treatments-xml/data/8E/10/87/8E1087A9FFFC896CFF2D1EF8A68AFD9A.xml

<document id="BD4F3F04A303AF361FFF03D33EB85860" ID-DOI="10.5281/zenodo.210960" ID-GBIF-Dataset="2ebe66ef-7e88-4eac-8085-b2df5c025467" ID-ISSN="1175-5326" ID-Zenodo-Dep="210960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460485119356" checkinUser="plazi" docAuthor="Teslenko, Valentina A." docDate="2012" docId="8E1087A9FFFC896CFF2D1EF8A68AFD9A" docLanguage="en" docName="zt03329p018.pdf" docOrigin="Zootaxa 3329" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Skwala compacta McLachlan 1872, comb. nov." docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="7229FFD1FFF1897CFFBA1E6FA253FFA6" masterDocTitle="A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="14" updateTime="1698308897074" updateUser="plazi">
<mods:mods id="8542C0A201232C9CECA403343C40011B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5F7FA54E0A5D8C1317C894F38C628896">
<mods:title id="FD9F53BFCBFC620D91FA7CBFC74DCB2E">A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae)</mods:title>
</mods:titleInfo>
<mods:name id="B03B192F848F41BA3F6096DF56DB5A9F" type="personal">
<mods:role id="60091EFD6556338D9AD7ED832A4D5E04">
<mods:roleTerm id="5CF03947D41B7647D35A968CFCAEB518">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2AF4626B09C89FABEF19589F26F46416">Teslenko, Valentina A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D60D9F694D7CF440B4FF42D24E22F39E">text</mods:typeOfResource>
<mods:relatedItem id="9F45B987970197B7BAB5D64A3CF49E7A" type="host">
<mods:titleInfo id="06B6B1C416C309E3FA341D92DFE7DD31">
<mods:title id="4F4A12E9FCC83E5EBF8E1193A99E1DBB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9A474F51DB377F44947FF85A9CF33BC2">
<mods:date id="A21027B32C73359C0CE0FE2BD93C9C51">2012</mods:date>
<mods:detail id="ECADB082D7DFA45CFC1D258BCC92A5D6" type="volume">
<mods:number id="07FCEBBE0BC69D1EA940CF2E1F1CABC4">3329</mods:number>
</mods:detail>
<mods:extent id="02941536660237F80C2D870C01B14B5E" unit="page">
<mods:start id="DDCBAF9A3056A202FC3077F6C952A565">1</mods:start>
<mods:end id="266618F268A1288B7CEBE379A853D6A7">18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3AD8ED3F6855C0FB04473FE9E52B8FBE">journal article</mods:classification>
<mods:identifier id="0722C66041DC6FE6A9032AD82646B273" type="DOI">10.5281/zenodo.210960</mods:identifier>
<mods:identifier id="27BAF89155D51BDAA6ED325849566995" type="GBIF-Dataset">2ebe66ef-7e88-4eac-8085-b2df5c025467</mods:identifier>
<mods:identifier id="7603C30C35528AD38AF893D4C3FD4635" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FA3C0CA4A182837DA6A97355BAC08B9D" type="Zenodo-Dep">210960</mods:identifier>
</mods:mods>
<treatment id="8E1087A9FFFC896CFF2D1EF8A68AFD9A" ID-DOI="http://doi.org/10.5281/zenodo.6174160" ID-GBIF-Taxon="119518129" ID-Zenodo-Dep="6174160" LSID="urn:lsid:plazi:treatment:8E1087A9FFFC896CFF2D1EF8A68AFD9A" httpUri="http://treatment.plazi.org/id/8E1087A9FFFC896CFF2D1EF8A68AFD9A" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="4EA36534FFFC8971FF2D1EF8A321FE14" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="060636BFFFFC8971FF2D1EF8A0B9FF17" blockId="13.[151,746,151,210]" box="[151,746,151,177]" pageId="13" pageNumber="14">
<heading id="5D4E81D3FFFC8971FF2D1EF8A0B9FF17" bold="true" box="[151,746,151,177]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="34CDEAADFFFC8971FF2D1EF8A0B9FF17" bold="true" box="[151,746,151,177]" pageId="13" pageNumber="14">
<taxonomicName id="C1B94D3CFFFC8971FF2D1EF8A00FFF17" ID-CoL="6YRPF" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[151,604,151,177]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="compacta" status="comb. nov.">
<emphasis id="34CDEAADFFFC8971FF2D1EF8A33AFF17" bold="true" box="[151,361,151,177]" italics="true" pageId="13" pageNumber="14">Skwala compacta</emphasis>
(
<bibRefCitation id="62284B4EFFFC8971FEC21EF8A007FF17" author="McLachlan" box="[376,596,151,177]" pageId="13" pageNumber="14" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="2FFE57D6FFFC8971FDD91EF8A0B9FF17" box="[611,746,151,177]" pageId="13" pageNumber="14">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="060636BFFFFC8971FF2D1ED5A374FF74" blockId="13.[151,746,151,210]" box="[151,295,186,210]" pageId="13" pageNumber="14">
(
<figureCitation id="9E822A3AFFFC8971FF251ED5A34CFF74" box="[159,287,186,210]" captionStart-0="FIGURES 39 – 41" captionStart-1="FIGURES 42 – 45" captionStart-2="FIGURES 46 – 48" captionStartId-0="14.[151,264,1692,1715]" captionStartId-1="15.[151,264,967,990]" captionStartId-2="15.[151,264,1980,2003]" captionTargetBox-0="[268,1297,543,1657]" captionTargetBox-1="[267,1322,287,936]" captionTargetBox-2="[264,1322,1088,1953]" captionTargetId-0="figure@14.[264,1323,543,1668]" captionTargetId-1="figure@15.[264,1323,287,942]" captionTargetId-2="figure@15.[264,1323,1088,1959]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="15" captionText-0="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." captionText-1="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." captionText-2="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri-0="https://zenodo.org/record/210970/files/figure.png" httpUri-1="https://zenodo.org/record/210971/files/figure.png" httpUri-2="https://zenodo.org/record/210972/files/figure.png" pageId="13" pageNumber="14">Figs 39–48</figureCitation>
)
</paragraph>
<paragraph id="060636BFFFFC8971FF2D1F6DA321FE14" blockId="13.[151,1436,257,434]" pageId="13" pageNumber="14">
<bibRefCitation id="62284B4EFFFC8971FF2D1F6DA310FEBE" author="McLachlan" box="[151,323,258,280]" pageId="13" pageNumber="14" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan 1872</bibRefCitation>
: 52−53, pl. I, figs 6, 7−7b (
<taxonomicName id="C1B94D3CFFFC8971FDD91F6CA11CFEBE" box="[611,847,259,280]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFFC8971FDD91F6CA11CFEBE" box="[611,847,259,280]" italics="true" pageId="13" pageNumber="14">Dictyopteryx compacta</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FCDC1F6DA1ACFEBE" author="Klapalek" box="[870,1023,258,280]" pageId="13" pageNumber="14" refString="Klapalek, F. (1912) Plecopteres I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Selys-Longchamps. Bruxelles, 4, 1 - 66." type="journal article" year="1912">Klapálek 1912</bibRefCitation>
: 15 (
<taxonomicName id="C1B94D3CFFFC8971FB881F6CA770FEBE" box="[1074,1315,259,280]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFFC8971FB881F6CA770FEBE" box="[1074,1315,259,280]" italics="true" pageId="13" pageNumber="14">Arcynopteryx compacta</emphasis>
</taxonomicName>
<taxonomicName id="C1B94D3CFFFC8971FA901F6CA7CFFEBE" box="[1322,1436,258,280]" pageId="13" pageNumber="14" rank="variety" variety="pusilla">
var. 
<emphasis id="34CDEAADFFFC8971FAED1F6DA7CFFEBE" box="[1367,1436,258,280]" italics="true" pageId="13" pageNumber="14">pusilla</emphasis>
</taxonomicName>
nov.); 
<bibRefCitation id="62284B4EFFFC8971FEBD1F4DA3CCFE91" author="Koponen" box="[263,415,290,311]" pageId="13" pageNumber="14" refString="Koponen, J. (1949) Neue oder wenig bekannte Plecoptera. Annales Entomologici Fennici. Helsinki, 15, 1 - 21." type="journal article" year="1949">Koponen 1949</bibRefCitation>
: 12−13, figs 10 А −С (
<taxonomicName id="C1B94D3CFFFC8971FD3A1F4DA119FE91" box="[640,842,289,311]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="34CDEAADFFFC8971FD3A1F4DA119FE91" box="[640,842,289,311]" italics="true" pageId="13" pageNumber="14">Arcynopteryx brevis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FCDA1F4EA19DFE91" author="Illies" box="[864,974,289,311]" pageId="13" pageNumber="14" refString="Illies, J. (1966) Katalog der rezenten Plecoptera. Das Tierreich, Berlin, 82, 632 s." type="journal article" year="1966">Illies 1966</bibRefCitation>
: 510 (sp. inquirenda); 
<bibRefCitation id="62284B4EFFFC8971FB0A1F4DA761FE91" author="Rauser" box="[1200,1330,289,311]" pageId="13" pageNumber="14" refString="Rauser, J. (1968) Plecoptera. Ergebnisse der zoologischen Forschungen von Dr. Z. Kazsab in der Mongolia. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 34, 398 s." type="journal article" year="1968">Raušer 1968</bibRefCitation>
: 352−354, figs 47−57 (
<taxonomicName id="C1B94D3CFFFC8971FEF91F2FA00AFEF0" box="[323,601,320,342]" class="Insecta" family="Perlodidae" genus="Arcynopterygoides" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vernalis">
<emphasis id="34CDEAADFFFC8971FEF91F2FA00AFEF0" box="[323,601,320,342]" italics="true" pageId="13" pageNumber="14">Arcynopterygoides vernalis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2FFE57D6FFFC8971FDDC1F2EA0F8FEF0" box="[614,683,321,342]" pageId="13" pageNumber="14">syn. n.</taxonomicNameLabel>
); 
<bibRefCitation id="62284B4EFFFC8971FD7A1F2FA124FEF3" author="Zwick" box="[704,887,320,342]" pageId="13" pageNumber="14" refString="Zwick, P., Levanidova, I. &amp; Zhiltzova, L. (1971) On the fauna of Plecoptera from the Soviet Far East. Revue d'Entomologie de I'URSS, 50, 849 - 869." type="journal article" year="1971">
Zwick 
<emphasis id="34CDEAADFFFC8971FCBD1F2EA168FEF0" box="[775,827,320,342]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
1971
</bibRefCitation>
: 852−853, figs 6−8 (
<taxonomicName id="C1B94D3CFFFC8971FBF61F2FA68BFEF0" box="[1100,1240,320,342]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="34CDEAADFFFC8971FBF61F2FA68BFEF0" box="[1100,1240,320,342]" italics="true" pageId="13" pageNumber="14">Skwala brevis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FB541F2FA73FFEF3" author="Zwick" box="[1262,1388,320,342]" pageId="13" pageNumber="14" refString="Zwick, P. (1973) Insecta: Plecoptera. Phylogenetisches System und Katalog. Das Tierreich. Walter de Gruyter, Berlin, 94, ixxxii + 465 pp." type="book" year="1973">Zwick 1973</bibRefCitation>
: 239 (
<taxonomicName id="C1B94D3CFFFC8971FF751F30A3C9FED3" box="[207,410,351,373]" class="Insecta" family="Perlodidae" genus="Arcynopteryx" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="34CDEAADFFFC8971FF751F30A3C9FED3" box="[207,410,351,373]" italics="true" pageId="13" pageNumber="14">Arcynopteryx brevis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FE0B1F31A0FBFED2" author="Stark" box="[433,680,350,372]" pageId="13" pageNumber="14" refString="Stark, B. &amp; Szczytko, W. (1981) Skwala brevis (Koponen) from Japan (Plecoptera: Perlodidae). Aquatic Insects, 3, 61 - 63." type="journal article" year="1981">Stark &amp; Szczytko, 1981</bibRefCitation>
: 61−63, figs 1−4 (
<taxonomicName id="C1B94D3CFFFC8971FCDF1F30A1A1FED3" box="[869,1010,351,373]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="34CDEAADFFFC8971FCDF1F30A1A1FED3" box="[869,1010,351,373]" italics="true" pageId="13" pageNumber="14">Skwala brevis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FBB21F31A6F5FED2" author="Zhiltzova" box="[1032,1190,350,372]" pageId="13" pageNumber="14" refString="Zhiltzova, L. (1982) New data on stonefly fauna (Plecoptera) of Sakhalin. Biology of the freshwater animals of the Far East. FESC AS USSR. Vladivostok, 115 - 124 (in Russian)." type="book chapter" year="1982">Zhiltzova 1982</bibRefCitation>
: 116 (
<taxonomicName id="C1B94D3CFFFC8971FB5F1F30A728FED3" box="[1253,1403,351,373]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="34CDEAADFFFC8971FB5F1F30A728FED3" box="[1253,1403,351,373]" italics="true" pageId="13" pageNumber="14">Skwala pusilla</emphasis>
</taxonomicName>
= 
<taxonomicName id="C1B94D3CFFFC8971FA331F30A357FE35" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="34CDEAADFFFC8971FA331F30A357FE35" italics="true" pageId="13" pageNumber="14">S. brevis</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FEA01F12A0DCFE35" author="Zhiltzova" box="[282,655,381,403]" pageId="13" pageNumber="14" refString="Zhiltzova, L. &amp; Zapekina-Dulkeit, Y. (1986) Order Plecoptera - stoneflies. In: P. A. Ler (Ed.), Key for Insects of the Soviet Far East. I. Apterygota, Palaeaodictyoptera, and Hemimetabola. Nauka Publishers, Leningrad, pp. 172 - 234 (in Russian)." type="book chapter" year="1986">Zhiltzova &amp; Zapekina-Dulkeit 1986</bibRefCitation>
: 184, figs 4−6 (
<taxonomicName id="C1B94D3CFFFC8971FC881F12A19BFE35" box="[818,968,381,403]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="34CDEAADFFFC8971FC881F12A19BFE35" box="[818,968,381,403]" italics="true" pageId="13" pageNumber="14">Skwala pusilla</emphasis>
</taxonomicName>
); 
<bibRefCitation id="62284B4EFFFC8971FC651F12A6A8FE34" author="Teslenko" box="[991,1275,381,403]" pageId="13" pageNumber="14" refString="Teslenko, V. &amp; Zhiltzova, L. (2009) Keys to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and larvae. Dalnauka, Vladivostok, 1 - 382 pp (in Russian)." type="book chapter" year="2009">Teslenko &amp; Zhiltzova 2009</bibRefCitation>
: 17, figs 49−53 (
<taxonomicName id="C1B94D3CFFFC8971FF751FF3A330FE14" box="[207,355,412,434]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="34CDEAADFFFC8971FF751FF3A330FE14" box="[207,355,412,434]" italics="true" pageId="13" pageNumber="14">Skwala pusilla</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFFC8971FF2D1FB1A080F9D0" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="060636BFFFFC8971FF2D1FB1A032FBFC" blockId="13.[151,1437,478,2014]" pageId="13" pageNumber="14">
<emphasis id="34CDEAADFFFC8971FF2D1FB1A342FE51" bold="true" box="[151,273,478,503]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Abdominal segments 1−2 are divided by a pleural fold. The submental gills are long and conical. The arms of the mesosternal ridge meet the anterior corners of the furcal pits, the transverse suture is absent. The stylet of the epiproct as a long, fine bristle is absent. The male of 
<taxonomicName id="C1B94D3CFFFC8971FC811C49A652FD99" box="[827,1025,550,575]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="compacta" status="comb. nov.">
<emphasis id="34CDEAADFFFC8971FC811C49A652FD99" box="[827,1025,550,575]" italics="true" pageId="13" pageNumber="14">Skwala compacta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FFE57D6FFFC8971FBB31C48A6D9FD98" box="[1033,1162,551,575]" pageId="13" pageNumber="14">
<emphasis id="34CDEAADFFFC8971FBB31C48A6D0FD99" bold="true" box="[1033,1155,551,575]" pageId="13" pageNumber="14">comb. nov</emphasis>
.
</taxonomicNameLabel>
can be distinguished by the shape of the hemitergal lobes, which are simple, cylindrical, wide, short, flat and slightly curved upward along the posterior margin. The hemitergal lobes are directed towards the middle transversely, and covered by small, stout setae (
<figureCitation id="9E822A3AFFFC8971FEA51CFDA3F5FD0C" box="[287,422,658,682]" captionStart-0="FIGURES 39 – 41" captionStart-1="FIGURES 42 – 45" captionStartId-0="14.[151,264,1692,1715]" captionStartId-1="15.[151,264,967,990]" captionTargetBox-0="[268,1297,543,1657]" captionTargetBox-1="[267,1322,287,936]" captionTargetId-0="figure@14.[264,1323,543,1668]" captionTargetId-1="figure@15.[264,1323,287,942]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." captionText-1="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri-0="https://zenodo.org/record/210970/files/figure.png" httpUri-1="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Figs 40−43</figureCitation>
). The lever arm is short, stout and slightly curved; the epiproct is short, blunt and spoon shaped, with a soft distal part and a finger-shaped top (
<figureCitation id="9E822A3AFFFC8971FCBF1CD9A10AFD68" box="[773,857,694,718]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
), the top is densely covered with thin, clear spines. The lateral stylets are attached laterally to the folds of the cowl only the pointed sclerotized tips are clearly pronounced (
<figureCitation id="9E822A3AFFFC8971FEBC1C91A308FCB0" box="[262,347,766,790]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
). The everted aedeagus is large and membranous, with a pair of distinct lateral rounded lobes at the dorsolateral margins; one large lobe rounded dorsally is present between the lateral lobes; one small knob is located apically; two broadly rounded swellings project posteriolaterally and slightly below the large lobe; tiny, clear, erect spinules densely cover the aedeagal surface ventrally and laterally; reddish spinules are grouped in a triangle on the dorsal surface of the large lobe basally and continue dorsomedially to the small knob (
<figureCitation id="9E822A3AFFFC8971FABC1DE1A70FFC00" box="[1286,1372,910,934]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Fig. 42</figureCitation>
). The female has a large subgenital plate, its posterior margin slightly rounded and sclerotized, with two shallow notches medially; the plate is covered with small, darkish setae (
<figureCitation id="9E822A3AFFFC8971FCB71DB9A132FC48" box="[781,865,982,1006]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
). The egg is spindle-shaped (
<figureCitation id="9E822A3AFFFC8971FB121DB9A6AFFC48" box="[1192,1276,982,1006]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
). The collar is stalked; the sides of the collar bear several sharp longitudinal carinae (
<figureCitation id="9E822A3AFFFC8971FC1D1D95A1A8FBB4" box="[935,1019,1018,1042]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="13" pageNumber="14">Fig. 47</figureCitation>
). The chorion is covered with closely packed hexagonal FCIs (
<figureCitation id="9E822A3AFFFC8971FE161A71A06BFB90" box="[428,568,1054,1078]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="13" pageNumber="14">Figs. 46−48</figureCitation>
); the FCI walls are thin, consisting of shallow furrows; the floors are flat with 4-8 small punctations medially (
<figureCitation id="9E822A3AFFFC8971FDBA1A2DA007FBFC" box="[512,596,1090,1114]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="13" pageNumber="14">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="060636BFFFFC8971FF731A09A080F9D0" blockId="13.[151,1437,478,2014]" pageId="13" pageNumber="14">
<emphasis id="34CDEAADFFFC8971FF731A09A325FBD9" bold="true" box="[201,374,1126,1151]" pageId="13" pageNumber="14">Adult habitus.</emphasis>
The head is brown, with a yellow spear-shaped spot in the interocellar area, not reaching the median ocellus and continued to the occiput medially, where the spot is slightly enlarged (
<figureCitation id="9E822A3AFFFC8971FB2F1AE5A6B9FB04" box="[1173,1258,1162,1186]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Fig. 39</figureCitation>
). A dark brown spot projects onto the clypeus in front of the brownish M-line. The tentorial pits are brown; the lateral margins of the clypeus are pale. The pair of tentorial pits in front of the lateral ocelli and the pair of small oval patches lateral to the lateral ocelli are brown. A posterolateral spot with brown callosities is weakly recognizable behind each compound eye. The antennae and palpi are brown. The pronotum is almost square or slightly longer in width than in length, brown with a yellow medial band and prominent dark brown rugosities (
<figureCitation id="9E822A3AFFFC8971FBD21B51A693FAF0" box="[1128,1216,1342,1366]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Fig. 39</figureCitation>
). The medial band expands in the last quarter of the length of the pronotum. The meso- and metanotum are dark brown. The anterior margin of the femur is dark brown. The tibia is light brown with a thin dark brown stripe basally. The wings of the male and female are slightly shortened and extend to the top of the abdomen. The RS of the forewing has three or more branches; the apex exhibits a random network of cells formed by the transverse veins and branches of the RS. The cerci are longer than the abdomen, ringed, and covered with long colorless hairs. Each cercal segment is pale basally and blackish distally. The apical cercal segments are completely blackish with the exception of the pale bases (
<figureCitation id="9E822A3AFFFC8971FF591855A36BF9F4" box="[227,312,1594,1618]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Fig. 39</figureCitation>
). The submental gills are long and conical. The arms of the mesosternal ridge meet the anterior corners of the furcal pits, a transverse suture is absent.
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFFC8973FF7318ECA716FF51" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="060636BFFFFC8972FF7318ECA138FE52" blockId="13.[151,1437,478,2014]" lastBlockId="14.[151,1436,151,500]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="34CDEAADFFFC8971FF7318ECA35EF93D" bold="true" box="[201,269,1667,1691]" pageId="13" pageNumber="14">Male.</emphasis>
Body length 
<quantity id="C1419B5AFFFC8971FE1418EDA004F93C" box="[430,599,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="13" pageNumber="14" unit="mm" value="13.0" valueMax="14.5" valueMin="11.5">11.5−14.5 mm</quantity>
, forewing 8.0−11.0 mm, wingspan 
<quantity id="C1419B5AFFFC8971FC4F18EDA6CDF93C" box="[1013,1182,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="2.035" metricValueMax="2.35" metricValueMin="1.72" pageId="13" pageNumber="14" unit="mm" value="20.35" valueMax="23.5" valueMin="17.2">17.2–23.5 mm</quantity>
. Abdominal tergum 9 exhibits a thin, transversal, membranous, brownish median band and one indistinct small pale spot anteromedially (
<figureCitation id="9E822A3AFFFC8971FF2518A5A2A8F944" box="[159,251,1738,1762]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Figs. 41</figureCitation>
&amp; 
<figureCitation id="9E822A3AFFFC8971FEA518A5A36CF944" box="[287,319,1738,1762]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">42</figureCitation>
). The posterior margin of tergum 9 bears a notch along 1/3 of the length of tergum 9 and two transversely elongated and rounded swellings, which are covered by small stout setae and fine colorless hairs posteriolaterally (
<figureCitation id="9E822A3AFFFC8971FEB9197DA304F88C" box="[259,343,1810,1834]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Fig. 41</figureCitation>
). Sternum 9 is scoop-shaped, extended backward and upward, pointed at the top (
<figureCitation id="9E822A3AFFFC8971FB4E197DA7DCF88C" box="[1268,1423,1810,1834]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="13" pageNumber="14">Figs 40 &amp; 41</figureCitation>
). Tergum 10 is divided into two hemiterga. The hemitergal lobes in dorsal view are simple, cylindrical, wide, short, flat and slightly curved upward along the posterior margin, directed towards the middle transversely, and covered by small, stout setae (
<figureCitation id="9E822A3AFFFC8971FE291911A049F830" box="[403,538,1918,1942]" captionStart-0="FIGURES 39 – 41" captionStart-1="FIGURES 42 – 45" captionStartId-0="14.[151,264,1692,1715]" captionStartId-1="15.[151,264,967,990]" captionTargetBox-0="[268,1297,543,1657]" captionTargetBox-1="[267,1322,287,936]" captionTargetId-0="figure@14.[264,1323,543,1668]" captionTargetId-1="figure@15.[264,1323,287,942]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." captionText-1="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri-0="https://zenodo.org/record/210970/files/figure.png" httpUri-1="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Figs 40−43</figureCitation>
). The cowl is membranous, folded, resembles a pouch between and under the hemitergal lobes, attached around the sclerites of the internal basal anchor. The cowl opens dorsally with a slit. The dorsolateral edges of the cowl are supported by flat, band-like sclerotized paragenital plates (
<figureCitation id="9E822A3AFFFC8971FB0219A9A75EF878" box="[1208,1293,1990,2014]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="13" pageNumber="14">Fig. 42</figureCitation>
). In a lateral view, the internal basal anchor is rounded anteriorly, the arrow-like sclerites are black and hidden under tergum 9 (
<figureCitation id="9E822A3AFFFF8972FF251ED3A2A6FF72" box="[159,245,188,212]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
). The lever arm is short, stout, slightly curved, hollowed and connected to the epiproct distally (
<figureCitation id="9E822A3AFFFF8972FA801ED3A7DCFF72" box="[1338,1423,188,212]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 44</figureCitation>
). The epiproct is short, blunt, spoon shaped, sclerotized basally and ventrally; its distal part is soft (
<figureCitation id="9E822A3AFFFF8972FB641EB0A761FF51" box="[1246,1330,223,247]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 44</figureCitation>
). The top of the distal part is finger-shaped, densely covered with thin, clear spines. Lateral stylets are attached to the folds of the cowl laterally; their acute sclerotized tops clearly pronounced (
<figureCitation id="9E822A3AFFFF8972FC1B1F48A1AAFE99" box="[929,1017,295,319]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 44</figureCitation>
). The everted aedeagus is large and membranous, in a dorsal view with a pair of finger-like lobes at the dorsolateral margins; one large dorsally rounded lobe is set between the lateral lobes; one small rounded knob located apically; two broadly rounded swellings project posteriolaterally and slightly below the large lobe; tiny, clear, erect spinules densely cover the aedeagal surface ventrally and laterally; reddish spinules are grouped in a triangle on the dorsal surface of the large lobe basally and continued dorsomedially to the small knob (
<figureCitation id="9E822A3AFFFF8972FCB01FB3A10DFE52" box="[778,862,476,500]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
).
</paragraph>
<caption id="52C66637FFFF8972FF2D18F3A613F977" httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="14" pageNumber="15" targetBox="[268,1297,543,1657]" targetPageId="14">
<paragraph id="060636BFFFFF8972FF2D18F3A613F977" blockId="14.[151,1437,1692,1746]" pageId="14" pageNumber="15">
<emphasis id="34CDEAADFFFF8972FF2D18F3A306F914" bold="true" box="[151,341,1692,1715]" pageId="14" pageNumber="15">FIGURES 39–41.</emphasis>
<taxonomicName id="C1B94D3CFFFF8972FEE718F2A043F915" box="[349,528,1693,1715]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFFF8972FEE718F2A043F915" box="[349,528,1693,1715]" italics="true" pageId="14" pageNumber="15">Skwala compacta</emphasis>
</taxonomicName>
: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova).
</paragraph>
</caption>
<paragraph id="060636BFFFFF8972FF731890A771F800" blockId="14.[151,1436,1790,2030]" pageId="14" pageNumber="15">
<emphasis id="34CDEAADFFFF8972FF731890A373F8B1" bold="true" box="[201,288,1791,1815]" pageId="14" pageNumber="15">Female</emphasis>
is larger than male and similar to male in shape and color. Body length 12.0− 
<quantity id="C1419B5AFFFF8972FB071891A774F8B0" box="[1213,1319,1790,1814]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="14" pageNumber="15" unit="mm" value="16.2">16.2 mm</quantity>
, forewing 14.0− 
<quantity id="C1419B5AFFFF8972FF62194DA36EF89C" box="[216,317,1826,1850]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="15" unit="mm" value="17.5">17.5 mm</quantity>
, wingspan 30.0−37.0 mm. Sternum 8 is pale medially. The subgenital plate is large, extends laterally from the sides of sternum 8, and reaches almost half the length of sternum 9; the posterior margin is slightly rounded and sclerotized, with two shallow notches medially (
<figureCitation id="9E822A3AFFFF8972FCF71905A1F1F824" box="[845,930,1898,1922]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 45</figureCitation>
). The subgenital plate is covered with small, darkish setae. Sterna 9, 10 are pale, sternum 9 with two brownish rounded spots mesolaterally (
<figureCitation id="9E822A3AFFFF8972FB7819E1A745F800" box="[1218,1302,1934,1958]" captionStart="FIGURES 42 – 45" captionStartId="15.[151,264,967,990]" captionTargetBox="[267,1322,287,936]" captionTargetId="figure@15.[264,1323,287,942]" captionTargetPageId="15" captionText="FIGURES 42 – 45. Skwala compacta: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0." httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="14" pageNumber="15">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="060636BFFFFF8973FF7319DCA716FF51" blockId="14.[151,1436,1790,2030]" lastBlockId="15.[151,1436,151,247]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="34CDEAADFFFF8972FF7319DCA2ABF86D" bold="true" box="[201,248,1971,1995]" pageId="14" pageNumber="15">Egg</emphasis>
is spindle-shaped (
<figureCitation id="9E822A3AFFFF8972FE7519DDA073F86C" box="[463,544,1970,1994]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="14" pageNumber="15">Fig. 46</figureCitation>
) and circular in cross-section, 315–350 x 229–250 µm. The collar is stalked, its rim irregularly incised; the sides of collar bear several sharp longitudinal carinae; the shoulder is low (
<figureCitation id="9E822A3AFFFF8972FAB019B9A70EF848" box="[1290,1373,2006,2030]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="14" pageNumber="15">Fig. 47</figureCitation>
). The chorion is covered with closely packed hexagonal FCIs; the FCI walls are thin, consisting of shallow furrows; the floors flat with 4-8 small punctations medially (
<figureCitation id="9E822A3AFFFE8973FD0B1ED3A155FF72" box="[689,774,188,212]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="15" pageNumber="16">Fig. 48</figureCitation>
). The row of micropyles is subequatorial; their orifices are small with indistinct lips, some set on micropylar mounds and surrounded by pentagonal rosettes (
<figureCitation id="9E822A3AFFFE8973FB5E1EB0A76BFF51" box="[1252,1336,223,247]" captionStart="FIGURES 46 – 48" captionStartId="15.[151,264,1980,2003]" captionTargetBox="[264,1322,1088,1953]" captionTargetId="figure@15.[264,1323,1088,1959]" captionTargetPageId="15" captionText="FIGURES 46 – 48. Skwala compacta, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view." httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="15" pageNumber="16">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="52C66637FFFE8973FF2D1DA8A30EFBBC" httpUri="https://zenodo.org/record/210971/files/figure.png" pageId="15" pageNumber="16" targetBox="[267,1322,287,936]" targetPageId="15">
<paragraph id="060636BFFFFE8973FF2D1DA8A30EFBBC" blockId="15.[151,1437,967,1051]" pageId="15" pageNumber="16">
<emphasis id="34CDEAADFFFE8973FF2D1DA8A306FC7A" bold="true" box="[151,341,967,990]" pageId="15" pageNumber="16">FIGURES 42–45.</emphasis>
<taxonomicName id="C1B94D3CFFFE8973FEE61DA8A05DFC7B" box="[348,526,967,989]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFFE8973FEE61DA8A05DFC7B" box="[348,526,967,989]" italics="true" pageId="15" pageNumber="16">Skwala compacta</emphasis>
</taxonomicName>
: 42. Male abdominal tip and aedeagus, dorsal. 43. Male left hemitergal lobe, dorsal. 44. Cowl: sclerites of the internal basal anchor, lever arm, epiproct, lateral stylet, lateral. 45. Female abdominal tip, ventral. Scale (mm): 43, 44 = 1.0.
</paragraph>
</caption>
<caption id="52C66637FFFE8973FF2D19D3A01EF857" httpUri="https://zenodo.org/record/210972/files/figure.png" pageId="15" pageNumber="16" targetBox="[264,1322,1088,1953]" targetPageId="15">
<paragraph id="060636BFFFFE8973FF2D19D3A01EF857" blockId="15.[151,1436,1980,2034]" pageId="15" pageNumber="16">
<emphasis id="34CDEAADFFFE8973FF2D19D3A304F874" bold="true" box="[151,343,1980,2003]" pageId="15" pageNumber="16">FIGURES 46–48.</emphasis>
<taxonomicName id="C1B94D3CFFFE8973FEE519D2A047F875" box="[351,532,1981,2003]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFFE8973FEE519D2A047F875" box="[351,532,1981,2003]" italics="true" pageId="15" pageNumber="16">Skwala compacta</emphasis>
</taxonomicName>
, egg: 46. Habitus lateral. 47. Collar, lateral. 48. Chorion structure, micropylar orifice, mound, and pentagon rosettes, dorsal view.
</paragraph>
</caption>
<subSubSection id="4EA36534FFE1896CFF7D1EF8A67FFE99" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="060636BFFFE1896CFF7D1EF8A67FFE99" blockId="16.[151,1436,151,573]" pageId="16" pageNumber="17">
<emphasis id="34CDEAADFFE1896CFF7D1EF8A3E2FF16" bold="true" box="[199,433,151,176]" pageId="16" pageNumber="17">Material examined.</emphasis>
<typeStatus id="D902881DFFE1896CFE001EF7A07EFF16" box="[442,557,152,176]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
, male (pinned), 
<taxonomicName id="C1B94D3CFFE1896CFD5C1EF6A6F8FF09" authority="McLachlan" authorityName="McLachlan" box="[742,1195,151,176]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFD5C1EF6A125FF16" box="[742,886,153,176]" italics="true" pageId="16" pageNumber="17">D. compacta</emphasis>
McLachlan (
<emphasis id="34CDEAADFFE1896CFBB51EF6A6F3FF16" box="[1039,1184,153,176]" italics="true" pageId="16" pageNumber="17">Dictyopteryx</emphasis>
)
</taxonomicName>
, [McL. label] / Sibir. orient. (Maa(c)k) / 
<taxonomicName id="C1B94D3CFFE1896CFED71ED2A389FF72" box="[365,474,189,212]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFED71ED2A389FF72" box="[365,474,189,212]" italics="true" pageId="16" pageNumber="17">compacta</emphasis>
</taxonomicName>
McL. / 
<taxonomicName id="C1B94D3CFFE1896CFD8E1ED2A16AFF72" box="[564,825,189,212]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFD8E1ED2A16AFF72" box="[564,825,189,212]" italics="true" pageId="16" pageNumber="17">Dictyopteryx compacta</emphasis>
</taxonomicName>
McL. The type-series in 
<collectionCode id="60A8AE7AFFE1896CFBEE1ED2A6FBFF72" box="[1108,1192,189,212]" pageId="16" pageNumber="17">BMNH</collectionCode>
includes three 
<typeStatus id="D902881DFFE1896CFAE81ED2A35AFF51" pageId="16" pageNumber="17" type="paralectotype">paralectotypes</typeStatus>
(
<bibRefCitation id="62284B4EFFE1896CFEAD1EB0A397FF51" author="Kimmins" box="[279,452,223,247]" pageId="16" pageNumber="17" refString="Kimmins, D. (1970) A list of the type-specimens of Plecoptera and Megaloptera in the British Museum (Natural History). Bulletin of the British Museum (Natural History) Entomology, 24, 335 - 361." type="journal article" year="1970">Kimmins 1970</bibRefCitation>
). Siberia, Eastern Sayan Mountains, Krasnoyarskyi Region, 
<specimenCount id="10BFFD36FFE1896CFBDD1EB0A69DFF51" box="[1127,1230,223,247]" pageId="16" pageNumber="17" type="male">12 males</specimenCount>
, 
<specimenCount id="10BFFD36FFE1896CFB631EB0A71BFF51" box="[1241,1352,223,247]" pageId="16" pageNumber="17" type="female">6 females</specimenCount>
, Bazaikha River, Yenisey R. Basin, 
<date id="7207107FFFE1896CFE581F6BA030FEBA" box="[482,611,260,284]" pageId="16" pageNumber="17" value="1959-05-20">20.05.1959</date>
, coll. I. Levanidova. Far East, Prymorskyi Region, 
<specimenCount id="10BFFD36FFE1896CFB1D1F6BA758FEBA" box="[1191,1291,260,284]" pageId="16" pageNumber="17" type="male">18 males</specimenCount>
, 
<specimenCount id="10BFFD36FFE1896CFAA01F6BA7CBFEBA" box="[1306,1432,260,284]" pageId="16" pageNumber="17" type="female">10 females</specimenCount>
, Partisanskaya River, near Frolovka settelment, 
<date id="7207107FFFE1896CFD1D1F48A17BFE99" box="[679,808,295,319]" pageId="16" pageNumber="17" value="1989-04-29">29.04.1989</date>
, coll. E. Makarchenko.
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFE1896CFF731F24A104FEC2" box="[201,855,331,356]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="060636BFFFE1896CFF731F24A104FEC2" blockId="16.[151,1436,151,573]" box="[201,855,331,356]" pageId="16" pageNumber="17">
<emphasis id="34CDEAADFFE1896CFF731F24A331FEC2" bold="true" box="[201,354,331,356]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="7EAE762FFFE1896CFED31F23A3E7FEC2" box="[361,436,332,356]" name="Russia" pageId="16" pageNumber="17">Russia</collectingCountry>
(Siberia, Far East), 
<collectingCountry id="7EAE762FFFE1896CFD2F1F23A156FEC2" box="[661,773,332,356]" name="Mongolia" pageId="16" pageNumber="17">Mongolia</collectingCountry>
, 
<collectingCountry id="7EAE762FFFE1896CFCAA1F22A100FEC2" box="[784,851,333,356]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4EA36534FFE1896CFF731F1FA68AFD9A" pageId="16" pageNumber="17" type="discussion">
<paragraph id="060636BFFFE1896CFF731F1FA68AFD9A" blockId="16.[151,1436,151,573]" pageId="16" pageNumber="17">
<emphasis id="34CDEAADFFE1896CFF731F1FA36EFE2E" bold="true" box="[201,317,368,392]" pageId="16" pageNumber="17">Remarks.</emphasis>
The 
<typeStatus id="D902881DFFE1896CFE381F1FA3A1FE2E" box="[386,498,368,392]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="C1B94D3CFFE1896CFD9E1F1EA0EAFE2E" box="[548,697,369,392]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFD9E1F1EA0EAFE2E" box="[548,697,369,392]" italics="true" pageId="16" pageNumber="17">D. compacta</emphasis>
</taxonomicName>
is in close agreement with the description of this species by 
<bibRefCitation id="62284B4EFFE1896CFF2D1FFBA33FFE0A" author="McLachlan" box="[151,364,404,428]" pageId="16" pageNumber="17" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan (1872)</bibRefCitation>
. The 
<typeStatus id="D902881DFFE1896CFE111FFAA388FE0B" box="[427,475,405,429]" pageId="16" pageNumber="17">type</typeStatus>
of 
<taxonomicName id="C1B94D3CFFE1896CFDB81FFAA0C2FE0A" box="[514,657,405,428]" class="Insecta" family="Perlodidae" genus="Dictyopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFDB81FFAA0C2FE0A" box="[514,657,405,428]" italics="true" pageId="16" pageNumber="17">D. compacta</emphasis>
</taxonomicName>
is in poor condition, the abdomen fragmented during cleaning, making a study of the internal structure of the epiproct impossible. However, the shape of the hemitergal lobes, lateral stylets present, and the short, blunt, spoon shaped epiproct (
<figureCitation id="9E822A3AFFE1896CFC8F1FB3A1D4FE52" box="[821,903,476,500]" captionStart="FIGURES 39 – 41" captionStartId="14.[151,264,1692,1715]" captionTargetBox="[268,1297,543,1657]" captionTargetId="figure@14.[264,1323,543,1668]" captionTargetPageId="14" captionText="FIGURES 39 – 41. Skwala compacta: 39. Habitus (Photograph by M. Tiunov). 40. Type, male abdominal tip, dorsal, pinned. (Photograph by D. Goodger). 41. Same, recent material, dorsal. (Photograph by T. Tiunova)." httpUri="https://zenodo.org/record/210970/files/figure.png" pageId="16" pageNumber="17">Fig. 40</figureCitation>
) suggests that the species should be transferred to 
<taxonomicName id="C1B94D3CFFE1896CFF0E1C6FA3CBFDB1" authority="Ricker 1943" authorityName="Ricker" authorityYear="1943" box="[180,408,511,536]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="34CDEAADFFE1896CFF0E1C6FA356FDBE" box="[180,261,512,536]" italics="true" pageId="16" pageNumber="17">Skwala</emphasis>
<bibRefCitation id="62284B4EFFE1896CFEB71F90A3CBFDB1" author="Ricker" box="[269,408,511,535]" pageId="16" pageNumber="17" refString="Ricker, W. (1943) Stoneflies of Southwestern British Columbia. Indiana University Publications, Science Series 12. Indiana University Press, Bloomington, 145 pp." type="book" year="1943">Ricker 1943</bibRefCitation>
</taxonomicName>
with the valid name 
<taxonomicName id="C1B94D3CFFE1896CFD321C6FA1BAFDB1" authority="McLachlan" authorityName="McLachlan" box="[648,1001,511,536]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta" status="comb. nov.">
<emphasis id="34CDEAADFFE1896CFD321C6FA11DFDBE" box="[648,846,512,536]" italics="true" pageId="16" pageNumber="17">Skwala compacta</emphasis>
(McLachlan)
</taxonomicName>
<taxonomicNameLabel id="2FFE57D6FFE1896CFC4A1C6FA622FDBE" box="[1008,1137,512,536]" pageId="16" pageNumber="17">
<emphasis id="34CDEAADFFE1896CFC4A1C6FA622FDBE" bold="true" box="[1008,1137,512,536]" pageId="16" pageNumber="17">comb. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="C1B94D3CFFE1896CFBC31C6FA28BFD9A" authority="Klapalek, 1912" authorityName="Klapalek" authorityYear="1912" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="34CDEAADFFE1896CFBC31C6FA74DFDBE" box="[1145,1310,512,536]" italics="true" pageId="16" pageNumber="17">Skwala pusilla</emphasis>
(
<bibRefCitation id="62284B4EFFE1896CFA941F90A283FD9A" author="Klapalek" pageId="16" pageNumber="17" refString="Klapalek, F. (1912) Plecopteres I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Selys-Longchamps. Bruxelles, 4, 1 - 66." type="journal article" year="1912">Klapálek, 1912</bibRefCitation>
)
</taxonomicName>
is therefore a subjective junior synonym of 
<taxonomicName id="C1B94D3CFFE1896CFD7D1C4BA639FD9A" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" box="[711,1130,548,572]" class="Insecta" family="Perlodidae" genus="Skwala" kingdom="Animalia" order="Plecoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="34CDEAADFFE1896CFD7D1C4BA1D8FD9A" box="[711,907,548,572]" italics="true" pageId="16" pageNumber="17">Skwala compacta</emphasis>
(
<bibRefCitation id="62284B4EFFE1896CFC201C4BA631FD9A" author="McLachlan" box="[922,1122,548,572]" pageId="16" pageNumber="17" refString="McLachlan, R. (1872) Perlides. In: Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps &amp; McLachlan. Annales de la Societe entomologique de Belgique (Bruxelles), 15, 47 - 55." type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
<emphasis id="34CDEAADFFE1896CFBCB1C4BA68AFD9A" bold="true" box="[1137,1241,548,572]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="2FFE57D6FFE1896CFBCB1C4BA68AFD9A" box="[1137,1241,548,572]" pageId="16" pageNumber="17">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>