189,342,1172,1198]" pageId="53" pageNumber="54">Distribution</emphasis>
(
<figureCitation id="11CF23E3FFBEFFB1FE81078BFE17BE92" box="[358,443,1172,1199]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="53" pageNumber="54">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A07DCFD76BEE0" blockId="53.[189,730,1219,1245]" box="[189,730,1219,1245]" pageId="53" pageNumber="54">This species is known from Northwest Africa.</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFBEFFB1FF5A061AFADBBFE7" pageId="53" pageNumber="54" type="discussion">
<paragraph id="894B3F66FFBEFFB1FF5A061AFE83BF22" blockId="53.[189,303,1285,1311]" box="[189,303,1285,1311]" pageId="53" pageNumber="54">
<heading id="D203880AFFBEFFB1FF5A061AFE83BF22" bold="true" box="[189,303,1285,1311]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="BB80E374FFBEFFB1FF5A061AFE83BF22" bold="true" box="[189,303,1285,1311]" pageId="53" pageNumber="54">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFBEFFB1FF5A062CFADBBFE7" blockId="53.[189,1400,1331,1498]" pageId="53" pageNumber="54">
The
<typeStatus id="564F81C4FFBEFFB1FF13062CFEF7BF70" box="[244,347,1331,1357]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
was on loan and not available for study at the time we requested it. Consequently, this description is based on the interpretation of the short differential diagnosis and illustration of the head in lateral view provided by
<bibRefCitation id="ED654297FFBEFFB1FDE30666FD6CBFA9" author="Villiers A." box="[516,704,1401,1428]" pageId="53" pageNumber="54" pagination="969 - 994" refId="ref52220" refString="Villiers A. 1963 a. Contribution a l'etude de la faune de la basse Casamance. Hemipteres Reduviides et Henicocephalides. Bulletin de l'Instiut Francais d'Afrique Noire, Ser. A 25 (3): 969 - 994." type="journal article" year="1963">Villiers (1963a)</bibRefCitation>
. Due to the limited data that we were able to obtain, it was excluded from cladistic analyses, but we are confident that it likely forms a clade with
<taxonomicName id="4EF444E5FFBEFFB1FB2B0682FADABF8A" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" baseAuthorityName="Villiers" baseAuthorityYear="1964" box="[1228,1398,1437,1463]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="BB80E374FFBEFFB1FB2B0682FADABF8A" box="[1228,1398,1437,1463]" italics="true" pageId="53" pageNumber="54">V. schoutedeni</emphasis>
</taxonomicName>
and
<taxonomicName id="4EF444E5FFBEFFB1FF0A06DEFEF5BFE7" authority="Castillo &amp; Rédei &amp; Weirauch, 2022" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" box="[237,345,1472,1498]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tridens" status="sp. nov.">
<emphasis id="BB80E374FFBEFFB1FF0A06DEFEF5BFE7" box="[237,345,1472,1498]" italics="true" pageId="53" pageNumber="54">V. tridens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B35E0FFFBEFFB1FE8706DFFE15BFE7" box="[352,441,1472,1498]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
because of the unique head morphology, microptery, and Afrotropical distribution.
</paragraph>
</subSubSection>
</treatment>
</document>