273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document id="FBDDA6D57EAF97936B4CB9D9AC6C107F" ID-CLB-Dataset="2598" ID-DOI="10.5852/ejt.2022.788.1625" ID-GBIF-Dataset="d6e6119f-4100-4cfc-9cc3-0d6951ee2c28" ID-ISSN="2118-9773" ID-Zenodo-Dep="5846636" ID-ZooBank="46C3CFCA-0CED-4432-AFD8-F4CFC1E0E1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642092502222" checkinUser="felipe" docAuthor="Castillo, Stephanie, Rédei, Dávid & Weirauch, Christiane" docDate="2022" docId="015D8E70FFB3FFBFFD9F05B5FD09B8EF" docLanguage="en" docName="ejt-788_1-95.pdf" docOrigin="European Journal of Taxonomy 788 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.788.1625" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Voconia lasiosoma Castillo & Rédei & Weirauch 2022, sp. nov." docType="treatment" docUuid="C5495703-AEE2-49FC-8B08-099ED0F398D3" docUuidSource="ZooBank" docVersion="10" lastPageNumber="60" masterDocId="FD64F608FF8BFF84FFE7031FFFACBA3D" masterDocTitle="Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs" masterLastPageNumber="95" masterPageNumber="1" pageNumber="57" updateTime="1699302197121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D9475116F0BD48E4C8CF757E67D5B9D1">Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs</mods:title>
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<mods:namePart id="584A8596BB966245C385E8E2D59C3077">Castillo, Stephanie</mods:namePart>
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<mods:affiliation id="582BD56211BA74144A4FE1BA15E87FE0">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<mods:nameIdentifier id="8AB986B8E0E33D89449D31F2FFDA7908" type="email">scast032@ucr.edu</mods:nameIdentifier>
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<mods:namePart id="16BF4D73D9772921CD1D0FC08F061AAE">Rédei, Dávid</mods:namePart>
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<mods:affiliation id="F4D99895738694D2A02EFE0E909ADA4D">National Chung Hsing University, 145 Xingda Rd., 40227 Taichung, Taiwan.</mods:affiliation>
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<mods:nameIdentifier id="E476BA6133EA11B08E47811341C8D5C4" type="email">david.redei@gmail.com</mods:nameIdentifier>
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<mods:namePart id="70DABE3C669D27B5E389F7FED8554AA3">Weirauch, Christiane</mods:namePart>
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<mods:affiliation id="65885A6794862DD40042085864CEB841">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
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<mods:nameIdentifier id="98E2F38035C28E16BC8CB9AE6EC2A291" type="email">christiane.weirauch@ucr.edu</mods:nameIdentifier>
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<mods:title id="4D86945C5B6DF844EEC0443E6B15F598">European Journal of Taxonomy</mods:title>
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<mods:part id="B5445F688AAF3515A92B53DF72DEDF1C">
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<mods:date id="8C2458B9B6546F10063B83055FE440A3">2022</mods:date>
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<mods:number id="F9E012848D0B2386D246C6538CE0E6EC">2022-01-11</mods:number>
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<treatment id="015D8E70FFB3FFBFFD9F05B5FD09B8EF" ID-DOI="http://doi.org/10.5281/zenodo.5849721" ID-GBIF-Taxon="192123849" ID-Zenodo-Dep="5849721" LSID="urn:lsid:zoobank.org:act:C5495703-AEE2-49FC-8B08-099ED0F398D3" httpUri="http://treatment.plazi.org/id/015D8E70FFB3FFBFFD9F05B5FD09B8EF" lastPageId="59" lastPageNumber="60" pageId="56" pageNumber="57">
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<subSubSection id="C1EE6CEDFFB3FFBCFD9F05B5FC10BCF8" box="[632,956,1705,1733]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="894B3F66FFB3FFBCFD9F05B5FC10BCF8" blockId="56.[369,1218,1705,1803]" box="[632,956,1705,1733]" pageId="56" pageNumber="57">
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<heading id="D203880AFFB3FFBCFD9F05B5FC10BCF8" box="[632,956,1705,1733]" centered="true" fontSize="11" level="2" pageId="56" pageNumber="57" reason="2">
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<taxonomicName id="4EF444E5FFB3FFBCFD9F05B5FCF6BCF9" authority="Castillo & Rédei & Weirauch, 2022" authorityName="Castillo & Rédei & Weirauch" authorityYear="2022" box="[632,858,1705,1732]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="lasiosoma" status="sp. nov.">
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<emphasis id="BB80E374FFB3FFBCFD9F05B5FCF6BCF9" bold="true" box="[632,858,1705,1732]" italics="true" pageId="56" pageNumber="57">Voconia lasiosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0B35E0FFFB3FFBCFC8605B4FC10BCF8" box="[865,956,1707,1733]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C1EE6CEDFFB3FFBCFE9605D2FC00BD36" pageId="56" pageNumber="57" type="description">
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<uri id="FD653364FFB3FFBCFE9605D2FB6EBCD5" box="[369,1218,1741,1768]" pageId="56" pageNumber="57">
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urn:lsid:zoobank.org:act:
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<uuid id="FD5205B3FFB3FFBCFD7705D2FB6EBCD5" box="[656,1218,1741,1768]" pageId="56" pageNumber="57">C5495703-AEE2-49FC-8B08-099ED0F398D3</uuid>
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</uri>
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</paragraph>
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<paragraph id="894B3F66FFB3FFBCFD6005EFFC00BD36" blockId="56.[369,1218,1705,1803]" box="[647,940,1776,1803]" pageId="56" pageNumber="57">
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<figureCitation id="11CF23E3FFB3FFBCFD6005EFFD5CBD36" box="[647,752,1776,1803]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[190,233,1950,1976]" captionStartId-1="9.[189,232,1883,1909]" captionTargetBox-0="[204,1371,808,1898]" captionTargetBox-1="[199,1395,651,1845]" captionTargetId-0="figure-168@8.[189,1398,793,1908]" captionTargetId-1="figure-137@9.[189,1398,644,1845]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 1. Strict consensus tree resulting from equal weighting analysis. Jackknife values over 51 are reported for tree one of eight." captionText-1="Fig. 2. Tree resulting from implied weighting analysis using K = 12 with characters unambiguously optimized. Symmetric resampling values over 51 are represented for trees obtained with K = 3, K = 6, K = 9, and K = 12 analyses. Characters and character states are described in pp. 11–22. Specimens not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5846638" figureDoi-1="http://doi.org/10.5281/zenodo.5846640" httpUri-0="https://zenodo.org/record/5846638/files/figure.png" httpUri-1="https://zenodo.org/record/5846640/files/figure.png" pageId="56" pageNumber="57">Figs 1–2</figureCitation>
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,
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<figureCitation id="11CF23E3FFB3FFBCFD1B05EEFCA1BD36" box="[764,781,1777,1803]" captionStart="Fig" captionStartId="32.[189,232,1960,1986]" captionTargetBox="[189,1398,643,1917]" captionTargetId="figure-72@32.[189,1398,643,1917]" captionTargetPageId="32" captionText="Fig. 7. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846650" httpUri="https://zenodo.org/record/5846650/files/figure.png" pageId="56" pageNumber="57">7</figureCitation>
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||
,
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<figureCitation id="11CF23E3FFB3FFBCFCFE05EEFC86BD36" box="[793,810,1777,1803]" captionStart="Fig" captionStartId="37.[189,232,1982,2008]" captionTargetBox="[189,1398,910,1943]" captionTargetId="figure-247@37.[189,1398,910,1943]" captionTargetPageId="37" captionText="Fig. 9. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846654" httpUri="https://zenodo.org/record/5846654/files/figure.png" pageId="56" pageNumber="57">9</figureCitation>
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,
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<figureCitation id="11CF23E3FFB3FFBCFCD005EEFCFBBD36" box="[823,855,1777,1803]" captionStart="Fig" captionStartId="57.[189,232,1985,2011]" captionTargetBox="[189,1397,687,1944]" captionTargetId="figure-102@57.[189,1399,686,1944]" captionTargetPageId="57" captionText="Fig. 12. Pygophore of species of Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5846662" httpUri="https://zenodo.org/record/5846662/files/figure.png" pageId="56" pageNumber="57">12</figureCitation>
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,
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<figureCitation id="11CF23E3FFB3FFBCFC8405EEFC2FBD36" box="[867,899,1777,1803]" captionStart="Fig" captionStartId="75.[188,231,1989,2015]" captionTargetBox="[189,1397,760,1946]" captionTargetId="figure-180@75.[189,1397,760,1946]" captionTargetPageId="75" captionText="Fig. 14. Aedeagus of species of Gerbelius Distant, 1903 and Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5846668" httpUri="https://zenodo.org/record/5846668/files/figure.png" pageId="56" pageNumber="57">14</figureCitation>
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,
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<figureCitation id="11CF23E3FFB3FFBCFC6805EEFC00BD36" box="[911,940,1777,1803]" captionStart="Fig" captionStartId="88.[188,231,1983,2009]" captionTargetBox="[189,1398,1222,1944]" captionTargetId="figure-267@88.[189,1398,1222,1944]" captionTargetPageId="88" captionText="Fig. 20. The distribution of species of Gerbelius Distant, 1903 and Voconia Stål, 1866 in Southeast Asia." figureDoi="http://doi.org/10.5281/zenodo.5846682" httpUri="https://zenodo.org/record/5846682/files/figure.png" pageId="56" pageNumber="57">20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1EE6CEDFFB3FFBDFF5A042DFCE0BBB0" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph id="894B3F66FFB3FFBCFF5A042DFE9ABD71" blockId="56.[189,310,1842,1868]" box="[189,310,1842,1868]" pageId="56" pageNumber="57">
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<heading id="D203880AFFB3FFBCFF5A042DFE9ABD71" bold="true" box="[189,310,1842,1868]" fontSize="11" level="3" pageId="56" pageNumber="57" reason="3">
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<emphasis id="BB80E374FFB3FFBCFF5A042DFE9ABD71" bold="true" box="[189,310,1842,1868]" pageId="56" pageNumber="57">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="894B3F66FFB3FFBDFF5A047FFCE0BBB0" blockId="56.[186,1400,1888,2021]" lastBlockId="57.[189,1400,265,397]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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Recognized from other
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<taxonomicName id="4EF444E5FFB3FFBCFE2E047EFD84BD47" authorityName="Stal" authorityYear="1866" box="[457,552,1889,1914]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="BB80E374FFB3FFBCFE2E047EFD84BD47" box="[457,552,1889,1914]" italics="true" pageId="56" pageNumber="57">Voconia</emphasis>
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</taxonomicName>
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by the densely long setation on the pronotum, corium, legs, and abdomen. This species most closely resembles two other Southeast Asian species,
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<taxonomicName id="4EF444E5FFB3FFBCFBEA049BFB3EBDA0" authority="Castillo & Rédei & Weirauch, 2022" authorityName="Castillo & Rédei & Weirauch" authorityYear="2022" box="[1037,1170,1923,1949]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="isosceles" status="sp. nov.">
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<emphasis id="BB80E374FFB3FFBCFBEA049BFB3EBDA0" box="[1037,1170,1923,1949]" italics="true" pageId="56" pageNumber="57">V. isosceles</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0B35E0FFFB3FFBCFB7E049BFB58BDA3" box="[1177,1268,1924,1950]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4EF444E5FFB3FFBCFAC9049BFADBBDA0" authority="Castillo & Rédei & Weirauch, 2022" authorityName="Castillo & Rédei & Weirauch" authorityYear="2022" box="[1326,1399,1923,1949]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="loki" status="sp. nov.">
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<emphasis id="BB80E374FFB3FFBCFAC9049BFADBBDA0" box="[1326,1399,1923,1949]" italics="true" pageId="56" pageNumber="57">V. loki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0B35E0FFFB3FFBCFF5A04B8FEBABDFC" box="[189,278,1959,1985]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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due to the finely granulose head and pronotum and dark pronotum with contrasting pale posterior margin. While
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<taxonomicName id="4EF444E5FFB3FFBCFE9504D4FE11BDD9" authority="Castillo & Rédei & Weirauch, 2022" authorityName="Castillo & Rédei & Weirauch" authorityYear="2022" box="[370,445,1994,2020]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="loki" status="sp. nov.">
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<emphasis id="BB80E374FFB3FFBCFE9504D4FE11BDD9" box="[370,445,1994,2020]" italics="true" pageId="56" pageNumber="57">V. loki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0B35E0FFFB3FFBCFE2204D4FD8EBDD8" box="[453,546,1995,2021]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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is also hairy,
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<taxonomicName id="4EF444E5FFB3FFBCFD2904D4FCC8BDD9" authority="Castillo & Rédei & Weirauch, 2022" authorityName="Castillo & Rédei & Weirauch" authorityYear="2022" box="[718,868,1994,2020]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="lasiosoma" status="sp. nov.">
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<emphasis id="BB80E374FFB3FFBCFD2904D4FCC8BDD9" box="[718,868,1994,2020]" italics="true" pageId="56" pageNumber="57">V. lasiosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0B35E0FFFB3FFBCFC8A04D4FC66BDD8" box="[877,970,1995,2021]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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is larger (about
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<quantity id="4E0C9283FFB3FFBCFB7604D5FB43BDD8" box="[1169,1263,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="56" pageNumber="57" unit="mm" value="9.4">9.4 mm</quantity>
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) than these two species; legs, antennae, and corium are dark yellowish-brown rather than yellow; dark portions of corium and clavus are black rather than reddish-brown; Cu-An
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<subScript id="15703D23FFB2FFBDFC4C0220FC18BB70" attach="left" box="[939,948,319,333]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
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cell stout (less than half the length of M-Cu cell); and proximal half of corium not entirely yellow (abducted corium with anteroproximal yellowish-brown stripe and posteromedial yellow spot).
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</paragraph>
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</subSubSection>
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<subSubSection id="C1EE6CEDFFB2FFBDFF5A02ABFEF1B87E" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="894B3F66FFB2FFBDFF5A02ABFEE8BBF3" blockId="57.[189,324,436,462]" box="[189,324,436,462]" pageId="57" pageNumber="58">
|
||
<heading id="D203880AFFB2FFBDFF5A02ABFEE8BBF3" bold="true" box="[189,324,436,462]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
|
||
<emphasis id="BB80E374FFB2FFBDFF5A02ABFEE8BBF3" bold="true" box="[189,324,436,462]" pageId="57" pageNumber="58">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB2FFBDFF5A02FDFEF1B87E" blockId="57.[189,1399,482,579]" pageId="57" pageNumber="58">
|
||
Derived from the Latinized Greek prefix ‘
|
||
<emphasis id="BB80E374FFB2FFBDFD5202FDFD54BBC1" box="[693,760,482,508]" italics="true" pageId="57" pageNumber="58">lasio-</emphasis>
|
||
’ meaning ‘hairy’ combined with the Latinized Greek noun ‘-
|
||
<emphasis id="BB80E374FFB2FFBDFEF40118FEFDB81D" box="[275,337,519,544]" italics="true" pageId="57" pageNumber="58">soma</emphasis>
|
||
’ meaning ‘body’. Refers to the long macrosetae covering the body and hemelytra. A noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DD8B6FEEFFB2FFBDFF5A04DEFB8BBDE1" ID-DOI="http://doi.org/10.5281/zenodo.5846662" ID-Zenodo-Dep="5846662" box="[189,1063,1985,2012]" httpUri="https://zenodo.org/record/5846662/files/figure.png" pageId="57" pageNumber="58" startId="57.[189,232,1985,2011]" targetBox="[189,1397,687,1944]" targetPageId="57">
|
||
<paragraph id="894B3F66FFB2FFBDFF5A04DEFB8BBDE1" blockId="57.[189,1063,1985,2012]" box="[189,1063,1985,2012]" pageId="57" pageNumber="58">
|
||
<emphasis id="BB80E374FFB2FFBDFF5A04DEFEB6BDE6" bold="true" box="[189,282,1985,2011]" pageId="57" pageNumber="58">Fig. 12.</emphasis>
|
||
Pygophore of species of
|
||
<taxonomicName id="4EF444E5FFB2FFBDFDA304DDFC88BDE6" authority="Stal, 1866" authorityName="Stal" authorityYear="1866" box="[580,804,1985,2012]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BB80E374FFB2FFBDFDA304DDFD0FBDE6" box="[580,675,1986,2011]" italics="true" pageId="57" pageNumber="58">Voconia</emphasis>
|
||
Stål, 1866
|
||
</taxonomicName>
|
||
. Scale bars = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C1EE6CEDFFB1FFBEFF5A0216FE2FBBE3" pageId="58" pageNumber="59" type="materials_examined">
|
||
<paragraph id="894B3F66FFB1FFBEFF5A0216FEC0BB1F" blockId="58.[189,364,264,291]" box="[189,364,264,291]" pageId="58" pageNumber="59">
|
||
<heading id="D203880AFFB1FFBEFF5A0216FEC0BB1F" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="BB80E374FFB1FFBEFF5A0216FEC0BB1F" bold="true" box="[189,364,264,291]" pageId="58" pageNumber="59">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBEFF0D0228FE2FBBE3" blockId="58.[189,1401,311,478]" pageId="58" pageNumber="59">
|
||
<materialsCitation id="399C353BFFB1FFBEFF0D0228FE2FBBE3" ID-GBIF-Occurrence="3436573373" collectingDate="2017-10-09" collectingDateMax="2017-10-10" collectingDateMin="2017-10-09" collectionCode="ZRC" collectorName="W. Hwang" country="Malaysia" elevation="3" latitude="1.55" location="Lanjak Entimau Wildlife Sanctuary" longLatPrecision="785" longitude="110.36" municipality="Lanjak Entimau Wildlife Sanctuary" pageId="58" pageNumber="59" specimenCode="UCR_ENT 00127826" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
|
||
<emphasis id="BB80E374FFB1FFBEFF0D0228FEF0BB6C" bold="true" box="[234,348,311,337]" pageId="58" pageNumber="59">
|
||
<typeStatus id="564F81C4FFB1FFBEFF0D0228FEF0BB6C" box="[234,348,311,337]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F1E37FF6FFB1FFBEFF5A0245FEF8BB49" box="[189,340,346,372]" name="Malaysia" pageId="58" pageNumber="59">MALAYSIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9FF2F4EFFFB1FFBEFE9E0244FE23BB48" box="[377,399,347,373]" pageId="58" pageNumber="59" type="male">♂</specimenCount>
|
||
(dissected pygophore and aedeagus in vial);
|
||
<collectingRegion id="4B30F184FFB1FFBEFC2B0245FB98BB49" box="[972,1076,346,372]" country="Malaysia" name="Sarawak" pageId="58" pageNumber="59">Sarawak</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="692FA51CFFB1FFBEFBA10245FE9ABBA5" country="Malaysia" latitude="1.55" longLatPrecision="785" longitude="110.36" name="Lanjak Entimau Wildlife Sanctuary" pageId="58" pageNumber="59" stateProvince="Sarawak">Lanjak Entimau Wildlife Sanctuary</collectingMunicipality>
|
||
; [
|
||
<geoCoordinate id="ECC059A1FFB1FFBEFEAC0262FE01BBA5" box="[331,429,381,408]" degrees="1.55" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="555" value="1.55">
|
||
1.55
|
||
<emphasis id="BB80E374FFB1FFBEFE660262FE21BBAA" bold="true" box="[385,397,381,407]" pageId="58" pageNumber="59">°</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ECC059A1FFB1FFBEFE5C0262FD99BBA5" box="[443,565,381,408]" degrees="110.36" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="555" value="110.36">
|
||
110.36
|
||
<emphasis id="BB80E374FFB1FFBEFDEC0262FDBBBBAA" bold="true" box="[523,535,381,407]" pageId="58" pageNumber="59">°</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
]; [elev.
|
||
<quantity id="4E0C9283FFB1FFBEFD720261FD65BBA5" box="[661,713,381,408]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="58" pageNumber="59" unit="m" value="3.0">
|
||
<elevation id="02D9D855FFB1FFBEFD720261FD65BBA5" box="[661,713,381,408]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="58" pageNumber="59" unit="m" value="3.0">3 m</elevation>
|
||
</quantity>
|
||
];
|
||
<date id="FD4A19A6FFB1FFBEFD380261FC30BBA5" box="[735,924,382,408]" pageId="58" pageNumber="59" value="2017-10-09" valueMax="2017-10-10" valueMin="2017-10-09">
|
||
<collectingDate id="ED0EE04EFFB1FFBEFD380261FC30BBA5" box="[735,924,382,408]" pageId="58" pageNumber="59" value="2017-10-09" valueMax="2017-10-10" valueMin="2017-10-09">9–10 Oct. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="24015AB0FFB1FFBEFC4E0261FB8BBBA5" box="[937,1063,382,408]" pageId="58" pageNumber="59">W. Hwang</collectorName>
|
||
<emphasis id="BB80E374FFB1FFBEFBD50261FBC9BBAA" box="[1074,1125,381,407]" italics="true" pageId="58" pageNumber="59">et al</emphasis>
|
||
. leg.; collecting event: SW17_L_27; MV light trap in dipterocarp forest; DNA voucher R_CW 5472; USI:
|
||
<specimenCode id="D952971DFFB1FFBEFB1202BEFE9ABBE3" pageId="58" pageNumber="59">UCR_ENT 00127826</specimenCode>
|
||
;
|
||
<collectionCode id="EFE5A7A3FFB1FFBEFEA502DBFED3BBE3" box="[322,383,452,478]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="58" pageNumber="59">ZRC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1EE6CEDFFB1FFBFFF5A011AFE95BB96" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
|
||
<paragraph id="894B3F66FFB1FFBEFF5A011AFEE2B822" blockId="58.[189,334,517,543]" box="[189,334,517,543]" pageId="58" pageNumber="59">
|
||
<heading id="D203880AFFB1FFBEFF5A011AFEE2B822" bold="true" box="[189,334,517,543]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="BB80E374FFB1FFBEFF5A011AFEE2B822" bold="true" box="[189,334,517,543]" pageId="58" pageNumber="59">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBEFF0D012BFE06B873" blockId="58.[189,711,564,626]" box="[234,426,564,590]" pageId="58" pageNumber="59">
|
||
<emphasis id="BB80E374FFB1FFBEFF0D012BFE86B873" bold="true" box="[234,298,564,590]" pageId="58" pageNumber="59">Male</emphasis>
|
||
(
|
||
<figureCitation id="11CF23E3FFB1FFBEFEDE012BFE2AB873" box="[313,390,564,590]" captionStart="Fig" captionStartId="32.[189,232,1960,1986]" captionTargetBox="[189,1398,643,1917]" captionTargetId="figure-72@32.[189,1398,643,1917]" captionTargetPageId="32" captionText="Fig. 7. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846650" httpUri="https://zenodo.org/record/5846650/files/figure.png" pageId="58" pageNumber="59">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="11CF23E3FFB1FFBEFE75012BFE0DB873" box="[402,417,564,590]" captionStart="Fig" captionStartId="37.[189,232,1982,2008]" captionTargetBox="[189,1398,910,1943]" captionTargetId="figure-247@37.[189,1398,910,1943]" captionTargetPageId="37" captionText="Fig. 9. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846654" httpUri="https://zenodo.org/record/5846654/files/figure.png" pageId="58" pageNumber="59">9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBEFF5A0147FD6BB84F" blockId="58.[189,711,564,626]" box="[189,711,599,626]" pageId="58" pageNumber="59">
|
||
BODY LENGTH. About
|
||
<quantity id="4E0C9283FFB1FFBEFE510148FDBFB84F" box="[438,531,599,626]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="58" pageNumber="59" unit="mm" value="9.4">9.4 mm</quantity>
|
||
; macropterous.
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBEFF5A0186FD8AB994" blockId="58.[187,1400,664,937]" pageId="58" pageNumber="59">COLORATION. Head: dark brown; postocular region with dark patches adjacent to medial ocellar margin, semicircular pale stripe encircling both ocelli; labium lighter than head, yellow. Thorax: anterior pronotal lobe darker than head and posterior pronotal lobe; posterior pronotal lobe with dark yellowish-brown gradient; scutellum dark with contrasting dark yellowish-brown apical spine. Hemelytron (abducted): clavus black with proximal dark yellowish-brown spot and distal yellow stripe; corium black with anteroproximal dark yellowish-brown stripe, posteromedial and distal yellow spots; membrane mostly dark with pale stripe along most of M vein; membranal veins R and M proximally pale. Legs: yellowishbrown. Abdomen: dark brown.</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBEFF5A00CEFD31BE80" blockId="58.[187,1400,976,1214]" pageId="58" pageNumber="59">INTEGUMENT AND VESTITURE. Head and pronotum: finely granulose with dense, long macrosetae interspersed among sparse pubescence; interocular region with two pairs of macrosetae paramedially; antennifer with short lateral setigerous tubercle; morphologically ventral surface of labium with dense, short macrosetae. Thorax: anterolateral angles of pronotal collar with macrosetae; scutellar lateral carinae with long setation. Hemelytron: corium with dense long setation. Legs: posterior row of protuberances on mid and hind femora with three large and three small spines on distal half. Abdomen: ventral surface pubescent, long macrosetae interspersed.</paragraph>
|
||
<paragraph id="894B3F66FFB1FFBFFF5A07FAFC5ABB7B" blockId="58.[186,1400,1253,2019]" lastBlockId="59.[189,1399,265,327]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region about one third of head length, shorter than postocular region (measured to anterior margin of neck); postocular region in dorsal view shorter than eye, lateral margins gently rounded; length of pedicel subequal to head width; maxillary plates ellipsoidal, adjacent to and shorter than clypeus; apices of maxillary plates in dorsal view directed straight; clypeus in dorsal view wider than maxillary plates; clypeal apex round, not narrowed; interocular glabrous markings anterolaterally curved, joined at interocular sulcus paramedially; interocular sulcus in dorsal view nearly straight; eye width in dorsal view about as wide as synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula without lateral protrusion that surpasses buccular margin, flat margin; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.5 times length of segment I. Thorax: pronotal collar in dorsal view narrow medially, with anterolateral angles short, flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins rounded; glabrous markings on pronotum conspicuous and not depressed; median apodeme depression of pronotum deep, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum smoothly rounded, without paramedial lobes; anterior margin of stridulitrum projected into small protuberance; proepimeron with acute protuberance on posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Abdomen: anterior margin of terga prominently carinulate; terga II and III with paired prominent longitudinal carinae, reaching posterior margin of tergum III. Pygophore (
|
||
<figureCitation id="11CF23E3FFB1FFBEFE300441FD9DBD44" box="[471,561,1886,1913]" captionStart="Fig" captionStartId="57.[189,232,1985,2011]" captionTargetBox="[189,1397,687,1944]" captionTargetId="figure-102@57.[189,1399,686,1944]" captionTargetPageId="57" captionText="Fig. 12. Pygophore of species of Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5846662" httpUri="https://zenodo.org/record/5846662/files/figure.png" pageId="58" pageNumber="59">Fig. 12</figureCitation>
|
||
): transverse bridge with rounded posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; short median apical process, upright in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres sinusoidal, apex tapered into squarely rounded tip. Aedeagus (
|
||
<figureCitation id="11CF23E3FFB0FFBFFF210216FEB2BB1E" box="[198,286,265,291]" captionStart="Fig" captionStartId="75.[188,231,1989,2015]" captionTargetBox="[189,1397,760,1946]" captionTargetId="figure-180@75.[189,1397,760,1946]" captionTargetPageId="75" captionText="Fig. 14. Aedeagus of species of Gerbelius Distant, 1903 and Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5846668" httpUri="https://zenodo.org/record/5846668/files/figure.png" pageId="59" pageNumber="60">Fig. 14</figureCitation>
|
||
): endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongueshaped in dorsal view; basal plate extension 5.6 times as long as wide.
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB0FFBFFF0D0271FEE9BBB5" blockId="59.[189,325,366,427]" box="[234,325,366,392]" pageId="59" pageNumber="60">
|
||
<emphasis id="BB80E374FFB0FFBFFF0D0271FEE9BBB5" bold="true" box="[234,325,366,392]" pageId="59" pageNumber="60">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB0FFBFFF5A028EFE95BB96" blockId="59.[189,325,366,427]" box="[189,313,401,427]" pageId="59" pageNumber="60">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1EE6CEDFFB0FFBFFF5A02CDFBA0B826" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="894B3F66FFB0FFBFFF5A02CDFE68BBD1" blockId="59.[189,452,466,492]" box="[189,452,466,492]" pageId="59" pageNumber="60">
|
||
<emphasis id="BB80E374FFB0FFBFFF5A02CDFEFABBD1" bold="true" box="[189,342,466,492]" pageId="59" pageNumber="60">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="11CF23E3FFB0FFBFFE8102CDFE10BBD1" box="[358,444,466,492]" captionStart="Fig" captionStartId="88.[188,231,1983,2009]" captionTargetBox="[189,1398,1222,1944]" captionTargetId="figure-267@88.[189,1398,1222,1944]" captionTargetPageId="88" captionText="Fig. 20. The distribution of species of Gerbelius Distant, 1903 and Voconia Stål, 1866 in Southeast Asia." figureDoi="http://doi.org/10.5281/zenodo.5846682" httpUri="https://zenodo.org/record/5846682/files/figure.png" pageId="59" pageNumber="60">Fig. 20</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB0FFBFFF5A011EFBA0B826" blockId="59.[189,1036,513,539]" box="[189,1036,513,539]" pageId="59" pageNumber="60">
|
||
This species is only known from the
|
||
<typeStatus id="564F81C4FFB0FFBFFD89011EFD0DB826" box="[622,673,513,539]" pageId="59" pageNumber="60">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="F1E37FF6FFB0FFBFFCC2011EFC21B826" box="[805,909,513,539]" name="Malaysia" pageId="59" pageNumber="60">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F1E37FF6FFB0FFBFFC7E011EFBA4B826" box="[921,1032,513,539]" name="Malaysia" pageId="59" pageNumber="60">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1EE6CEDFFB0FFBFFF5A015DFD09B8EF" pageId="59" pageNumber="60" type="discussion">
|
||
<paragraph id="894B3F66FFB0FFBFFF5A015DFE83B861" blockId="59.[189,303,578,604]" box="[189,303,578,604]" pageId="59" pageNumber="60">
|
||
<heading id="D203880AFFB0FFBFFF5A015DFE83B861" bold="true" box="[189,303,578,604]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="BB80E374FFB0FFBFFF5A015DFE83B861" bold="true" box="[189,303,578,604]" pageId="59" pageNumber="60">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="894B3F66FFB0FFBFFF5A016EFD09B8EF" blockId="59.[189,1398,625,722]" pageId="59" pageNumber="60">
|
||
The
|
||
<typeStatus id="564F81C4FFB0FFBFFF10016EFEF2B8B6" box="[247,350,625,651]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
was preserved in ethanol when the abdomen was dissected for DNA extraction. We acquired sufficient DNA in our NGS library for Illumina sequencing for future phylogenomic studies, referred to as DNA voucher R_CW 5472.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |