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<document id="064B40900269CAC52A457E06E0FDCC60" ID-DOI="10.11646/zootaxa.3985.2.4" ID-GBIF-Dataset="4642635d-3cb2-4749-bd66-ecf4bb47697a" ID-ISSN="1175-5326" ID-Zenodo-Dep="253999" ID-ZooBank="77F77A09-2767-4EC5-ADA8-46A28BF5AE76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461257459696" checkinUser="plazi" docAuthor="Marusik, Yuri M., Omelko, Mikhail M. &amp; Koponen, Seppo" docDate="2015" docId="CA3F87D3F469A40C66C4FACDFDD01586" docLanguage="en" docName="zt03985p264.pdf" docOrigin="Zootaxa 3985 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melecosa Marusik, Omelko &amp; Koponen, 2015, gen. n." docType="treatment" docVersion="8" lastPageNumber="259" masterDocId="3606FFABF46FA40B6653FFA5FFB5126C" masterDocTitle="A survey of East Palaearctic Lycosidae (Araneae). 11. Two new genera from the Acantholycosa complex" masterLastPageNumber="264" masterPageNumber="252" pageNumber="258" updateTime="1698608407070" updateUser="plazi">
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<mods:title id="01CBB65A6E7941591A932D106573EE18">A survey of East Palaearctic Lycosidae (Araneae). 11. Two new genera from the Acantholycosa complex</mods:title>
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<mods:namePart id="770850DF7CAC85F84A6BE27421679CB6">Marusik, Yuri M.</mods:namePart>
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<mods:namePart id="F29C975D11C7A936BEA899A8FD232163">Omelko, Mikhail M.</mods:namePart>
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<mods:namePart id="FFA5942647594F1C747858716368E303">Koponen, Seppo</mods:namePart>
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<mods:date id="4C986E2A001BF13A1A1C00299B833EC7">2015</mods:date>
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<treatment id="CA3F87D3F469A40C66C4FACDFDD01586" ID-DOI="http://doi.org/10.5281/zenodo.5661051" ID-GBIF-Taxon="119636273" ID-Zenodo-Dep="5661051" LSID="urn:lsid:plazi:treatment:CA3F87D3F469A40C66C4FACDFDD01586" httpUri="http://treatment.plazi.org/id/CA3F87D3F469A40C66C4FACDFDD01586" lastPageId="7" lastPageNumber="259" pageId="6" pageNumber="258">
<subSubSection id="0A8C654EF469A40D66C4FACDFED717EF" box="[151,354,1384,1411]" pageId="6" pageNumber="258" type="nomenclature">
<paragraph id="422936C5F469A40D66C4FACDFED717EF" blockId="6.[151,354,1384,1411]" box="[151,354,1384,1411]" pageId="6" pageNumber="258">
<heading id="196181A9F469A40D66C4FACDFED717EF" bold="true" box="[151,354,1384,1411]" fontSize="11" level="1" pageId="6" pageNumber="258" reason="1">
<emphasis id="70E2EAD7F469A40D66C4FACDFED717EF" bold="true" box="[151,354,1384,1411]" pageId="6" pageNumber="258">
<taxonomicName id="85964D46F469A40D66C4FACDFEBC17EE" ID-CoL="5NNB" box="[151,265,1384,1410]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F469A40D66C4FACDFEBC17EE" bold="true" box="[151,265,1384,1410]" italics="true" pageId="6" pageNumber="258">Melecosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BD157ACF469A40D6743FACCFED717EF" box="[272,354,1385,1411]" pageId="6" pageNumber="258" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="0A8C654EF469A40D66C4FA0CFC0517AD" box="[151,944,1449,1474]" pageId="6" pageNumber="258" type="materials_examined">
<paragraph id="422936C5F469A40D66C4FA0CFC0517AD" blockId="6.[151,1436,1449,2037]" box="[151,944,1449,1474]" pageId="6" pageNumber="258">
<emphasis id="70E2EAD7F469A40D66C4FA0CFE9E17AE" bold="true" box="[151,299,1449,1474]" pageId="6" pageNumber="258">
<typeStatus id="9D2D8867F469A40D66C4FA0CFF6717AE" box="[151,210,1449,1474]" pageId="6" pageNumber="258">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="85964D46F469A40D6769FA0CFC1917AD" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[314,940,1449,1474]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="70E2EAD7F469A40D6769FA0CFE4B17AE" box="[314,510,1449,1474]" italics="true" pageId="6" pageNumber="258">Sibirocosa alpina</emphasis>
<bibRefCitation id="26074B34F469A40D6455FA0CFC1917AD" author="Marusik" box="[518,940,1449,1474]" pageId="6" pageNumber="258" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12, 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0A8C654EF469A40D6694FA69FC031464" pageId="6" pageNumber="258" type="etymology">
<paragraph id="422936C5F469A40D6694FA69FC031464" blockId="6.[151,1436,1449,2037]" pageId="6" pageNumber="258">
<emphasis id="70E2EAD7F469A40D6694FA69FEF31789" bold="true" box="[199,326,1484,1509]" pageId="6" pageNumber="258">Etymology</emphasis>
. The generic name is formed from two elements, with the first element derived from the scientific name of the badger (
<taxonomicName id="85964D46F469A40D672DFA4AFE741464" box="[382,449,1519,1544]" class="Mammalia" family="Mustelidae" genus="Meles" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="258" phylum="Chordata" rank="genus">
<emphasis id="70E2EAD7F469A40D672DFA4AFE741464" box="[382,449,1519,1544]" italics="true" pageId="6" pageNumber="258">Meles</emphasis>
</taxonomicName>
) and ending -cosa. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="0A8C654EF469A40D6694F9B7FEA214B7" pageId="6" pageNumber="258" type="diagnosis">
<paragraph id="422936C5F469A40D6694F9B7FEA214B7" blockId="6.[151,1436,1449,2037]" pageId="6" pageNumber="258">
<emphasis id="70E2EAD7F469A40D6694F9B7FE8F1447" bold="true" box="[199,314,1554,1579]" pageId="6" pageNumber="258">Diagnosis</emphasis>
.
<taxonomicName id="85964D46F469A40D671AF9B7FE011447" box="[329,436,1554,1579]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F469A40D671AF9B7FE011447" box="[329,436,1554,1579]" italics="true" pageId="6" pageNumber="258">Melecosa</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F469A40D67EFF9B7FDB91447" bold="true" box="[444,524,1554,1579]" pageId="6" pageNumber="258">
<taxonomicNameLabel id="6BD157ACF469A40D67EFF9B7FDB91447" box="[444,524,1554,1579]" pageId="6" pageNumber="258" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from the somewhat similar
<taxonomicName id="85964D46F469A40D6269F9B7FB041447" box="[1082,1201,1554,1579]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D6269F9B7FB041447" box="[1082,1201,1554,1579]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
by having 3 pairs of ventral tibial spines on leg I (
<figureCitation id="DAAD2A40F469A40D67BFF990FDE01422" box="[492,597,1589,1614]" captionStart="FIGURES 1 ‒ 6" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[292,1298,194,1807]" captionTargetId="figure@3.[275,1305,194,1811]" captionTargetPageId="3" captionText="FIGURES 1 ‒ 6. Habitus and male leg I of Gulocosa eskovi sp. n. (1 ‒ 3) and Melecosa alpina (4 ‒ 6). 1, 4 female, dorsal; 2, 5 male, dorsal; 3, 6 leg I prolateral." httpUri="https://zenodo.org/record/254000/files/figure.png" pageId="6" pageNumber="258">Figs 5‒6</figureCitation>
; 5–6 pairs in
<taxonomicName id="85964D46F469A40D64A7F990FCDF1422" box="[756,874,1589,1614]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D64A7F990FCDF1422" box="[756,874,1589,1614]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
); tegular apophysis with two pointed arms (
<figureCitation id="DAAD2A40F469A40D6339F990FF55141D" captionStart="FIGURES 40 ‒ 44" captionStartId="8.[151,264,1667,1690]" captionTargetBox="[197,1398,774,1647]" captionTargetId="figure@8.[188,1399,763,1650]" captionTargetPageId="8" captionText="FIGURES 40 ‒ 44. Male palp of Melecosa alpina. 40, 42 ventral; 41 retrolateral; 43 ‒ 44 embolic division, retrolateral and from above, respectively. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254005/files/figure.png" pageId="6" pageNumber="258">Figs 40‒42</figureCitation>
,
<figureCitation id="DAAD2A40F469A40D66BEF9FCFE80141E" box="[237,309,1625,1650]" captionStart="FIGURES 45 ‒ 50" captionStartId="9.[151,264,913,936]" captionTargetBox="[193,1394,193,892]" captionTargetId="figure@9.[193,1394,193,893]" captionTargetPageId="9" captionText="FIGURES 45 ‒ 50. Copulatory organs of Melecosa alpina. 45 ‒ 46 male palp, subapical and subapical-retrolateral respectively; 47 ‒ 48 embolic division, ventral and from above respectively; 49 ‒ 50 epigyne, ventral and dorsal respectively. Abbreviations: Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254006/files/figure.png" pageId="6" pageNumber="258">45‒46</figureCitation>
) (basal arm in
<taxonomicName id="85964D46F469A40D67B7F9FCFDEE141E" box="[484,603,1625,1650]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D67B7F9FCFDEE141E" box="[484,603,1625,1650]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
only, cf.
<figureCitation id="DAAD2A40F469A40D6499F9FCFCAA141D" box="[714,799,1625,1650]" captionStart="FIGURES 7 ‒ 12" captionStartId="4.[151,264,1432,1455]" captionTargetBox="[175,1391,212,1374]" captionTargetId="figure@4.[171,1399,193,1379]" captionTargetPageId="4" captionText="FIGURES 7 ‒ 12. Male palp of Gulocosa eskovi sp. n. (7 ‒ 9), Sibirocosa nadolnyi (10: after Omelko &amp; Marusik 2013) and Acantholycosa norvegica (11 ‒ 12). 7 ‒ 8, 10 ‒ 11 ventral; 9, 12 from above. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Et embolic tooth, Pa palea, Pp process of palea, Pw wart of palea, Ta terminal apophysis." httpUri="https://zenodo.org/record/254001/files/figure.png" pageId="6" pageNumber="258">Fig. 10</figureCitation>
), large paleal process (
<emphasis id="70E2EAD7F469A40D627BF9FFFBF2141D" box="[1064,1095,1626,1649]" italics="true" pageId="6" pageNumber="258">Pp</emphasis>
) and lack of distinct terminal apophysis (
<figureCitation id="DAAD2A40F469A40D674EF9D9FEC314F9" box="[285,374,1660,1685]" captionStart="FIGURES 45 ‒ 50" captionStartId="9.[151,264,913,936]" captionTargetBox="[193,1394,193,892]" captionTargetId="figure@9.[193,1394,193,893]" captionTargetPageId="9" captionText="FIGURES 45 ‒ 50. Copulatory organs of Melecosa alpina. 45 ‒ 46 male palp, subapical and subapical-retrolateral respectively; 47 ‒ 48 embolic division, ventral and from above respectively; 49 ‒ 50 epigyne, ventral and dorsal respectively. Abbreviations: Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254006/files/figure.png" pageId="6" pageNumber="258">Fig. 46</figureCitation>
) (terminal apophysis well developed, more robust than embolus in
<taxonomicName id="85964D46F469A40D62E3F9D9FA9214F9" box="[1200,1319,1660,1685]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D62E3F9D9FA9214F9" box="[1200,1319,1660,1685]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
), shallow epigynal fovea (
<figureCitation id="DAAD2A40F469A40D671DF93AFE6714D4" box="[334,466,1695,1720]" captionStart="FIGURES 45 ‒ 50" captionStartId="9.[151,264,913,936]" captionTargetBox="[193,1394,193,892]" captionTargetId="figure@9.[193,1394,193,893]" captionTargetPageId="9" captionText="FIGURES 45 ‒ 50. Copulatory organs of Melecosa alpina. 45 ‒ 46 male palp, subapical and subapical-retrolateral respectively; 47 ‒ 48 embolic division, ventral and from above respectively; 49 ‒ 50 epigyne, ventral and dorsal respectively. Abbreviations: Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254006/files/figure.png" pageId="6" pageNumber="258">Figs 49‒50</figureCitation>
), two apical pockets (one in
<taxonomicName id="85964D46F469A40D654FF93AFC2614D4" box="[796,915,1695,1720]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D654FF93AFC2614D4" box="[796,915,1695,1720]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
,
<figureCitation id="DAAD2A40F469A40D65F1F93AFC4114D4" box="[930,1012,1695,1720]" captionStart="FIGURES 28 ‒ 39" captionStartId="7.[151,264,1590,1613]" captionTargetBox="[182,1379,183,1576]" captionTargetId="figure@7.[182,1399,183,1578]" captionTargetPageId="7" captionText="FIGURES 28 ‒ 39. Epigyne of Gulocosa eskovi sp. n. (28 ‒ 31), Melecosa alpina (32 ‒ 33), Acantholycosa norvegica (34 ‒ 36), Sibirocosa koponeni (37 ‒ 38) an S. trilikauskasi (39). 28 ‒ 30, 32, 34 ‒ 35, 37 ventral; 31, 33, 36, 38 ‒ 39 dorsal. 37 ‒ 39 after Omelko &amp; Marusik (2013). Abbreviations: Ap apical pocket, Ba basal arm, Fo fovea, Lm lateral margin, Rh head of receptacle, Sb septal base, Se septum, Ss septal stem." httpUri="https://zenodo.org/record/254004/files/figure.png" pageId="6" pageNumber="258">Fig. 37</figureCitation>
) and large heads of receptacles (
<figureCitation id="DAAD2A40F469A40D6339F93AFF5514B7" captionStart="FIGURES 28 ‒ 39" captionStartId="7.[151,264,1590,1613]" captionTargetBox="[182,1379,183,1576]" captionTargetId="figure@7.[182,1399,183,1578]" captionTargetPageId="7" captionText="FIGURES 28 ‒ 39. Epigyne of Gulocosa eskovi sp. n. (28 ‒ 31), Melecosa alpina (32 ‒ 33), Acantholycosa norvegica (34 ‒ 36), Sibirocosa koponeni (37 ‒ 38) an S. trilikauskasi (39). 28 ‒ 30, 32, 34 ‒ 35, 37 ventral; 31, 33, 36, 38 ‒ 39 dorsal. 37 ‒ 39 after Omelko &amp; Marusik (2013). Abbreviations: Ap apical pocket, Ba basal arm, Fo fovea, Lm lateral margin, Rh head of receptacle, Sb septal base, Se septum, Ss septal stem." httpUri="https://zenodo.org/record/254004/files/figure.png" pageId="6" pageNumber="258">Figs 32‒33</figureCitation>
,
<figureCitation id="DAAD2A40F469A40D66B8F967FEBE14B7" box="[235,267,1730,1755]" captionStart="FIGURES 45 ‒ 50" captionStartId="9.[151,264,913,936]" captionTargetBox="[193,1394,193,892]" captionTargetId="figure@9.[193,1394,193,893]" captionTargetPageId="9" captionText="FIGURES 45 ‒ 50. Copulatory organs of Melecosa alpina. 45 ‒ 46 male palp, subapical and subapical-retrolateral respectively; 47 ‒ 48 embolic division, ventral and from above respectively; 49 ‒ 50 epigyne, ventral and dorsal respectively. Abbreviations: Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254006/files/figure.png" pageId="6" pageNumber="258">50</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A8C654EF469A40C6694F940FDD01586" lastPageId="7" lastPageNumber="259" pageId="6" pageNumber="258" type="description">
<paragraph id="422936C5F469A40D6694F940FDDE1492" blockId="6.[151,1436,1449,2037]" box="[199,619,1765,1790]" pageId="6" pageNumber="258">
<emphasis id="70E2EAD7F469A40D6694F940FEE41492" bold="true" box="[199,337,1765,1790]" pageId="6" pageNumber="258">Description</emphasis>
. See species description.
</paragraph>
<paragraph id="422936C5F469A40D6694F8ACFB281599" blockId="6.[151,1436,1449,2037]" pageId="6" pageNumber="258">
<emphasis id="70E2EAD7F469A40D6694F8ACFEDC154E" bold="true" box="[199,361,1801,1826]" pageId="6" pageNumber="258">Relationships</emphasis>
. As in
<taxonomicName id="85964D46F469A40D6790F8ACFD9A154E" box="[451,559,1801,1826]" class="Arachnida" family="Lycosidae" genus="Gulocosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F469A40D6790F8ACFD9A154E" box="[451,559,1801,1826]" italics="true" pageId="6" pageNumber="258">Gulocosa</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F469A40D646AF8ACFD3C154E" bold="true" box="[569,649,1801,1826]" pageId="6" pageNumber="258">
<taxonomicNameLabel id="6BD157ACF469A40D646AF8ACFD3C154E" box="[569,649,1801,1826]" pageId="6" pageNumber="258" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
, it is difficult to trace the relationships of this new genus based on somatic characters or the features of copulatory organs. The epigyne in this genus is rather similar to that of
<taxonomicName id="85964D46F469A40D66C4F8EAFEBB1504" box="[151,270,1871,1896]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D66C4F8EAFEBB1504" box="[151,270,1871,1896]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
and that is why it was originally placed in the genus. The two genera, although generally similar in appearance, clearly differ in the fovea depth (
<figureCitation id="DAAD2A40F469A40D64CBF8D7FCAF15E7" box="[664,794,1906,1931]" captionStart="FIGURES 28 ‒ 39" captionStartId="7.[151,264,1590,1613]" captionTargetBox="[182,1379,183,1576]" captionTargetId="figure@7.[182,1399,183,1578]" captionTargetPageId="7" captionText="FIGURES 28 ‒ 39. Epigyne of Gulocosa eskovi sp. n. (28 ‒ 31), Melecosa alpina (32 ‒ 33), Acantholycosa norvegica (34 ‒ 36), Sibirocosa koponeni (37 ‒ 38) an S. trilikauskasi (39). 28 ‒ 30, 32, 34 ‒ 35, 37 ventral; 31, 33, 36, 38 ‒ 39 dorsal. 37 ‒ 39 after Omelko &amp; Marusik (2013). Abbreviations: Ap apical pocket, Ba basal arm, Fo fovea, Lm lateral margin, Rh head of receptacle, Sb septal base, Se septum, Ss septal stem." httpUri="https://zenodo.org/record/254004/files/figure.png" pageId="6" pageNumber="258">Figs 37‒39</figureCitation>
,
<figureCitation id="DAAD2A40F469A40D6575F8D7FCC515E7" box="[806,880,1906,1931]" captionStart="FIGURES 45 ‒ 50" captionStartId="9.[151,264,913,936]" captionTargetBox="[193,1394,193,892]" captionTargetId="figure@9.[193,1394,193,893]" captionTargetPageId="9" captionText="FIGURES 45 ‒ 50. Copulatory organs of Melecosa alpina. 45 ‒ 46 male palp, subapical and subapical-retrolateral respectively; 47 ‒ 48 embolic division, ventral and from above respectively; 49 ‒ 50 epigyne, ventral and dorsal respectively. Abbreviations: Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254006/files/figure.png" pageId="6" pageNumber="258">49‒50</figureCitation>
), number of apical pockets and size of receptacle heads (
<figureCitation id="DAAD2A40F469A40D66B5F830FE0815C2" box="[230,445,1941,1966]" captionStart="FIGURES 28 ‒ 39" captionStartId="7.[151,264,1590,1613]" captionTargetBox="[182,1379,183,1576]" captionTargetId="figure@7.[182,1399,183,1578]" captionTargetPageId="7" captionText="FIGURES 28 ‒ 39. Epigyne of Gulocosa eskovi sp. n. (28 ‒ 31), Melecosa alpina (32 ‒ 33), Acantholycosa norvegica (34 ‒ 36), Sibirocosa koponeni (37 ‒ 38) an S. trilikauskasi (39). 28 ‒ 30, 32, 34 ‒ 35, 37 ventral; 31, 33, 36, 38 ‒ 39 dorsal. 37 ‒ 39 after Omelko &amp; Marusik (2013). Abbreviations: Ap apical pocket, Ba basal arm, Fo fovea, Lm lateral margin, Rh head of receptacle, Sb septal base, Se septum, Ss septal stem." httpUri="https://zenodo.org/record/254004/files/figure.png" pageId="6" pageNumber="258">Figs 32‒33, 38‒39</figureCitation>
). The males of the two genera differ by the shape of the tegular apophysis and the width and thickness of the embolus. The most striking difference between the two genera is the lack of a distinct terminal apophysis in
<taxonomicName id="85964D46F469A40D6778F879FE231599" box="[299,406,2012,2037]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F469A40D6778F879FE231599" box="[299,406,2012,2037]" italics="true" pageId="6" pageNumber="258">Melecosa</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F469A40D67CEF879FE591599" bold="true" box="[413,492,2012,2037]" pageId="6" pageNumber="258">
<taxonomicNameLabel id="6BD157ACF469A40D67CEF879FE591599" box="[413,492,2012,2037]" pageId="6" pageNumber="258" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
and a strongly developed one in
<taxonomicName id="85964D46F469A40D650DF879FC611599" box="[862,980,2012,2037]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F469A40D650DF879FC611599" box="[862,980,2012,2037]" italics="true" pageId="6" pageNumber="258">Sibirocosa</emphasis>
</taxonomicName>
(
<figureCitation id="DAAD2A40F469A40D65B7F879FB8F1599" box="[996,1082,2012,2037]" captionStart="FIGURES 7 ‒ 12" captionStartId="4.[151,264,1432,1455]" captionTargetBox="[175,1391,212,1374]" captionTargetId="figure@4.[171,1399,193,1379]" captionTargetPageId="4" captionText="FIGURES 7 ‒ 12. Male palp of Gulocosa eskovi sp. n. (7 ‒ 9), Sibirocosa nadolnyi (10: after Omelko &amp; Marusik 2013) and Acantholycosa norvegica (11 ‒ 12). 7 ‒ 8, 10 ‒ 11 ventral; 9, 12 from above. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Et embolic tooth, Pa palea, Pp process of palea, Pw wart of palea, Ta terminal apophysis." httpUri="https://zenodo.org/record/254001/files/figure.png" pageId="6" pageNumber="258">Figs 10</figureCitation>
,
<figureCitation id="DAAD2A40F469A40D6215F878FB241599" box="[1094,1169,2013,2037]" captionStart="FIGURES 40 ‒ 44" captionStartId="8.[151,264,1667,1690]" captionTargetBox="[197,1398,774,1647]" captionTargetId="figure@8.[188,1399,763,1650]" captionTargetPageId="8" captionText="FIGURES 40 ‒ 44. Male palp of Melecosa alpina. 40, 42 ventral; 41 retrolateral; 43 ‒ 44 embolic division, retrolateral and from above, respectively. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254005/files/figure.png" pageId="6" pageNumber="258">40‒42</figureCitation>
).
</paragraph>
<caption id="16E9664DF468A40C66C4F993FD8E14C5" httpUri="https://zenodo.org/record/254004/files/figure.png" pageId="7" pageNumber="259" targetBox="[182,1379,183,1576]" targetPageId="7">
<paragraph id="422936C5F468A40C66C4F993FD8E14C5" blockId="7.[151,1436,1590,1705]" pageId="7" pageNumber="259">
<emphasis id="70E2EAD7F468A40C66C4F993FEE01420" bold="true" box="[151,341,1590,1613]" pageId="7" pageNumber="259">FIGURES 28‒39.</emphasis>
Epigyne of
<taxonomicName id="85964D46F468A40C6784F993FD351420" box="[471,640,1590,1612]" class="Arachnida" family="Lycosidae" genus="Gulocosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="eskovi" status="sp. nov.">
<emphasis id="70E2EAD7F468A40C6784F993FD351420" box="[471,640,1590,1612]" italics="true" pageId="7" pageNumber="259">Gulocosa eskovi</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F468A40C64DBF992FD761420" bold="true" box="[648,707,1591,1612]" pageId="7" pageNumber="259">
<taxonomicNameLabel id="6BD157ACF468A40C64DBF992FD761420" box="[648,707,1591,1612]" pageId="7" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(28‒31),
<taxonomicName id="85964D46F468A40C6579F993FC611420" box="[810,980,1590,1612]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="70E2EAD7F468A40C6579F993FC611420" box="[810,980,1590,1612]" italics="true" pageId="7" pageNumber="259">Melecosa alpina</emphasis>
</taxonomicName>
(32‒33),
<taxonomicName id="85964D46F468A40C6268F993FA881421" box="[1083,1341,1590,1613]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="norvegica">
<emphasis id="70E2EAD7F468A40C6268F993FA881421" box="[1083,1341,1590,1613]" italics="true" pageId="7" pageNumber="259">Acantholycosa norvegica</emphasis>
</taxonomicName>
(34‒36),
<taxonomicName id="85964D46F468A40C66C4F9F0FEDC1407" box="[151,361,1621,1643]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="70E2EAD7F468A40C66C4F9F0FEDC1407" box="[151,361,1621,1643]" italics="true" pageId="7" pageNumber="259">Sibirocosa koponeni</emphasis>
</taxonomicName>
(37‒38) an
<taxonomicName id="85964D46F468A40C67BFF9F3FD331407" box="[492,646,1621,1643]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="trilikauskasi">
<emphasis id="70E2EAD7F468A40C67BFF9F3FD331407" box="[492,646,1621,1643]" italics="true" pageId="7" pageNumber="259">S. trilikauskasi</emphasis>
</taxonomicName>
(39). 28‒30, 32, 34‒35, 37 ventral; 31, 33, 36, 38‒39 dorsal. 37‒39 after Omelko &amp; Marusik (2013). Abbreviations:
<emphasis id="70E2EAD7F468A40C641CF9D0FDD914E6" box="[591,620,1653,1674]" italics="true" pageId="7" pageNumber="259">Ap</emphasis>
apical pocket,
<emphasis id="70E2EAD7F468A40C6557F9D0FC9514E6" box="[772,800,1653,1674]" italics="true" pageId="7" pageNumber="259">Ba</emphasis>
basal arm,
<emphasis id="70E2EAD7F468A40C65C6F9D0FC0414E6" box="[917,945,1653,1674]" italics="true" pageId="7" pageNumber="259">Fo</emphasis>
fovea,
<emphasis id="70E2EAD7F468A40C65AFF9D0FBA914E6" box="[1020,1052,1653,1674]" italics="true" pageId="7" pageNumber="259">Lm</emphasis>
lateral margin,
<emphasis id="70E2EAD7F468A40C62EFF9D1FB6D14E6" box="[1212,1240,1652,1674]" italics="true" pageId="7" pageNumber="259">Rh</emphasis>
head of receptacle,
<emphasis id="70E2EAD7F468A40C66C4F936FF0414C5" box="[151,177,1683,1705]" italics="true" pageId="7" pageNumber="259">Sb</emphasis>
septal base,
<emphasis id="70E2EAD7F468A40C6762F936FEFC14C4" box="[305,329,1683,1704]" italics="true" pageId="7" pageNumber="259">Se</emphasis>
septum,
<emphasis id="70E2EAD7F468A40C67F6F936FE0914C4" box="[421,444,1683,1704]" italics="true" pageId="7" pageNumber="259">Ss</emphasis>
septal stem.
</paragraph>
</caption>
<paragraph id="422936C5F468A40C6694F95CFE9B1512" blockId="7.[151,1437,1785,2026]" pageId="7" pageNumber="259">
Three genera,
<taxonomicName id="85964D46F468A40C6722F95CFE69157E" box="[369,476,1785,1810]" class="Arachnida" family="Lycosidae" genus="Gulocosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F468A40C6722F95CFE69157E" box="[369,476,1785,1810]" italics="true" pageId="7" pageNumber="259">Gulocosa</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F468A40C67BBF95CFD89157E" bold="true" box="[488,572,1785,1810]" pageId="7" pageNumber="259">
<taxonomicNameLabel id="6BD157ACF468A40C67BBF95CFD89157E" box="[488,572,1785,1810]" pageId="7" pageNumber="259" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="85964D46F468A40C641DF95CFD0F157E" box="[590,698,1785,1810]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70E2EAD7F468A40C641DF95CFD0F157E" box="[590,698,1785,1810]" italics="true" pageId="7" pageNumber="259">Melecosa</emphasis>
</taxonomicName>
<emphasis id="70E2EAD7F468A40C6496F95CFCAC157E" bold="true" box="[709,793,1785,1810]" pageId="7" pageNumber="259">
<taxonomicNameLabel id="6BD157ACF468A40C6496F95CFCAC157E" box="[709,793,1785,1810]" pageId="7" pageNumber="259" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="85964D46F468A40C650BF95CFC7A157E" box="[856,975,1785,1810]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="70E2EAD7F468A40C650BF95CFC7A157E" box="[856,975,1785,1810]" italics="true" pageId="7" pageNumber="259">Sibirocosa</emphasis>
</taxonomicName>
have, a high position (location) of the embolic base, which lies almost at the same level as the upper edge of palea (
<figureCitation id="DAAD2A40F468A40C625AF8B8FBCB155A" box="[1033,1150,1821,1846]" captionStart="FIGURES 7 ‒ 12" captionStartId="4.[151,264,1432,1455]" captionTargetBox="[175,1391,212,1374]" captionTargetId="figure@4.[171,1399,193,1379]" captionTargetPageId="4" captionText="FIGURES 7 ‒ 12. Male palp of Gulocosa eskovi sp. n. (7 ‒ 9), Sibirocosa nadolnyi (10: after Omelko &amp; Marusik 2013) and Acantholycosa norvegica (11 ‒ 12). 7 ‒ 8, 10 ‒ 11 ventral; 9, 12 from above. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Et embolic tooth, Pa palea, Pp process of palea, Pw wart of palea, Ta terminal apophysis." httpUri="https://zenodo.org/record/254001/files/figure.png" pageId="7" pageNumber="259">Figs 8, 10</figureCitation>
,
<figureCitation id="DAAD2A40F468A40C62D8F8BBFB1E155A" box="[1163,1195,1822,1846]" captionStart="FIGURES 40 ‒ 44" captionStartId="8.[151,264,1667,1690]" captionTargetBox="[197,1398,774,1647]" captionTargetId="figure@8.[188,1399,763,1650]" captionTargetPageId="8" captionText="FIGURES 40 ‒ 44. Male palp of Melecosa alpina. 40, 42 ventral; 41 retrolateral; 43 ‒ 44 embolic division, retrolateral and from above, respectively. Abbreviations: Aa apical arm, Ba basal arm, Em embolus, Pp process of palea." httpUri="https://zenodo.org/record/254005/files/figure.png" pageId="7" pageNumber="259">40</figureCitation>
), which is unique for Pardosinae. These characters can be connected with the thickness of embolus but do not indicate close relationships.
</paragraph>
<paragraph id="422936C5F468A40C6694F82CFDD01586" blockId="7.[151,1437,1785,2026]" pageId="7" pageNumber="259">
<emphasis id="70E2EAD7F468A40C6694F82CFF4A15CE" bold="true" box="[199,255,1929,1954]" pageId="7" pageNumber="259">Note</emphasis>
. We cannot homologise parts of the embolic division in this genus with certainty with other Pardosinae genera. It appears that this genus lacks a terminal apophysis and instead has a huge palear process.
<emphasis id="70E2EAD7F468A40C6694F874FEEA1586" bold="true" box="[199,351,2001,2026]" pageId="7" pageNumber="259">Composition</emphasis>
. Only the
<typeStatus id="9D2D8867F468A40C6786F877FDB01586" box="[469,517,2002,2026]" pageId="7" pageNumber="259">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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