299 lines
29 KiB
XML
299 lines
29 KiB
XML
<document id="41E5C0FD625834459EA4CB8240119E4A" ID-DOI="10.1649/0010-065X-68.4.663" ID-Zenodo-Dep="5369368" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630263369791" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2014" docId="4A5987B5315C1E33FABACB4EE0058433" docLanguage="en" docName="0010-065X-68.4.663.pdf" docOrigin="The Coleopterists Bulletin 68 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-68.4.663" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Neoryctes similis Ratcliffe 2014, new species" docType="treatment" docVersion="3" lastPageNumber="680" masterDocId="B660FFCD31541E3AF861C828E47C820E" masterDocTitle="A New Genus and Species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other New Species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a Revised Key to the Genera of New World Pentodontini" masterLastPageNumber="680" masterPageNumber="663" pageNumber="663" updateTime="1699203105727" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="B275D021B6B0E3BCED73C7F5F2A4A21C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A31AC55E0892A86095EF6CA95FFF3875">
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<mods:title id="CA08D37913611F35408B69C8C4E063BC">A New Genus and Species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other New Species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a Revised Key to the Genera of New World Pentodontini</mods:title>
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</mods:titleInfo>
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<mods:name id="D8F3ACAF968841890EBE74366B591C50" type="personal">
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<mods:role id="006EE63113850F8F1065B0E7FD2B3863">
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<mods:roleTerm id="46E0A562DA5454DA3EDA1AD1011B6A7C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="03AC76A52A2ED4E89BEA90925FDA0713">Ratcliffe, Brett C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A4237A0A2402F44DFBF2071AF87F2BDC">text</mods:typeOfResource>
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<mods:titleInfo id="12E3BE7D97AA14621C4579DD5FDE6BF2">
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<mods:title id="C878BC4C8EABE17DFE96B70C745E4BB0">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="6859DA8468B0B4F285952C074413E4A2">
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<mods:date id="075222BCD1D4CA6501E0190194070F68">2014</mods:date>
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<mods:detail id="130C5B0FB3401E262775D76145435C54" type="pubDate">
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<mods:number id="85597605865D59CE2CF4298134943D4B">2014-12-31</mods:number>
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</mods:detail>
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<mods:detail id="4C2432E9E90082875DE3A30F3E26177F" type="volume">
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<mods:number id="D30970F169265DBFA28A218FFA237BC0">68</mods:number>
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</mods:detail>
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<mods:detail id="80CDD3EF05EE9744793FECD382BCFE98" type="issue">
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<mods:number id="482BBD7E12FC521DBE467F88774C6B95">4</mods:number>
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</mods:detail>
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<mods:extent id="20B449D30276F38A689028137B77595D" unit="page">
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<mods:start id="59BEF3BB192CA5FD1305AA69A4707666">663</mods:start>
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<mods:end id="3E94BF5937BF480CC320CBA1A1DC4090">680</mods:end>
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<mods:location id="706D4FAFB2C43997DF05385CF2EA3BDD">
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<mods:url id="AFD7171CAB2BBFAAAE6987502398270D">http://dx.doi.org/10.1649/0010-065x-68.4.663</mods:url>
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</mods:location>
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<mods:classification id="8C9BE73C778D85E112DB318AE1441EE3">journal article</mods:classification>
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<mods:identifier id="984D80DE76031A3F5D593A3EB1031B4A" type="DOI">10.1649/0010-065X-68.4.663</mods:identifier>
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<mods:identifier id="4502690D0DB81ED1483EE82D3551B3AD" type="Zenodo-Dep">5369368</mods:identifier>
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</mods:mods>
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<treatment id="4A5987B5315C1E33FABACB4EE0058433" LSID="urn:lsid:plazi:treatment:4A5987B5315C1E33FABACB4EE0058433" httpUri="http://treatment.plazi.org/id/4A5987B5315C1E33FABACB4EE0058433" lastPageId="9" pageId="8">
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<subSubSection id="8AEA6528315C1E32FABACB4EE79E8198" pageId="8" type="nomenclature">
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<paragraph id="C24F36A3315C1E32FABACB4EE79E8198" blockId="8.[731,1125,870,918]" pageId="8">
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<emphasis id="F084EAB1315C1E32FABACB4EE0198175" bold="true" box="[731,1125,870,891]" pageId="8">
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<taxonomicName id="05F04D20315C1E32FABACB4EE79F8175" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2014" box="[731,995,870,891]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
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<emphasis id="F084EAB1315C1E32FABACB4EE7FE8175" bold="true" box="[731,898,870,891]" italics="true" pageId="8">Neoryctes similis</emphasis>
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Ratcliffe
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="EBB757CA315C1E32FB91CB4EE0198175" box="[1008,1125,870,891]" pageId="8" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
(
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||
<figureCitation id="5ACB2A26315C1E32FB07CBA9E7A78198" box="[870,987,896,918]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="8.[175,215,1710,1729]" captionStartId-1="9.[197,231,945,964]" captionTargetBox-0="[170,1166,1133,1688]" captionTargetBox-1="[173,659,193,922]" captionTargetId-0="figure-355@8.[151,1172,1121,1698]" captionTargetId-1="figure-592@9.[172,659,192,922]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figs. 13–15. Neoryctes similis, holotype. 13) Habitus; 14) Parameres, caudal view; 15) Parameres, lateral view." captionText-1="Fig. 16. Isla Isabela, Galápagos Islands. Image courtesy of Frans Lanting/Lanting.com." figureDoi-0="http://doi.org/10.5281/zenodo.5369384" figureDoi-1="http://doi.org/10.5281/zenodo.5369386" httpUri-0="https://zenodo.org/record/5369384/files/figure.png" httpUri-1="https://zenodo.org/record/5369386/files/figure.png" pageId="8">Figs. 13–16</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="8AEA6528315C1E33FAA4CB83E7FE83EF" lastPageId="9" pageId="8" type="materials_examined">
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||
<paragraph id="C24F36A3315C1E33FAA4CB83E507864A" blockId="8.[685,1172,939,1093]" lastBlockId="9.[173,660,1044,1729]" lastPageId="9" pageId="8">
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<emphasis id="F084EAB1315C1E32FAA4CB83E72181CE" bold="true" box="[709,861,939,960]" pageId="8">Type Material.</emphasis>
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<materialsCitation id="72983CFE315C1E32FB0BCB83E7698625" collectorName="F. X. Williams" location="Cawley Mt." pageId="8" specimenCount="2" stateProvince="Galapagos" typeStatus="holotype">
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<typeStatus id="1D4B8801315C1E32FB0BCB83E7C381CE" box="[874,959,939,960]" pageId="8" type="holotype">Holotype</typeStatus>
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male labeled: “
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<location id="C72F6078315C1E32FC2CCB83E6AD81D5" LSID="urn:lsid:plazi:treatment:4A5987B5315C1E33FABACB4EE0058433:C72F6078315C1E32FC2CCB83E6AD81D5" name="Cawley Mt." pageId="8" stateProvince="Galapagos">Cawley Mt.</location>
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/Alt. Vii-06/Albermarle I/
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<collectingRegion id="0034F841315C1E32FBB2CBEEE04681D5" box="[979,1082,966,987]" country="Ecuador" name="Galapagos" pageId="8">Galapagos</collectingRegion>
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Is//Coll. by/
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<collectorName id="6F055375315C1E32FAAECBC8E71681FB" box="[719,874,992,1013]" pageId="8">F. X. Williams</collectorName>
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||
” and with my red
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<typeStatus id="1D4B8801315C1E32FC5BCBC8E0E881FB" box="[1082,1172,992,1013]" pageId="8" type="holotype">holotype</typeStatus>
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||
label. Right wing mounted on small card beneath specimen
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</materialsCitation>
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.
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<materialsCitation id="72983CFE315C1E33FB46CC3EE50B864A" collectionCode="CA" collectorName="F. X. Williams" lastPageId="9" location="Academy of Sciences" municipality="Types" pageId="8" specimenCount="1" stateProvince="California" typeStatus="allotype">
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<typeStatus id="1D4B8801315C1E32FB46CC3EE7F88625" box="[807,900,1046,1067]" pageId="8" type="allotype">Allotype</typeStatus>
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female with same data and wing mounted beneath specimen.
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<collectingMunicipality id="222BACD9315C1E32FC36CC18E0EF864B" box="[1111,1171,1072,1093]" pageId="8">Types</collectingMunicipality>
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deposited at the
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<collectingRegion id="0034F841315D1E33F934CC3CE5CF8627" box="[341,435,1044,1065]" country="United States of America" name="California" pageId="9">California</collectingRegion>
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<location id="C72F6078315D1E33F9DFCC3CE6EF8627" LSID="urn:lsid:plazi:treatment:4A5987B5315C1E33FABACB4EE0058433:C72F6078315D1E33F9DFCC3CE6EF8627" box="[446,659,1044,1065]" municipality="Types" name="Academy of Sciences" pageId="9" stateProvince="California">Academy of Sciences</location>
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(San Francisco,
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<collectionCode id="A4E1AE66315D1E33F92DCC07E50C864A" box="[332,368,1071,1092]" country="USA" name="Chicago Academy of Sciences" pageId="9" type="Museum">CA</collectionCode>
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)
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</materialsCitation>
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||
.
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</paragraph>
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<caption id="968F662B315C1E32F8CECE86E0F184CF" ID-DOI="http://doi.org/10.5281/zenodo.5369384" ID-Zenodo-Dep="5369384" box="[175,1165,1710,1729]" httpUri="https://zenodo.org/record/5369384/files/figure.png" pageId="8" startId="8.[175,215,1710,1729]" targetBox="[170,1166,1133,1688]" targetPageId="8">
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||
<paragraph id="C24F36A3315C1E32F8CECE86E0F184CF" blockId="8.[175,1165,1710,1729]" box="[175,1165,1710,1729]" pageId="8">
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<emphasis id="F084EAB1315C1E32F8CECE86E55C84CF" bold="true" box="[175,288,1710,1729]" pageId="8">Figs. 13–15.</emphasis>
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<taxonomicName id="05F04D20315C1E32F957CE86E5B484CE" authorityName="Ratcliffe" authorityYear="2014" box="[310,456,1710,1728]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="similis">
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<emphasis id="F084EAB1315C1E32F957CE86E5B484CE" box="[310,456,1710,1728]" italics="true" pageId="8">Neoryctes similis</emphasis>
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</taxonomicName>
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||
, holotype.
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<emphasis id="F084EAB1315C1E32FA4BCE86E63584CF" bold="true" box="[554,585,1710,1729]" pageId="8">13)</emphasis>
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Habitus;
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<emphasis id="F084EAB1315C1E32FAFFCE86E6C084CF" bold="true" box="[670,700,1710,1729]" pageId="8">14)</emphasis>
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Parameres, caudal view;
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<emphasis id="F084EAB1315C1E32FBFBCE86E7C584CF" bold="true" box="[922,953,1710,1729]" pageId="8">15)</emphasis>
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Parameres, lateral view.
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</paragraph>
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</caption>
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<caption id="968F662B315D1E33F8A4CB99E67A81D2" ID-DOI="http://doi.org/10.5281/zenodo.5369386" ID-Zenodo-Dep="5369386" httpUri="https://zenodo.org/record/5369386/files/figure.png" pageId="9" startId="9.[197,231,945,964]" targetBox="[173,659,193,922]" targetPageId="9">
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||
<paragraph id="C24F36A3315D1E33F8A4CB99E67A81D2" blockId="9.[173,659,945,988]" pageId="9">
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||
<emphasis id="F084EAB1315D1E33F8A4CB99E56E81CA" bold="true" box="[197,274,945,964]" pageId="9">Fig. 16.</emphasis>
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Isla Isabela, Galápagos Islands. Image courtesy of Frans Lanting/Lanting.com.
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||
</paragraph>
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||
</caption>
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<paragraph id="C24F36A3315D1E33F8A4CC61E7918352" blockId="9.[173,660,1044,1729]" lastBlockId="9.[707,1194,194,1597]" pageId="9">
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||
<emphasis id="F084EAB1315D1E33F8A4CC61E5598650" bold="true" box="[197,293,1097,1118]" pageId="9">
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||
<typeStatus id="1D4B8801315D1E33F8A4CC61E55D8650" box="[197,289,1097,1118]" pageId="9" type="holotype">Holotype</typeStatus>
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||
.
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</emphasis>
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Male. Length 16.0 mm; width
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<quantity id="05089B46315D1E33FA26CC61E6EC8650" box="[583,656,1097,1118]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" unit="mm" value="8.7">8.7 mm</quantity>
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. Color reddish brown.
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<emphasis id="F084EAB1315D1E33F9F2CC4CE5AE8677" bold="true" box="[403,466,1124,1145]" pageId="9">Head:</emphasis>
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Frons completely, coarsely rugopunctate, punctures large. Frontoclypeal ridge distinctly elevated, arcuate in anterior view. Clypeus subtriangular, apex weakly bidentate; surface rugopunctate, punctures moderate in size. Ocular canthus lacking acute lateral process but with small, distinct lobe at base. Mandibles large, exposed, arcuate on external edge. Interocular width equals 4.5 transverse eye diameters. Antenna 9-segmented, club subequal in length to antennomeres 2–6.
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<emphasis id="F084EAB1315D1E33F8BFCD45E537878C" bold="true" box="[222,331,1389,1410]" pageId="9">Pronotum:</emphasis>
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Surface convex, lacking fovea or tubercles, punctures moderately large, moderate in density, becoming slightly sparser on basal third. Base with complete marginal bead.
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<emphasis id="F084EAB1315D1E33FA77CD95E61C87DC" bold="true" box="[534,608,1469,1490]" pageId="9">Elytra:</emphasis>
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Surface with punctures moderate in size and density, shallow, glabrous, some in distinct rows. Apices bluntly rounded, appearing foreshortened, elytra together as wide as long. Hind wings
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<quantity id="05089B46315D1E33FA75CE0FE6218432" box="[532,605,1575,1596]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" unit="mm" value="8.7">8.7 mm</quantity>
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long.
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<emphasis id="F084EAB1315D1E33F8CCCE6AE5698459" bold="true" box="[173,277,1602,1623]" pageId="9">Pygidium:</emphasis>
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||
Surface with minute, sparse punctures, shiny. In lateral view, surface strongly convex.
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<emphasis id="F084EAB1315D1E33F8CCCE5FE49A8482" bold="true" box="[173,230,1655,1676]" pageId="9">Legs:</emphasis>
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Protibia quadridentate, basal tooth nearly obsolete. Metatibia at apex slightly expanded, subtruncate, with about 20 short spinules (some absent).
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<emphasis id="F084EAB1315D1E33FB77C8EAE72282D9" bold="true" box="[790,862,194,215]" pageId="9">Venter:</emphasis>
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Prosternal process short, triangular, laminate. long, subrectangular, thick, slightly recurving, apex truncate.
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<emphasis id="F084EAB1315D1E33FBAFC8DFE03A8302" bold="true" box="[974,1094,247,268]" pageId="9">Parameres:</emphasis>
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||
Together subrectangular, apices broadly expanded, median edge of each paramere just below middle with prominent tooth (
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<figureCitation id="5ACB2A26315D1E33FB0AC96FE79D8352" box="[875,993,326,348]" captionStart="Figs" captionStartId="8.[175,215,1710,1729]" captionTargetBox="[170,1166,1133,1688]" captionTargetId="figure-355@8.[151,1172,1121,1698]" captionTargetPageId="8" captionText="Figs. 13–15. Neoryctes similis, holotype. 13) Habitus; 14) Parameres, caudal view; 15) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5369384" httpUri="https://zenodo.org/record/5369384/files/figure.png" pageId="9">Figs. 14–15</figureCitation>
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||
).
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</paragraph>
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||
<paragraph id="C24F36A3315D1E33FABAC949E7FE83EF" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABAC949E7458378" bold="true" box="[731,825,353,374]" pageId="9">
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||
<typeStatus id="1D4B8801315D1E33FABAC949E7498378" box="[731,821,353,374]" pageId="9" type="allotype">Allotype</typeStatus>
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||
.
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</emphasis>
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Female. Length
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||
<quantity id="05089B46315D1E33FB9BC949E0228378" box="[1018,1118,353,374]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="9" unit="mm" value="16.8">16.8 mm</quantity>
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||
; width
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||
<quantity id="05089B46315D1E33FAA2C954E76B839F" box="[707,791,380,401]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="9" unit="mm" value="8.8">8.8 mm</quantity>
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||
. As
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||
<typeStatus id="1D4B8801315D1E33FB2FC954E7D9839F" box="[846,933,380,401]" pageId="9" type="holotype">holotype</typeStatus>
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||
except in the following respects:
|
||
<emphasis id="F084EAB1315D1E33FB45C9BFE7E983A2" bold="true" box="[804,917,407,428]" pageId="9">Pronotum:</emphasis>
|
||
Punctures slightly denser. Hind wing
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||
<quantity id="05089B46315D1E33FB52C999E70283C8" box="[819,894,433,454]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="9" unit="mm" value="8.3">8.3 mm</quantity>
|
||
in length.
|
||
<emphasis id="F084EAB1315D1E33FB8AC999E02983C8" bold="true" box="[1003,1109,433,454]" pageId="9">Pygidium:</emphasis>
|
||
Convex but not strongly so.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AEA6528315D1E33FABAC9CEE0D68045" pageId="9" type="etymology">
|
||
<paragraph id="C24F36A3315D1E33FABAC9CEE0D68045" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABAC9CEE73183F5" bold="true" box="[731,845,486,507]" pageId="9">Etymology.</emphasis>
|
||
The specific epithet is Latin that indicates “similar to...”, in reference to the similarity between
|
||
<taxonomicName id="05F04D20315D1E33FB3ECA34E7BC803F" authorityName="Ratcliffe" authorityYear="2014" box="[863,960,540,561]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FB3ECA34E7BC803F" box="[863,960,540,561]" italics="true" pageId="9">N. similis</emphasis>
|
||
</taxonomicName>
|
||
from
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<collectingCountry id="BAE77633315D1E33FC67CA33E055803E" box="[1030,1065,539,560]" name="Iceland" pageId="9">Isla</collectingCountry>
|
||
Isabela and
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<taxonomicName id="05F04D20315D1E33FAA2CA1EE7B98045" authority="(Mutchler)" baseAuthorityName="Mutchler" box="[707,965,566,587]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="linelli">
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<emphasis id="F084EAB1315D1E33FAA2CA1EE7278045" box="[707,859,566,587]" italics="true" pageId="9">Neoryctes linelli</emphasis>
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(Mutchler)
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||
</taxonomicName>
|
||
from
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<collectingCountry id="BAE77633315D1E33FB9FCA1EE0DA8045" box="[1022,1190,566,587]" name="Isla San Cristobal" pageId="9">Isla San Cristóbal</collectingCountry>
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.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="8AEA6528315D1E33FABACA79E05880E4" pageId="9" type="distribution">
|
||
<paragraph id="C24F36A3315D1E33FABACA79E0188095" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABACA79E7218068" bold="true" box="[731,861,593,614]" pageId="9">Distribution.</emphasis>
|
||
<taxonomicName id="05F04D20315D1E33FB0ACA79E06D8068" authorityName="Ratcliffe" authorityYear="2014" box="[875,1041,593,614]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FB0ACA79E06D8068" box="[875,1041,593,614]" italics="true" pageId="9">Neoryctes similis</emphasis>
|
||
</taxonomicName>
|
||
is known only from Isabela Island (formerly Albermarle) in the
|
||
<collectingRegion id="0034F841315D1E33FAA2CAAEE75B8095" box="[707,807,646,667]" country="Ecuador" name="Galapagos" pageId="9">Galápagos</collectingRegion>
|
||
Archipelago,
|
||
<collectingCountry id="BAE77633315D1E33FBD3CAAEE07D8095" box="[946,1025,646,667]" name="Ecuador" pageId="9">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="5ACB2A26315D1E33FC71CAAEE0248095" box="[1040,1112,646,667]" captionStart="Fig" captionStartId="9.[197,231,945,964]" captionTargetBox="[173,659,193,922]" captionTargetId="figure-592@9.[172,659,192,922]" captionTargetPageId="9" captionText="Fig. 16. Isla Isabela, Galápagos Islands. Image courtesy of Frans Lanting/Lanting.com." figureDoi="http://doi.org/10.5281/zenodo.5369386" httpUri="https://zenodo.org/record/5369386/files/figure.png" pageId="9">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C24F36A3315D1E33FABACA88E72780DE" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABACA88E05D80BB" bold="true" box="[731,1057,672,693]" pageId="9">
|
||
Locality Records.
|
||
<collectingCountry id="BAE77633315D1E33FBE6CA88E78680BB" box="[903,1018,672,693]" name="Ecuador" pageId="9">ECUADOR</collectingCountry>
|
||
(2):
|
||
</emphasis>
|
||
<collectingRegion id="0034F841315D1E33FC45CA88E0D580BB" box="[1060,1193,672,693]" country="Ecuador" name="Galapagos" pageId="9">GALÁPAGOS</collectingRegion>
|
||
(2): Isla Isabela.
|
||
</paragraph>
|
||
<paragraph id="C24F36A3315D1E33FABACAFDE05880E4" blockId="9.[707,1194,194,1597]" box="[731,1060,725,746]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABACAFDE7B980E4" bold="true" box="[731,965,725,746]" pageId="9">Temporal Distribution.</emphasis>
|
||
July (2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AEA6528315D1E33FABACAD8E0058433" pageId="9" type="diagnosis">
|
||
<paragraph id="C24F36A3315D1E33FABACAD8E05A86C7" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABACAD8E738810B" bold="true" box="[731,836,752,773]" pageId="9">Diagnosis.</emphasis>
|
||
This species will key out to couplet
|
||
<quantity id="05089B46315D1E33FA8FCB22E7688111" box="[750,788,778,799]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" unit="in" value="3.0">3 in</quantity>
|
||
<bibRefCitation id="A6614B52315D1E33FB7CCB22E7AE8111" author="Cook" box="[797,978,778,800]" pageId="9" pagination="177 - 193" refId="ref10850" refString="Cook, J., H. F. Howden, and S. B. Peck. 1995. The Galapagos Islands' genus Neoryctes Arrow (Coleoptera: Scarabaeidae: Dynastinae). The Canadian Entomologist 127: 177 - 193." type="journal article" year="1995">
|
||
Cook
|
||
<emphasis id="F084EAB1315D1E33FB38CB23E7F8812E" box="[857,900,779,800]" italics="true" pageId="9">et al</emphasis>
|
||
. (1995)
|
||
</bibRefCitation>
|
||
because of the length of the wing (
|
||
<quantity id="05089B46315D1E33FB23CB0DE7EC8134" box="[834,912,805,826]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" unit="mm" value="8.7">8.7 mm</quantity>
|
||
), but then only parts of each of the remaining two choices conform to the specimen.
|
||
<taxonomicName id="05F04D20315D1E33FB5BCB72E0048161" authority="Dechambre" authorityName="Dechambre" authorityYear="1991" box="[826,1144,858,879]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="moreti">
|
||
<emphasis id="F084EAB1315D1E33FB5BCB72E78F8161" box="[826,1011,858,879]" italics="true" pageId="9">Neoryctes moreti</emphasis>
|
||
Dechambre
|
||
</taxonomicName>
|
||
has reduced eyes and an acute lateral process on the ocular canthus, whereas
|
||
<taxonomicName id="05F04D20315D1E33FC6DCBB8E00481AB" authorityName="Ratcliffe" authorityYear="2014" box="[1036,1144,912,933]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FC6DCBB8E00481AB" box="[1036,1144,912,933]" italics="true" pageId="9">N. similis</emphasis>
|
||
</taxonomicName>
|
||
has neither;
|
||
<taxonomicName id="05F04D20315D1E33FB77CB82E70F81B1" authorityName="Dechambre" authorityYear="1991" box="[790,883,938,959]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="moreti">
|
||
<emphasis id="F084EAB1315D1E33FB77CB82E70F81B1" box="[790,883,938,959]" italics="true" pageId="9">N. moreti</emphasis>
|
||
</taxonomicName>
|
||
has a reduced medial process on the parameres, whereas this process is prominent in
|
||
<taxonomicName id="05F04D20315D1E33FB6FCBF7E71581FA" authorityName="Ratcliffe" authorityYear="2014" box="[782,873,991,1012]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FB6FCBF7E71581FA" box="[782,873,991,1012]" italics="true" pageId="9">N. similis</emphasis>
|
||
</taxonomicName>
|
||
. In the second part of couplet 3,
|
||
<taxonomicName id="05F04D20315D1E33FAA2CBD2E7A48601" authority="Cook" authorityName="Cook" box="[707,984,1018,1039]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="F084EAB1315D1E33FAA2CBD2E7E98601" box="[707,917,1018,1039]" italics="true" pageId="9">Neoryctes williamsi</emphasis>
|
||
Cook
|
||
</taxonomicName>
|
||
has the hind wing greatly reduced and lacking an anal fold, whereas
|
||
<taxonomicName id="05F04D20315D1E33FAA2CC07E761864A" authorityName="Ratcliffe" authorityYear="2014" box="[707,797,1071,1092]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FAA2CC07E761864A" box="[707,797,1071,1092]" italics="true" pageId="9">N. similis</emphasis>
|
||
</taxonomicName>
|
||
has a larger wing with an anal fold. The parameres of
|
||
<taxonomicName id="05F04D20315D1E33FB2ACC62E7DB8651" authorityName="Ratcliffe" authorityYear="2014" box="[843,935,1098,1119]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="F084EAB1315D1E33FB2ACC62E7DB8651" box="[843,935,1098,1119]" italics="true" pageId="9">N. similis</emphasis>
|
||
</taxonomicName>
|
||
are most similar to those of
|
||
<taxonomicName id="05F04D20315D1E33FA83CC4CE7428677" box="[738,830,1124,1145]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="linelli">
|
||
<emphasis id="F084EAB1315D1E33FA83CC4CE7428677" box="[738,830,1124,1145]" italics="true" pageId="9">N. linelli</emphasis>
|
||
</taxonomicName>
|
||
, but in that species the wings are longer than
|
||
<quantity id="05089B46315D1E33FB5ACC57E7F8869A" box="[827,900,1151,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" unit="mm" value="10.0">10 mm</quantity>
|
||
.
|
||
<taxonomicName id="05F04D20315D1E33FBF0CC57E041869A" box="[913,1085,1151,1172]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="F084EAB1315D1E33FBF0CC57E78C869A" box="[913,1008,1151,1172]" italics="true" pageId="9">Neoryctes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have variable character states (
|
||
<bibRefCitation id="A6614B52315D1E33FBFFCCB1E02C86A0" author="Cook" box="[926,1104,1177,1198]" pageId="9" pagination="177 - 193" refId="ref10850" refString="Cook, J., H. F. Howden, and S. B. Peck. 1995. The Galapagos Islands' genus Neoryctes Arrow (Coleoptera: Scarabaeidae: Dynastinae). The Canadian Entomologist 127: 177 - 193." type="journal article" year="1995">
|
||
Cook
|
||
<emphasis id="F084EAB1315D1E33FBBFCCB1E07186A0" box="[990,1037,1177,1198]" italics="true" pageId="9">et al</emphasis>
|
||
. 1995
|
||
</bibRefCitation>
|
||
), and so identifications can be problematical.
|
||
</paragraph>
|
||
<paragraph id="C24F36A3315D1E33FABACCE6E05B8409" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<bibRefCitation id="A6614B52315D1E33FABACCE6E7EB86ED" author="Cook" box="[731,919,1230,1251]" pageId="9" pagination="177 - 193" refId="ref10850" refString="Cook, J., H. F. Howden, and S. B. Peck. 1995. The Galapagos Islands' genus Neoryctes Arrow (Coleoptera: Scarabaeidae: Dynastinae). The Canadian Entomologist 127: 177 - 193." type="journal article" year="1995">
|
||
Cook
|
||
<emphasis id="F084EAB1315D1E33FB78CCE6E73A86ED" box="[793,838,1230,1251]" italics="true" pageId="9">et al</emphasis>
|
||
. (1995)
|
||
</bibRefCitation>
|
||
was aware of, but did not see, the Cawley Mountain specimens referred to in
|
||
<bibRefCitation id="A6614B52315D1E33FABDCD2BE7FD8716" author="Van Dyke" box="[732,897,1283,1304]" pageId="9" pagination="1 - 181" refId="ref11854" refString="Van Dyke, E. C. 1953. The Coleoptera of the Galapagos Islands. Occasional Papers of the California Academy of Sciences 22: 1 - 181." type="journal article" year="1953">Van Dyke (1953)</bibRefCitation>
|
||
, and the new species described here is undoubtedly those specimens. They were in the collections of the
|
||
<collectingRegion id="0034F841315D1E33FBA1CD11E0588740" box="[960,1060,1337,1358]" country="United States of America" name="California" pageId="9">California</collectingRegion>
|
||
Academy of Sciences and placed in a series with an old header label of
|
||
<taxonomicName id="05F04D20315D1E33FB75CD46E03C878D" authority="(Waterhouse)" baseAuthorityName="Waterhouse" box="[788,1088,1390,1411]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="galapagoensis">
|
||
<emphasis id="F084EAB1315D1E33FB75CD46E7C6878D" box="[788,954,1390,1411]" italics="true" pageId="9">N. galapagoensis</emphasis>
|
||
(Waterhouse)
|
||
</taxonomicName>
|
||
(L. Saylor determination). Henry Howden arranged for the loan from the Academy’ s
|
||
<taxonomicName id="05F04D20315D1E33FBA1CD8BE06087B6" authorityName="Arrow" authorityYear="1908" box="[960,1052,1443,1464]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F084EAB1315D1E33FBA1CD8BE06087B6" box="[960,1052,1443,1464]" italics="true" pageId="9">Neoryctes</emphasis>
|
||
</taxonomicName>
|
||
specimens for the revision by
|
||
<bibRefCitation id="A6614B52315D1E33FB34CD95E07887DC" author="Cook" box="[853,1028,1469,1490]" pageId="9" pagination="177 - 193" refId="ref10850" refString="Cook, J., H. F. Howden, and S. B. Peck. 1995. The Galapagos Islands' genus Neoryctes Arrow (Coleoptera: Scarabaeidae: Dynastinae). The Canadian Entomologist 127: 177 - 193." type="journal article" year="1995">
|
||
Cook
|
||
<emphasis id="F084EAB1315D1E33FBEECD95E7C487DC" box="[911,952,1469,1490]" italics="true" pageId="9">et al</emphasis>
|
||
. (1995)
|
||
</bibRefCitation>
|
||
, but it seems that not all were sent since the Cawley Mountain material was not included in the revision.
|
||
</paragraph>
|
||
<paragraph id="C24F36A3315D1E33FABACE25E0058433" blockId="9.[707,1194,194,1597]" pageId="9">
|
||
<emphasis id="F084EAB1315D1E33FABACE25E7FF842C" bold="true" box="[731,899,1549,1570]" pageId="9">Natural History.</emphasis>
|
||
Nothing is known of the life history for any of the five
|
||
<taxonomicName id="05F04D20315D1E33FBA6CE00E0058433" box="[967,1145,1576,1597]" class="Insecta" family="Dynastidae" genus="Neoryctes" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="F084EAB1315D1E33FBA6CE00E05A8433" box="[967,1062,1576,1597]" italics="true" pageId="9">Neoryctes</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |