378 lines
36 KiB
XML
378 lines
36 KiB
XML
<document id="276A53B929B4BBF891E9AC678AC36A0E" ID-DOI="10.1649/0010-065X-68.4.663" ID-Zenodo-Dep="5369368" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630263369791" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2014" docId="4A5987B531521E32FADECA46E6168127" docLanguage="en" docName="0010-065X-68.4.663.pdf" docOrigin="The Coleopterists Bulletin 68 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-68.4.663" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Hiekeianus halophilus Ratcliffe 2014, new species" docType="treatment" docVersion="4" lastPageNumber="680" masterDocId="B660FFCD31541E3AF861C828E47C820E" masterDocTitle="A New Genus and Species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other New Species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a Revised Key to the Genera of New World Pentodontini" masterLastPageNumber="680" masterPageNumber="663" pageNumber="663" updateTime="1699203105727" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A223AB044D09B32B0914431AC4B81767">
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<mods:title id="51C6F20D7B76601FB6C6277F59BF9ADB">A New Genus and Species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other New Species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a Revised Key to the Genera of New World Pentodontini</mods:title>
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</mods:titleInfo>
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<mods:name id="135391198D0327DE81618F8EDF957883" type="personal">
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<mods:roleTerm id="D05A7E1C96B3F41219F335EF1C8C4F78">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5EE7CB205B17AFC8C5D2B75A005C3158">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="9CA5836095D0358802877FFCDA7E341E">text</mods:typeOfResource>
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<mods:title id="933527832D6A6DC178C6E2708A0510B2">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="87B2CBCE22AF221F1FE5C963B6B8FC22">
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<mods:date id="21444601F511CFB228931BFB60210F0B">2014</mods:date>
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<mods:detail id="846FADBCF6C312AE34B74D695506330F" type="pubDate">
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<mods:number id="42E04F80262DE3A31EE676C44CB347D7">2014-12-31</mods:number>
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<mods:detail id="5F07BD282C9DA4B75295467C52FD85CE" type="volume">
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<mods:number id="A622BA590A8256B6D3289399BBD02767">68</mods:number>
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<mods:number id="E057DB64CFF9BD669AC4EA34A5CAC24B">4</mods:number>
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<mods:start id="0772D5C4ABFEB80A9B7D83F46DBB4D61">663</mods:start>
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<mods:end id="70D8330FD39FAFFF1EF108F23E3CA5FB">680</mods:end>
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<mods:location id="BE024BAFC58869E7459C03CABE05EEC4">
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<mods:url id="3CBD42F271DE953F17E2262A322BF983">http://dx.doi.org/10.1649/0010-065x-68.4.663</mods:url>
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<mods:classification id="7ECE193FFADB33FEF2F3D479C7CDE89A">journal article</mods:classification>
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<mods:identifier id="F4F9C0C112818477EE1A2248991FED3D" type="DOI">10.1649/0010-065X-68.4.663</mods:identifier>
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<mods:identifier id="34FAA246D7FFE46887AE960C8A5E5ED1" type="Zenodo-Dep">5369368</mods:identifier>
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<treatment id="4A5987B531521E32FADECA46E6168127" ID-DOI="http://doi.org/10.5281/zenodo.5461073" ID-Zenodo-Dep="5461073" LSID="urn:lsid:plazi:treatment:4A5987B531521E32FADECA46E6168127" httpUri="http://treatment.plazi.org/id/4A5987B531521E32FADECA46E6168127" lastPageId="8" pageId="6">
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<subSubSection id="8AEA652831521E3CFADECA46E79E8090" pageId="6" type="nomenclature">
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<paragraph id="C24F36A331521E3CFADECA46E79E8090" blockId="6.[703,1154,622,671]" pageId="6">
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<emphasis id="F084EAB131521E3CFADECA46E0FE808D" bold="true" box="[703,1154,622,643]" pageId="6">
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<taxonomicName id="05F04D2031521E3CFADECA46E783808D" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2014" box="[703,1023,622,643]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="halophilus" status="sp. nov.">
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<emphasis id="F084EAB131521E3CFADECA46E7E2808D" bold="true" box="[703,926,622,643]" italics="true" pageId="6">Hiekeianus halophilus</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="EBB757CA31521E3CFC6DCA46E0FE808D" box="[1036,1154,622,643]" pageId="6" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="5ACB2A2631521E3CFB07CAA1E7A68090" box="[870,986,649,671]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[197,229,844,863]" captionStartId-1="7.[197,229,1686,1705]" captionTargetBox-0="[230,588,199,821]" captionTargetBox-1="[183,1183,913,1663]" captionTargetId-0="figure-205@7.[172,659,183,833]" captionTargetId-1="figure-182@7.[182,1183,913,1663]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Fig. 11. Hiekeianus halophilus, holotype." captionText-1="Fig. 12. Salt catchments at Salineras de Maras, Cusco, Peru, type locality for Hiekeianus halophilus. Photograph by M. L. Jameson." figureDoi-0="http://doi.org/10.5281/zenodo.5369380" figureDoi-1="http://doi.org/10.5281/zenodo.5369382" httpUri-0="https://zenodo.org/record/5369380/files/figure.png" httpUri-1="https://zenodo.org/record/5369382/files/figure.png" pageId="6">Figs. 11–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="8AEA652831521E3DFAA4CA9CE0158351" lastPageId="7" pageId="6" type="materials_examined">
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<paragraph id="C24F36A331521E3CFAA4CA9CE7448166" blockId="6.[685,1173,692,1377]" pageId="6">
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<emphasis id="F084EAB131521E3CFAA4CA9CE72180C7" bold="true" box="[709,861,692,713]" pageId="6">Type Material.</emphasis>
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<materialsCitation id="72983CFE31521E3CFB0BCA9CE7448166" collectingDate="2012-01-05" collectorName="M. Jameson & L. Russell" country="Peru" elevation="3000" location="University of Nebraska State Museum" longitude="-13.302222" pageId="6" specimenCount="2" stateProvince="Cusco" typeStatus="holotype">
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<typeStatus id="1D4B880131521E3CFB0BCA9CE7B580C7" box="[874,969,692,713]" pageId="6" type="holotype">Holotype</typeStatus>
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female with handwritten label: “
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<collectingCountry id="BAE7763331521E3CFB23CAE6E7FF80ED" box="[834,899,718,739]" name="Peru" pageId="6">PERU</collectingCountry>
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:
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<collectingRegion id="0034F84131521E3CFBEECAE6E7B780ED" box="[911,971,718,739]" country="Peru" name="Cusco" pageId="6">Cusco</collectingRegion>
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/Salineras de Maras/ S
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<geoCoordinate id="A7C4506431521E3CFADACAC1E73980F0" box="[699,837,744,766]" degrees="13" direction="west" minutes="18" orientation="longitude" pageId="6" precision="15" seconds="08" value="-13.302222">13°18′08″ W</geoCoordinate>
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72°9′15″/
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<date id="B64E106331521E3CFBC7CAC1E06080F0" box="[934,1052,745,766]" pageId="6" value="2012-01-05">
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<collectingDate id="A60AE98B31521E3CFBC7CAC1E06080F0" box="[934,1052,745,766]" pageId="6" value="2012-01-05">5 Jan 2012</collectingDate>
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</date>
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,
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<quantity id="05089B4631521E3CFC4BCAC1E00E80F0" box="[1066,1138,745,766]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" unit="m" value="3000.0">
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<elevation id="49DDD19031521E3CFC4BCAC1E00E80F0" box="[1066,1138,745,766]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" unit="m" value="3000.0">3000m</elevation>
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</quantity>
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/
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<collectorName id="6F05537531521E3CFC1BCAC1E6828116" pageId="6">M. Jameson</collectorName>
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and
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<collectorName id="6F05537531521E3CFB50CB2BE7EE8116" box="[817,914,771,792]" pageId="6">L. Russell</collectorName>
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//Found dead in/salt catchment” and with my red
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<typeStatus id="1D4B880131521E3CFBEECB36E79D813D" box="[911,993,798,819]" pageId="6" type="holotype">holotype</typeStatus>
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label. Type deposited at the
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<location id="C72F607831521E3CFB7BCB10E0EF8143" LSID="urn:lsid:plazi:treatment:4A5987B531521E32FADECA46E6168127:C72F607831521E3CFB7BCB10E0EF8143" box="[794,1171,824,845]" country="Peru" longitude="-13.302222" name="University of Nebraska State Museum" pageId="6" stateProvince="Cusco">University of Nebraska State Museum</location>
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(Lincoln, NE).
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</materialsCitation>
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</paragraph>
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<paragraph id="C24F36A331521E3DFAA4CB45E0158351" blockId="6.[685,1173,692,1377]" lastBlockId="7.[707,1194,195,863]" lastPageId="7" pageId="6">
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||
<emphasis id="F084EAB131521E3CFAA4CB45E755818C" bold="true" box="[709,809,877,898]" pageId="6">
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<typeStatus id="1D4B880131521E3CFAA4CB45E758818C" box="[709,804,877,898]" pageId="6" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Female. Length 30.0 mm; width
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<quantity id="05089B4631521E3CFACCCBA0E77B8193" box="[685,775,904,925]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="6" unit="mm" value="14.5">14.5 mm</quantity>
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. Color dark reddish brown.
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<emphasis id="F084EAB131521E3CFC79CBA0E0298193" bold="true" box="[1048,1109,904,925]" pageId="6">Head:</emphasis>
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Frons with large, dense, deep, confluent punctures. Frontoclypeal region at center with short, transversely arcuate tubercle. Clypeus triangular, apex narrowly rounded, not reflexed, surface transversely rugose. Mandibles exposed, large, evenly rounded. Interocular width equals 4.0 transverse eye diameters. Antenna 10-segmented, club subequal in length to antennomeres 2–7.
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<emphasis id="F084EAB131521E3CFB17CC75E79A867C" bold="true" box="[886,998,1117,1138]" pageId="6">Pronotum:</emphasis>
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Surface convex, lacking fovea or tubercles, weakly crazed, anterior fifth half with moderately large, dense, deep punctures. Remainder of surface with moderately large punctures sparse to moderate in density. Base with thick marginal bead.
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<emphasis id="F084EAB131521E3CFBE5CCC9E7B086F8" bold="true" box="[900,972,1249,1270]" pageId="6">Elytra:</emphasis>
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Sutural stria a row of deep, elongate punctures. Surface completely, densely punctate, punctures mostly large, glabrous.
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<emphasis id="F084EAB131521E3CFACCCD19E76F8748" bold="true" box="[685,787,1329,1350]" pageId="6">Pygidium:</emphasis>
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Disc shiny, with small, sparse, glabrous punctures. Base with narrow, transverse band of, setigerous punctures; punctures small to moderate, and setae long, reddish brown. In lateral view, surface convex.
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<emphasis id="F084EAB131531E3DFB5EC8D1E7088300" bold="true" box="[831,884,249,270]" pageId="7">Legs:</emphasis>
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Protibia quadridentate. Metatibia at apex expanded, truncate and minutely crenulate with 17 stout spinules.
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<emphasis id="F084EAB131531E3DFBCAC907E78E834A" bold="true" box="[939,1010,303,324]" pageId="7">Venter:</emphasis>
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Prosternal process moderately long, laminate, apex subtruncate.
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</paragraph>
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</subSubSection>
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<caption id="968F662B31521E3CF8CECDA2E7D58793" box="[175,937,1418,1437]" pageId="6" startId="6.[175,223,1418,1437]">
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<paragraph id="C24F36A331521E3CF8CECDA2E7D58793" blockId="6.[175,937,1418,1437]" box="[175,937,1418,1437]" pageId="6">
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<emphasis id="F084EAB131521E3CF8CECDA2E48B8793" bold="true" box="[175,247,1418,1437]" pageId="6">Table 1.</emphasis>
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Morphological comparison of
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<taxonomicName id="05F04D2031521E3CFA72CDA2E6BB8792" authorityName="Ratcliffe" authorityYear="2014" box="[531,711,1418,1436]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="caiporae">
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<emphasis id="F084EAB131521E3CFA72CDA2E6BB8792" box="[531,711,1418,1436]" italics="true" pageId="6">Stenocrates caiporae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="05F04D2031521E3CFA95CDA2E7D88792" authorityName="Ratcliffe" authorityYear="1977" box="[756,932,1418,1436]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="haackae">
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<emphasis id="F084EAB131521E3CFA95CDA2E7D88792" box="[756,932,1418,1436]" italics="true" pageId="6">Stenocrates haackae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph id="C24F36A331521E3CF8D6CD95E7EF84BA" pageId="6">
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<table id="B0F0C4033152E1C5F8F6CD95E0ED84BA" box="[151,1169,1469,1716]" gridcols="3" gridrows="10" pageId="6">
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<tr id="7CC034E13152E1C5F8F6CD95E0ED87DE" box="[151,1169,1469,1488]" gridrow="0" pageId="6">
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<th id="3F115D9D3152E1C5F8F6CD95E54F87DE" box="[151,307,1469,1488]" gridcol="0" gridrow="0" pageId="6">
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<emphasis id="F084EAB131521E3CF8D6CD95E56F87DE" bold="true" box="[183,275,1469,1488]" pageId="6">Character</emphasis>
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</th>
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<th id="3F115D9D3152E1C5F9F9CD95E6C787DE" box="[408,699,1469,1488]" gridcol="1" gridrow="0" pageId="6">
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<taxonomicName id="05F04D2031521E3CF999CD95E62787DE" authorityName="Ratcliffe" authorityYear="2014" box="[504,603,1469,1488]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="caiporae">
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<emphasis id="F084EAB131521E3CF999CD95E62787DE" bold="true" box="[504,603,1469,1488]" italics="true" pageId="6">S. caiporae</emphasis>
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</taxonomicName>
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</th>
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<th id="3F115D9D3152E1C5FB40CD95E0ED87DE" box="[801,1169,1469,1488]" gridcol="2" gridrow="0" pageId="6">
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<taxonomicName id="05F04D2031521E3CFBCBCD95E07787DE" authorityName="Ratcliffe" authorityYear="1977" box="[938,1035,1469,1488]" class="Insecta" family="Dynastidae" genus="Stenocrates" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="haackae">
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<emphasis id="F084EAB131521E3CFBCBCD95E07787DE" bold="true" box="[938,1035,1469,1488]" italics="true" pageId="6">S. haackae</emphasis>
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</taxonomicName>
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</th>
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</tr>
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<tr id="7CC034E13152E1C5F8F6CDCAE0ED87FB" box="[151,1169,1506,1525]" gridrow="1" pageId="6">
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<th id="3F115D9D3152E1C5F8F6CDCAE54F87FB" box="[151,307,1506,1525]" gridcol="0" gridrow="1" pageId="6">Length</th>
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<td id="3F115D9D3152E1C5F9F9CDCAE6C787FB" box="[408,699,1506,1525]" gridcol="1" gridrow="1" pageId="6">25 mm</td>
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<td id="3F115D9D3152E1C5FB40CDCAE0ED87FB" box="[801,1169,1506,1525]" gridcol="2" gridrow="1" pageId="6">21 mm</td>
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||
</tr>
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<tr id="7CC034E13152E1C5F8F6CDD2E0ED8403" box="[151,1169,1530,1549]" gridrow="2" pageId="6">
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<th id="3F115D9D3152E1C5F8F6CDD2E54F8403" box="[151,307,1530,1549]" gridcol="0" gridrow="2" pageId="6">Clypeal apex</th>
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<td id="3F115D9D3152E1C5F9F9CDD2E6C78403" box="[408,699,1530,1549]" gridcol="1" gridrow="2" pageId="6">Slightly emarginate</td>
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<td id="3F115D9D3152E1C5FB40CDD2E0ED8403" box="[801,1169,1530,1549]" gridcol="2" gridrow="2" pageId="6">Subtruncate</td>
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||
</tr>
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<tr id="7CC034E13152E1C5F8F6CE3AE0ED842B" box="[151,1169,1554,1573]" gridrow="3" pageId="6">
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||
<th id="3F115D9D3152E1C5F8F6CE3AE54F842B" box="[151,307,1554,1573]" gridcol="0" gridrow="3" pageId="6">Frons</th>
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<td id="3F115D9D3152E1C5F9F9CE3AE6C7842B" box="[408,699,1554,1573]" gridcol="1" gridrow="3" pageId="6">Smooth</td>
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||
<td id="3F115D9D3152E1C5FB40CE3AE0ED842B" box="[801,1169,1554,1573]" gridcol="2" gridrow="3" pageId="6">With large, dense punctures</td>
|
||
</tr>
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||
<tr id="7CC034E13152E1C5F8F6CE02E0ED8433" box="[151,1169,1578,1597]" gridrow="4" pageId="6">
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||
<th id="3F115D9D3152E1C5F8F6CE02E54F8433" box="[151,307,1578,1597]" gridcol="0" gridrow="4" pageId="6">Interocular width</th>
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||
<td id="3F115D9D3152E1C5F9F9CE02E6C78433" box="[408,699,1578,1597]" gridcol="1" gridrow="4" pageId="6">2.2 transverse eye diameters</td>
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||
<td id="3F115D9D3152E1C5FB40CE02E0ED8433" box="[801,1169,1578,1597]" gridcol="2" gridrow="4" pageId="6">2.8 transverse eye diameters</td>
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||
</tr>
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||
<tr id="7CC034E13152E1C5F8F6CE69E0ED845A" box="[151,1169,1601,1620]" gridrow="5" pageId="6">
|
||
<th id="3F115D9D3152E1C5F8F6CE69E54F845A" box="[151,307,1601,1620]" gridcol="0" gridrow="5" pageId="6">Pronotum</th>
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<td id="3F115D9D3152E1C5F9F9CE69E6C7845A" box="[408,699,1601,1620]" gridcol="1" gridrow="5" pageId="6">Mostly smooth with small, sparse</td>
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<td id="3F115D9D3152E1C5FB40CE69E0ED845A" box="[801,1169,1601,1620]" gridcol="2" gridrow="5" pageId="6">Small, sparse punctures on disc with large</td>
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||
</tr>
|
||
<tr id="7CC034E13152E1C5F8F6CE71E0ED8462" box="[151,1169,1625,1644]" gridrow="6" pageId="6" rowspan-0="1">
|
||
<td id="3F115D9D3152E1C5F9F9CE71E6C78462" box="[408,699,1625,1644]" gridcol="1" gridrow="6" pageId="6">punctures in posterior angle</td>
|
||
<td id="3F115D9D3152E1C5FB40CE71E0ED8462" box="[801,1169,1625,1644]" gridcol="2" gridrow="6" pageId="6">dense punctures in posterior angle</td>
|
||
</tr>
|
||
<tr id="7CC034E13152E1C5F8F6CE59E0ED848A" box="[151,1169,1649,1668]" gridrow="7" pageId="6">
|
||
<th id="3F115D9D3152E1C5F8F6CE59E54F848A" box="[151,307,1649,1668]" gridcol="0" gridrow="7" pageId="6">Pygidium</th>
|
||
<td id="3F115D9D3152E1C5F9F9CE59E6C7848A" box="[408,699,1649,1668]" gridcol="1" gridrow="7" pageId="6">Completely, densely punctate</td>
|
||
<td id="3F115D9D3152E1C5FB40CE59E0ED848A" box="[801,1169,1649,1668]" gridcol="2" gridrow="7" pageId="6">Sparse punctures on apical half</td>
|
||
</tr>
|
||
<tr id="7CC034E13152E1C5F8F6CEA1E0ED8492" box="[151,1169,1673,1692]" gridrow="8" pageId="6">
|
||
<th id="3F115D9D3152E1C5F8F6CEA1E54F8492" box="[151,307,1673,1692]" gridcol="0" gridrow="8" pageId="6">Protibial teeth</th>
|
||
<td id="3F115D9D3152E1C5F9F9CEA1E6C78492" box="[408,699,1673,1692]" gridcol="1" gridrow="8" pageId="6">Spaced subequally</td>
|
||
<td id="3F115D9D3152E1C5FB40CEA1E0ED8492" box="[801,1169,1673,1692]" gridcol="2" gridrow="8" pageId="6">Basal tooth slightly removed</td>
|
||
</tr>
|
||
<tr id="7CC034E13152E1C5F8F6CE89E0ED84BA" box="[151,1169,1697,1716]" gridrow="9" pageId="6">
|
||
<th id="3F115D9D3152E1C5F8F6CE89E54F84BA" box="[151,307,1697,1716]" gridcol="0" gridrow="9" pageId="6">Prosternal process</th>
|
||
<td id="3F115D9D3152E1C5F9F9CE89E6C784BA" box="[408,699,1697,1716]" gridcol="1" gridrow="9" pageId="6">Apex broadly rounded</td>
|
||
<td id="3F115D9D3152E1C5FB40CE89E0ED84BA" box="[801,1169,1697,1716]" gridcol="2" gridrow="9" pageId="6">Apex conical</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="8AEA652831531E3DFABAC94DE015800F" pageId="7" type="etymology">
|
||
<paragraph id="C24F36A331531E3DFABAC94DE015800F" blockId="7.[707,1194,195,863]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DFABAC94DE7318374" bold="true" box="[731,845,357,378]" pageId="7">Etymology.</emphasis>
|
||
The specific epithet is derived from the Greek words
|
||
<emphasis id="F084EAB131531E3DFB3DC9A8E7F1839B" box="[860,909,384,405]" italics="true" pageId="7">halos</emphasis>
|
||
for salt and
|
||
<emphasis id="F084EAB131531E3DFB9AC9A8E053839B" box="[1019,1071,384,405]" italics="true" pageId="7">philia</emphasis>
|
||
for fondness. It is used here in reference to the salt catchments in which it was found. This is not to imply that this species actually has a predilection for salty habitats, but that it was simply found dead in such a place.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AEA652831531E3DFABACA2FE03980AC" pageId="7" type="distribution">
|
||
<paragraph id="C24F36A331531E3DFABACA2FE7CA805C" blockId="7.[707,1194,195,863]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DFABACA2FE7218012" bold="true" box="[731,861,519,540]" pageId="7">Distribution.</emphasis>
|
||
<taxonomicName id="05F04D2031531E3DFB0ACA2FE03A8012" authorityName="Ratcliffe" authorityYear="2014" box="[875,1094,519,540]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DFB0ACA2FE03A8012" box="[875,1094,519,540]" italics="true" pageId="7">Hiekeianus halophilus</emphasis>
|
||
</taxonomicName>
|
||
is known only from the area of the Maras salt pan area in
|
||
<collectingRegion id="0034F84131531E3DFAA2CA15E682805C" box="[707,766,573,594]" country="Peru" name="Cusco" pageId="7">Cusco</collectingRegion>
|
||
Department,
|
||
<collectingCountry id="BAE7763331531E3DFBE5CA15E7CE805C" box="[900,946,573,594]" name="Peru" pageId="7">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C24F36A331531E3DFABACA70E7FC8086" blockId="7.[707,1194,195,863]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DFABACA70E7A38063" bold="true" box="[731,991,600,621]" pageId="7">
|
||
Locality Record.
|
||
<collectingCountry id="BAE7763331531E3DFBF5CA70E7A78063" box="[916,987,600,621]" name="Peru" pageId="7">PERU</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="0034F84131531E3DFB88CA70E0438063" box="[1001,1087,600,621]" country="Peru" name="Cusco" pageId="7">CUSCO</collectingRegion>
|
||
: Salineras de Maras (
|
||
<figureCitation id="5ACB2A2631531E3DFB4ACA5BE70F8086" box="[811,883,627,648]" captionStart="Fig" captionStartId="7.[197,229,1686,1705]" captionTargetBox="[183,1183,913,1663]" captionTargetId="figure-182@7.[182,1183,913,1663]" captionTargetPageId="7" captionText="Fig. 12. Salt catchments at Salineras de Maras, Cusco, Peru, type locality for Hiekeianus halophilus. Photograph by M. L. Jameson." figureDoi="http://doi.org/10.5281/zenodo.5369382" httpUri="https://zenodo.org/record/5369382/files/figure.png" pageId="7">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C24F36A331531E3DFABACAA5E03980AC" blockId="7.[707,1194,195,863]" box="[731,1093,653,674]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DFABACAA5E7B980AC" bold="true" box="[731,965,653,674]" pageId="7">Temporal Distribution.</emphasis>
|
||
January (1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AEA652831531E32FABACA80E6168127" lastPageId="8" pageId="7" type="diagnosis">
|
||
<paragraph id="C24F36A331531E3DFABACA80E7DB8127" blockId="7.[707,1194,195,863]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DFABACA80E73880B3" bold="true" box="[731,836,680,701]" pageId="7">Diagnosis.</emphasis>
|
||
The female of
|
||
<taxonomicName id="05F04D2031531E3DFB96CA80E0F080B3" authorityName="Ratcliffe" authorityYear="2014" box="[1015,1164,680,701]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DFB96CA80E0F080B3" box="[1015,1164,680,701]" italics="true" pageId="7">H. halophilus</emphasis>
|
||
</taxonomicName>
|
||
is described as new because it is so different from the females of
|
||
<taxonomicName id="05F04D2031531E3DFB3ECAF6E78780FD" authorityName="Endrodi" authorityYear="1978" box="[863,1019,734,755]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tuberculatus">
|
||
<emphasis id="F084EAB131531E3DFB3ECAF6E78780FD" box="[863,1019,734,755]" italics="true" pageId="7">H. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
. The differences between
|
||
<taxonomicName id="05F04D2031531E3DFB7ECAD1E7D78100" authorityName="Ratcliffe" authorityYear="2014" box="[799,939,761,782]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DFB7ECAD1E7D78100" box="[799,939,761,782]" italics="true" pageId="7">H. halophilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05F04D2031531E3DFB82CAD1E0FD8100" authorityName="Endrodi" authorityYear="1978" box="[995,1153,761,782]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tuberculatus">
|
||
<emphasis id="F084EAB131531E3DFB82CAD1E0FD8100" box="[995,1153,761,782]" italics="true" pageId="7">H. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
are summarized in
|
||
<tableCitation id="8F72031831531E3DFB38CB3CE7DF8127" box="[857,931,788,809]" captionStart="Table 2" captionStartId="8.[175,223,193,212]" captionText="Table 2. Morphological comparison of Hiekeianus halophilus and Hiekeianus tuberculatus." pageId="7">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C24F36A331531E32FABACB07E58080FA" blockId="7.[707,1194,195,863]" lastBlockId="8.[151,638,545,809]" lastPageId="8" pageId="7">
|
||
In size and
|
||
<emphasis id="F084EAB131531E3DFB28CB07E7F4814A" box="[841,904,815,836]" italics="true" pageId="7">gestalt</emphasis>
|
||
,
|
||
<taxonomicName id="05F04D2031531E3DFBF4CB07E06B814A" authorityName="Ratcliffe" authorityYear="2014" box="[917,1047,815,836]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DFBF4CB07E06B814A" box="[917,1047,815,836]" italics="true" pageId="7">H. halophilus</emphasis>
|
||
</taxonomicName>
|
||
also resembles
|
||
<taxonomicName id="05F04D2031531E3DFAA2CB62E7948151" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="1981" box="[707,1000,842,863]" class="Insecta" family="Dynastidae" genus="Barutus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="hartmanni">
|
||
<emphasis id="F084EAB131531E3DFAA2CB62E7FA8151" box="[707,902,842,863]" italics="true" pageId="7">Barutus hartmanni</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="BAE7763331531E3DFC4DCB62E0078151" box="[1068,1147,842,863]" name="Panama" pageId="7">Panama</collectingCountry>
|
||
and
|
||
<collectingCountry id="BAE77633315C1E32F8F6CA09E5798038" box="[151,261,545,566]" name="Costa Rica" pageId="8">Costa Rica</collectingCountry>
|
||
, but in
|
||
<taxonomicName id="05F04D20315C1E32F935CA09E5A88038" authorityName="Ratcliffe" authorityYear="1981" box="[340,468,545,566]" class="Insecta" family="Dynastidae" genus="Barutus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="hartmanni">
|
||
<emphasis id="F084EAB1315C1E32F935CA09E51E8038" box="[340,354,545,566]" italics="true" pageId="8">B</emphasis>
|
||
.
|
||
<emphasis id="F084EAB1315C1E32F910CA09E5A88038" box="[369,468,545,566]" italics="true" pageId="8">hartmanni</emphasis>
|
||
</taxonomicName>
|
||
the clypeal apex is broader, the mandibles are weakly lobed, the frontoclypeal region has a binodose tubercle, the mentum is lanceolate rather than constricted at the subapex, the last abdominal sternite is subtruncate, and the prosternal process is large, thick, and rounded as opposed to the laminate, subtruncate process in
|
||
<taxonomicName id="05F04D20315C1E32F915CAF7E58A80FA" authorityName="Ratcliffe" authorityYear="2014" box="[372,502,735,756]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB1315C1E32F915CAF7E58A80FA" box="[372,502,735,756]" italics="true" pageId="8">H. halophilus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="968F662B31531E3DF8A4CB64E6478151" ID-DOI="http://doi.org/10.5281/zenodo.5369380" ID-Zenodo-Dep="5369380" box="[197,571,844,863]" httpUri="https://zenodo.org/record/5369380/files/figure.png" pageId="7" startId="7.[197,229,844,863]" targetBox="[230,588,199,821]" targetPageId="7">
|
||
<paragraph id="C24F36A331531E3DF8A4CB64E6478151" blockId="7.[197,571,844,863]" box="[197,571,844,863]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DF8A4CB64E5758151" bold="true" box="[197,265,844,863]" pageId="7">Fig. 11.</emphasis>
|
||
<taxonomicName id="05F04D2031531E3DF97ECB64E5A28150" authorityName="Ratcliffe" authorityYear="2014" box="[287,478,844,862]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DF97ECB64E5A28150" box="[287,478,844,862]" italics="true" pageId="7">Hiekeianus halophilus</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="968F662B31531E3DF8A4CEBEE52D84CF" ID-DOI="http://doi.org/10.5281/zenodo.5369382" ID-Zenodo-Dep="5369382" httpUri="https://zenodo.org/record/5369382/files/figure.png" pageId="7" startId="7.[197,229,1686,1705]" targetBox="[183,1183,913,1663]" targetPageId="7">
|
||
<paragraph id="C24F36A331531E3DF8A4CEBEE52D84CF" blockId="7.[173,1192,1686,1729]" pageId="7">
|
||
<emphasis id="F084EAB131531E3DF8A4CEBEE57784A7" bold="true" box="[197,267,1686,1705]" pageId="7">Fig. 12.</emphasis>
|
||
Salt catchments at Salineras de Maras, Cusco, Peru, type locality for
|
||
<taxonomicName id="05F04D2031531E3DFB1DCEBEE04784A6" authorityName="Ratcliffe" authorityYear="2014" box="[892,1083,1686,1704]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB131531E3DFB1DCEBEE04784A6" box="[892,1083,1686,1704]" italics="true" pageId="7">Hiekeianus halophilus</emphasis>
|
||
</taxonomicName>
|
||
. Photograph by M. L. Jameson.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="968F662B315C1E32F8CEC8E9E7AE82DA" box="[175,978,193,212]" pageId="8" startId="8.[175,223,193,212]">
|
||
<paragraph id="C24F36A3315C1E32F8CEC8E9E7AE82DA" blockId="8.[175,978,193,212]" box="[175,978,193,212]" pageId="8">
|
||
<emphasis id="F084EAB1315C1E32F8CEC8E9E48482DA" bold="true" box="[175,248,193,212]" pageId="8">Table 2.</emphasis>
|
||
Morphological comparison of
|
||
<taxonomicName id="05F04D20315C1E32FA72C8E9E6AE82DD" authorityName="Ratcliffe" authorityYear="2014" box="[531,722,193,211]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB1315C1E32FA72C8E9E6AE82DD" box="[531,722,193,211]" italics="true" pageId="8">Hiekeianus halophilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05F04D20315C1E32FA9EC8E9E7B182DD" authorityName="Endrodi" authorityYear="1978" box="[767,973,193,211]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tuberculatus">
|
||
<emphasis id="F084EAB1315C1E32FA9EC8E9E7B182DD" box="[767,973,193,211]" italics="true" pageId="8">Hiekeianus tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C24F36A3315C1E32F8AFC8DCE7E683E2" pageId="8">
|
||
<table id="B0F0C403315CE1C5F8F6C8DCE0EF83E2" box="[151,1171,244,492]" gridcols="3" gridrows="10" pageId="8">
|
||
<tr id="7CC034E1315CE1C5F8F6C8DCE0EF8309" box="[151,1171,244,263]" gridrow="0" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C8DCE51C8309" box="[151,352,244,263]" gridcol="0" gridrow="0" pageId="8">
|
||
<emphasis id="F084EAB1315C1E32F8AFC8DCE5568309" bold="true" box="[206,298,244,263]" pageId="8">Character</emphasis>
|
||
</th>
|
||
<th id="3F115D9D315CE1C5F9CAC8DCE6968309" box="[427,746,244,263]" gridcol="1" gridrow="0" pageId="8">
|
||
<taxonomicName id="05F04D20315C1E32FA6FC8DCE6FB8309" authorityName="Ratcliffe" authorityYear="2014" box="[526,647,244,263]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="halophilus">
|
||
<emphasis id="F084EAB1315C1E32FA6FC8DCE6FB8309" bold="true" box="[526,647,244,263]" italics="true" pageId="8">H. halophilus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="3F115D9D315CE1C5FB57C8DCE0EF8309" box="[822,1171,244,263]" gridcol="2" gridrow="0" pageId="8">
|
||
<taxonomicName id="05F04D20315C1E32FBC1C8DCE0568309" authorityName="Endrodi" authorityYear="1978" box="[928,1066,244,263]" class="Insecta" family="Dynastidae" genus="Hiekeianus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="tuberculatus">
|
||
<emphasis id="F084EAB1315C1E32FBC1C8DCE0568309" bold="true" box="[928,1066,244,263]" italics="true" pageId="8">H. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C931E0EF8322" box="[151,1171,281,300]" gridrow="1" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C931E51C8322" box="[151,352,281,300]" gridcol="0" gridrow="1" pageId="8">Length and depth</th>
|
||
<td id="3F115D9D315CE1C5F9CAC931E6968322" box="[427,746,281,300]" gridcol="1" gridrow="1" pageId="8">30 mm and 13 mm</td>
|
||
<td id="3F115D9D315CE1C5FB57C931E0EF8322" box="[822,1171,281,300]" gridcol="2" gridrow="1" pageId="8">24–26 mm and 11 mm</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C919E0EF834A" box="[151,1171,305,324]" gridrow="2" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C919E51C834A" box="[151,352,305,324]" gridcol="0" gridrow="2" pageId="8">Frontoclypeal suture</th>
|
||
<td id="3F115D9D315CE1C5F9CAC919E696834A" box="[427,746,305,324]" gridcol="1" gridrow="2" pageId="8">Tubercle short, transverse, arcuate</td>
|
||
<td id="3F115D9D315CE1C5FB57C919E0EF834A" box="[822,1171,305,324]" gridcol="2" gridrow="2" pageId="8">Tubercle simple, conical</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C961E0EF8352" box="[151,1171,329,348]" gridrow="3" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C961E51C8352" box="[151,352,329,348]" gridcol="0" gridrow="3" pageId="8">Clypeal apex</th>
|
||
<td id="3F115D9D315CE1C5F9CAC961E6968352" box="[427,746,329,348]" gridcol="1" gridrow="3" pageId="8">Narrowly rounded</td>
|
||
<td id="3F115D9D315CE1C5FB57C961E0EF8352" box="[822,1171,329,348]" gridcol="2" gridrow="3" pageId="8">Narrowly bidentate</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C949E0EF837A" box="[151,1171,353,372]" gridrow="4" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C949E51C837A" box="[151,352,353,372]" gridcol="0" gridrow="4" pageId="8">Mentum surface</th>
|
||
<td id="3F115D9D315CE1C5F9CAC949E696837A" box="[427,746,353,372]" gridcol="1" gridrow="4" pageId="8">Punctures sparse, shallow</td>
|
||
<td id="3F115D9D315CE1C5FB57C949E0EF837A" box="[822,1171,353,372]" gridcol="2" gridrow="4" pageId="8">Punctures dense, deep</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C951E0EF8382" box="[151,1171,377,396]" gridrow="5" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C951E51C8382" box="[151,352,377,396]" gridcol="0" gridrow="5" pageId="8">Mentum shape</th>
|
||
<td id="3F115D9D315CE1C5F9CAC951E6968382" box="[427,746,377,396]" gridcol="1" gridrow="5" pageId="8">Lyre-shaped and elongated anteriorly</td>
|
||
<td id="3F115D9D315CE1C5FB57C951E0EF8382" box="[822,1171,377,396]" gridcol="2" gridrow="5" pageId="8">Lyre-shaped but not elongated anteriorly</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C9B9E0EF83AA" box="[151,1171,401,420]" gridrow="6" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C9B9E51C83AA" box="[151,352,401,420]" gridcol="0" gridrow="6" pageId="8">Pronotum</th>
|
||
<td id="3F115D9D315CE1C5F9CAC9B9E69683AA" box="[427,746,401,420]" gridcol="1" gridrow="6" pageId="8">Basal bead thick, complete</td>
|
||
<td id="3F115D9D315CE1C5FB57C9B9E0EF83AA" box="[822,1171,401,420]" gridcol="2" gridrow="6" pageId="8">Basal bead slender, incomplete at center</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C981E0EF83B2" box="[151,1171,425,444]" gridrow="7" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C981E51C83B2" box="[151,352,425,444]" gridcol="0" gridrow="7" pageId="8">Elytra</th>
|
||
<td id="3F115D9D315CE1C5F9CAC981E69683B2" box="[427,746,425,444]" gridcol="1" gridrow="7" pageId="8">Surface glabrous</td>
|
||
<td id="3F115D9D315CE1C5FB57C981E0EF83B2" box="[822,1171,425,444]" gridcol="2" gridrow="7" pageId="8">With sparse, reddish brown setae,</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C9E9E0EF83DA" box="[151,1171,449,468]" gridrow="8" pageId="8" rowspan-0="1" rowspan-1="1">
|
||
<td id="3F115D9D315CE1C5FB57C9E9E0EF83DA" box="[822,1171,449,468]" gridcol="2" gridrow="8" pageId="8">especially near apices</td>
|
||
</tr>
|
||
<tr id="7CC034E1315CE1C5F8F6C9F1E0EF83E2" box="[151,1171,473,492]" gridrow="9" pageId="8">
|
||
<th id="3F115D9D315CE1C5F8F6C9F1E51C83E2" box="[151,352,473,492]" gridcol="0" gridrow="9" pageId="8">Last abdominal sternite</th>
|
||
<td id="3F115D9D315CE1C5F9CAC9F1E69683E2" box="[427,746,473,492]" gridcol="1" gridrow="9" pageId="8">Weakly emarginate</td>
|
||
<td id="3F115D9D315CE1C5FB57C9F1E0EF83E2" box="[822,1171,473,492]" gridcol="2" gridrow="9" pageId="8">Subtruncate</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="C24F36A3315C1E32F8CECAD1E6168127" blockId="8.[151,638,545,809]" pageId="8">
|
||
<emphasis id="F084EAB1315C1E32F8CECAD1E52B8100" bold="true" box="[175,343,761,782]" pageId="8">Natural History.</emphasis>
|
||
The single
|
||
<typeStatus id="1D4B8801315C1E32F9A8CAD1E6658100" box="[457,537,761,782]" pageId="8" type="holotype">holotype</typeStatus>
|
||
was found dead, and so nothing is known of its life history.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |