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<document id="5B8A22668F5B3B19824599EA51523E34" ID-DOI="10.5281/zenodo.183058" ID-GBIF-Dataset="049d6547-b411-4ce9-bd66-1aaf47a293d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="183058" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459890547261" checkinUser="plazi" docAuthor="Silva, Anjana, Maduwage, Kalana &amp; Pethiyagoda, Rohan" docDate="2008" docId="DB5D87B9271CC0050DFB3B0EAE44FE8B" docLanguage="en" docName="zt01824p064.pdf" docOrigin="Zootaxa 1824" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Puntius kamalika Silva, Maduwage &amp; Pethiyagoda, 2008, sp. nov." docType="treatment" docVersion="7" lastPageNumber="60" masterDocId="2764FFC1271DC0000D6C3F4CAB60FFA1" masterDocTitle="Puntius kamalika, a new species of barb from Sri Lanka (Teleostei: Cyprinidae)" masterLastPageNumber="64" masterPageNumber="55" pageNumber="56" updateTime="1698227996130" updateUser="plazi">
<mods:mods id="9696C605C691CB1398AD7DA5A9E4EA89" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C85C5A1FCC29F20B95D9F3FAD03C94FD">
<mods:title id="FD08B102182C80936EDA091D555B3B5C">Puntius kamalika, a new species of barb from Sri Lanka (Teleostei: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="F1E086DE427BE287C343D5BC38742D4E" type="personal">
<mods:role id="4D078338E0C9CD0062E18D7C88A9A3F3">
<mods:roleTerm id="1D70D20B02BA61CE69970DB389D535F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18208CC757D97BE15DFF538E38C86F82">Silva, Anjana</mods:namePart>
</mods:name>
<mods:name id="D1185B5C164B2234DCDC35138A678419" type="personal">
<mods:role id="DF81ADFFEC737130E098F4B4E88145DD">
<mods:roleTerm id="75F1B069A2E4A496A13BCB653E6B4D0D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78E59FAC97524E5F5E9D51655EA57FDB">Maduwage, Kalana</mods:namePart>
</mods:name>
<mods:name id="86D07BF1E469313223D3C22EAA0FF990" type="personal">
<mods:role id="B64927A6DDBBA746B7FF743BA885A5E3">
<mods:roleTerm id="0849E5F78228F8E67E15819CB1DCC3D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD7AD2A6AA6787ECFA378A80DD96E99C">Pethiyagoda, Rohan</mods:namePart>
</mods:name>
<mods:typeOfResource id="47075D8E04A13FD010B8AF332ED3112D">text</mods:typeOfResource>
<mods:relatedItem id="3C750C61614F4C41D7E95B3B5949ABC2" type="host">
<mods:titleInfo id="1972C57D70ABB25303423F848B47B83A">
<mods:title id="A79CAEE1D37E0AB3838EE5CE601E5F1E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="21B43C974D1D457FF5229BDF7D61A4E9">
<mods:date id="6EC06227F7C912A5DA44110864E259A0">2008</mods:date>
<mods:detail id="CD09FAE197E243E16D61E516B45C8AFC" type="volume">
<mods:number id="B5FD65118D823FD30B9F93C5632964A2">1824</mods:number>
</mods:detail>
<mods:extent id="0D64FC756F8767B1D5E89C368D7A983A" unit="page">
<mods:start id="DF6B987F0E1B23942B9A801203F51D4C">55</mods:start>
<mods:end id="0A6E7333763923690F533A4F25A35746">64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="32A0E5C392C544FE5B9B19CF81967616">journal article</mods:classification>
<mods:identifier id="46D23827333979BCEA6080395CBF341B" type="DOI">10.5281/zenodo.183058</mods:identifier>
<mods:identifier id="AF10595092F8F8F77784E1DA7FBEEA7E" type="GBIF-Dataset">049d6547-b411-4ce9-bd66-1aaf47a293d7</mods:identifier>
<mods:identifier id="F6DEB5C4F7C205F713EE87609FC55326" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="66318AB0A0F4931D73E2FEBD4CC335D0" type="Zenodo-Dep">183058</mods:identifier>
</mods:mods>
<treatment id="DB5D87B9271CC0050DFB3B0EAE44FE8B" ID-DOI="http://doi.org/10.5281/zenodo.5659251" ID-GBIF-Taxon="119371885" ID-Zenodo-Dep="5659251" LSID="urn:lsid:plazi:treatment:DB5D87B9271CC0050DFB3B0EAE44FE8B" httpUri="http://treatment.plazi.org/id/DB5D87B9271CC0050DFB3B0EAE44FE8B" lastPageId="5" lastPageNumber="60" pageId="1" pageNumber="56">
<subSubSection id="1BEE6524271CC0010DFB3B0EAB8FFBDE" pageId="1" pageNumber="56" type="nomenclature">
<paragraph id="534B36AF271CC0010DFB3B0EAAB9FBFD" blockId="1.[151,473,1090,1151]" box="[151,473,1090,1116]" pageId="1" pageNumber="56">
<heading id="080381C3271CC0010DFB3B0EAAB9FBFD" bold="true" box="[151,473,1090,1116]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
<emphasis id="6180EABD271CC0010DFB3B0EAAB9FBFD" bold="true" box="[151,473,1090,1116]" pageId="1" pageNumber="56">
<taxonomicName id="94F44D2C271CC0010DFB3B0EAA0CFBFD" ID-CoL="4QJ55" box="[151,364,1090,1116]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="kamalika" status="sp. nov.">
<emphasis id="6180EABD271CC0010DFB3B0EAA0CFBFD" bold="true" box="[151,364,1090,1116]" italics="true" pageId="1" pageNumber="56">Puntius kamalika</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="7AB357C6271CC0010C103B0EAAB9FBFD" box="[380,473,1090,1116]" pageId="1" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="534B36AF271CC0010DFB3B29AB8FFBDE" blockId="1.[151,473,1090,1151]" box="[151,239,1125,1151]" pageId="1" pageNumber="56">
(
<figureCitation id="CBCF2A2A271CC0010DF33B29AB83FBDE" box="[159,227,1125,1151]" captionStart="FIGURE 1" captionStartId="3.[151,255,1516,1540]" captionTargetBox="[280,1307,198,1478]" captionTargetId="figure@3.[280,1307,194,1012]" captionTargetPageId="3" captionText="FIGURE 1. Puntius kamalika. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo." httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BEE6524271CC0010DFB3BFCAA4CFAA6" pageId="1" pageNumber="56" type="reference_group">
<paragraph id="534B36AF271CC0010DFB3BFCAA4CFAA6" blockId="1.[151,1436,1200,1287]" pageId="1" pageNumber="56">
<treatmentCitationGroup id="73E41181271CC0010DFB3BFCAA4CFAA6" pageId="1" pageNumber="56">
<taxonomicName id="94F44D2C271CC0010DFB3BFCAA3AFB66" ID-CoL="KMX4" box="[151,346,1200,1223]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="amphibius">
<emphasis id="6180EABD271CC0010DFB3BFCAA3AFB66" box="[151,346,1200,1223]" italics="true" pageId="1" pageNumber="56">Barbus amphibius</emphasis>
</taxonomicName>
(non
<bibRefCitation id="37654B5E271CC0010CF43BFCA908FB66" author="Valenciennes" box="[408,616,1200,1223]" pageId="1" pageNumber="64" refString="Cuvier, G. &amp; Valenciennes, A. (1842) Histoire naturelle des poissons, vol. 16. P. Bertrand, Paris, xx + 472 pp., pls 465 - 487." type="book chapter" year="1842">Valenciennes, 1842</bibRefCitation>
)
<treatmentCitation id="D25510BE271CC0010F1A3BFCA82DFB66" author="Duncker" box="[630,845,1200,1223]" page="262" pageId="1" pageNumber="56" year="1912">
<bibRefCitation id="37654B5E271CC0010F1A3BFCA877FB66" author="Duncker" box="[630,791,1200,1223]" pageId="1" pageNumber="64" refString="Duncker, G. (1912) Die Susswasserfische Ceylons. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 241 - 272." type="journal article" year="1912">Duncker, 1912</bibRefCitation>
: 262
</treatmentCitation>
;
<treatmentCitation id="D25510BE271CC0010E343BFCAF23FB66" author="Senanayake" box="[856,1091,1200,1223]" page="84" pageId="1" pageNumber="56" year="1980">
<bibRefCitation id="37654B5E271CC0010E343BFCAF7DFB66" author="Senanayake" box="[856,1053,1200,1223]" pageId="1" pageNumber="64" refString="Senanayake, F. R. (1980) The biogeography and ecology of the inland fishes of Sri Lanka. Unpublished Ph. D. dissertation, Department of Wildlife and Fisheries Biology, University of California, Davis, 421 pp." type="book" year="1980">Senanayake, 1980</bibRefCitation>
: 84
</treatmentCitation>
(in part).
<taxonomicName id="94F44D2C271CC0010DFB3B9CAA00FB46" ID-CoL="4QHZ5" box="[151,352,1232,1255]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="amphibius">
<emphasis id="6180EABD271CC0010DFB3B9CAA00FB46" box="[151,352,1232,1255]" italics="true" pageId="1" pageNumber="56">Puntius amphibius</emphasis>
</taxonomicName>
(non
<bibRefCitation id="37654B5E271CC0010CCE3B9CA915FB46" author="Valenciennes" box="[418,629,1232,1255]" pageId="1" pageNumber="64" refString="Cuvier, G. &amp; Valenciennes, A. (1842) Histoire naturelle des poissons, vol. 16. P. Bertrand, Paris, xx + 472 pp., pls 465 - 487." type="book chapter" year="1842">Valenciennes, 1842</bibRefCitation>
)
<treatmentCitation id="D25510BE271CC0010FE83B9CA81DFB46" author="Deraniyagala" box="[644,893,1232,1255]" page="15" pageId="1" pageNumber="56" year="1930">
<bibRefCitation id="37654B5E271CC0010FE83B9CA834FB46" author="Deraniyagala" box="[644,852,1232,1255]" pageId="1" pageNumber="64" refString="Deraniyagala, P. E. P. (1930) The Eventognathi of Ceylon. Spolia Zeylanica, 16, 1 - 41." type="journal article" year="1930">Deraniyagala 1930</bibRefCitation>
: 15
</treatmentCitation>
;
<treatmentCitation id="D25510BE271CC0010EE73B9CAF20FB46" author="Herre" box="[907,1088,1232,1255]" page="175" pageId="1" pageNumber="56" year="1946">
<bibRefCitation id="37654B5E271CC0010EE73B9CAF69FB46" author="Herre" box="[907,1033,1232,1255]" pageId="1" pageNumber="64" refString="Herre, A. W. C. T. (1946) Fishes collected in Ceylon in April, 1934. Spolia Zeylanica, 24, 173 - 179." type="journal article" year="1946">Herre 1946</bibRefCitation>
: 175
</treatmentCitation>
;
<treatmentCitation id="D25510BE271CC00109213B9CAE27FB46" author="Deraniyagala" box="[1101,1351,1232,1255]" page="28" pageId="1" pageNumber="56" year="1952">
<bibRefCitation id="37654B5E271CC00109213B9CAE7DFB46" author="Deraniyagala" box="[1101,1309,1232,1255]" pageId="1" pageNumber="64" refString="Deraniyagala, P. E. P. (1952) A coloured atlas of some vertebrates from Ceylon. Vol. 1: Fishes. National Museums of Ceylon, Colombo, 149 pp., 34 pls." type="book" year="1952">Deraniyagala 1952</bibRefCitation>
: 28
</treatmentCitation>
;
<treatmentCitation id="D25510BE271CC001083F3B9CAA48FAA6" author="Munro" page="44" pageId="1" pageNumber="56" year="1955">
<bibRefCitation id="37654B5E271CC001083F3B9CAA60FAA6" author="Munro" pageId="1" pageNumber="64" refString="Munro, I. S. R. (1955) The Marine and Freshwater Fishes of Ceylon. Department of External Affairs, Canberra, 349 pp." type="book" year="1955">Munro 1955</bibRefCitation>
: 44
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BEE6524271CC0010DFB3A7EAE74F9C5" pageId="1" pageNumber="56" type="materials_examined">
<paragraph id="534B36AF271CC0010DFB3A7EAE66FAD5" blockId="1.[151,1437,1330,2036]" pageId="1" pageNumber="56">
<emphasis id="6180EABD271CC0010DFB3A7EAA31FAED" bold="true" box="[151,337,1330,1356]" pageId="1" pageNumber="56">
<typeStatus id="8C4F880D271CC0010DFB3A7EABB4FAED" box="[151,212,1330,1356]" pageId="1" pageNumber="56">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8C4F880D271CC0010C313A7EAAAFFAED" box="[349,463,1330,1356]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
: WHT 7639,
<quantity id="940C9B4A271CC0010F163A7EA987FAED" box="[634,743,1330,1356]" metricMagnitude="-2" metricUnit="m" metricValue="5.02" pageId="1" pageNumber="56" unit="mm" value="50.2">50.2 mm</quantity>
SL,
<collectingCountry id="2BE3763F271CC0010E443A7EA8C2FAED" box="[808,930,1330,1356]" name="Sri Lanka" pageId="1" pageNumber="56">Sri Lanka</collectingCountry>
, Kalu River at Walandure near Kuruwita (
<geoCoordinate id="36C05068271CC0010DCC3A16AB9AFAD5" box="[160,250,1370,1396]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="6.7666664">6°46’N</geoCoordinate>
,
<geoCoordinate id="36C05068271CC0010C6A3A16AA12FAD5" box="[262,370,1370,1396]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="80.38333">80°23’ E</geoCoordinate>
), elevation
<quantity id="940C9B4A271CC0010C963A16A927FAD5" box="[506,583,1370,1396]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="56" unit="m" value="120.0">120 m</quantity>
; coll. K. Manamendra-Arachchi &amp; M. M. Bahir,
<date id="274A106F271CC00109EB3A16AE62FAD5" box="[1159,1282,1370,1396]" pageId="1" pageNumber="56" value="1998-01-05">5 Jan1998</date>
.
</paragraph>
<paragraph id="534B36AF271CC0010DA93ACEAE74F9C5" blockId="1.[151,1437,1330,2036]" pageId="1" pageNumber="56">
<typeStatus id="8C4F880D271CC0010DA93ACEAA5BFA3D" box="[197,315,1410,1436]" pageId="1" pageNumber="56" type="paratype">Paratypes</typeStatus>
: WHT 7640, 11 ex.,
<quantity id="940C9B4A271CC0010F2D3ACEA98FFA3D" box="[577,751,1410,1436]" metricMagnitude="-2" metricUnit="m" metricValue="4.734999999999999" metricValueMax="5.54" metricValueMin="3.9299999999999997" pageId="1" pageNumber="56" unit="mm" value="47.349999999999994" valueMax="55.4" valueMin="39.3">39.3–55.4 mm</quantity>
SL,
<collectingCountry id="2BE3763F271CC0010E423ACEA8C8FA3D" box="[814,936,1410,1436]" name="Sri Lanka" pageId="1" pageNumber="56">Sri Lanka</collectingCountry>
, Kalu River at Walandure near Kuruwita (
<geoCoordinate id="36C05068271CC0010DCC3AE6AB9AFA65" box="[160,250,1450,1476]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="6.7666664">6°46’N</geoCoordinate>
,
<geoCoordinate id="36C05068271CC0010C693AE6AA11FA65" box="[261,369,1450,1476]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="80.38333">80°23’ E</geoCoordinate>
), elevation
<quantity id="940C9B4A271CC0010C953AE6A926FA65" box="[505,582,1450,1476]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="56" unit="m" value="120.0">120 m</quantity>
; coll. K. Manamendra-Arachchi &amp; M. M. Bahir,
<date id="274A106F271CC00109E83AE6AE65FA65" box="[1156,1285,1450,1476]" pageId="1" pageNumber="56" value="1998-01-05">5 Jan 1998</date>
. WHT 7641, 2 ex.,
<quantity id="940C9B4A271CC0010DB23A9EAA29FA4D" box="[222,329,1490,1516]" metricMagnitude="-2" metricUnit="m" metricValue="4.42" pageId="1" pageNumber="56" unit="mm" value="44.2">44.2 mm</quantity>
,
<quantity id="940C9B4A271CC0010C383A9EAADDFA4D" box="[340,445,1490,1516]" metricMagnitude="-2" metricUnit="m" metricValue="5.81" pageId="1" pageNumber="56" unit="mm" value="58.1">58.1 mm</quantity>
SL,
<collectingCountry id="2BE3763F271CC0010C983A9EA909FA4D" box="[500,617,1490,1516]" name="Sri Lanka" pageId="1" pageNumber="56">Sri Lanka</collectingCountry>
, Gin River at Wakwella near Galle (
<geoCoordinate id="36C05068271CC001097C3A9EAF0AFA4D" box="[1040,1130,1490,1516]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="6.1">6°06’N</geoCoordinate>
,
<geoCoordinate id="36C05068271CC00109193A9EAF80FA4D" box="[1141,1248,1490,1516]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="80.183334">80°11’ E</geoCoordinate>
), elevation
<quantity id="940C9B4A271CC001080B3A9EAEF7FA4D" box="[1383,1431,1490,1516]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="56" unit="m" value="5.0">5 m</quantity>
; coll. M. M. Bahir &amp; M. de Silva,
<date id="274A106F271CC0010F443AB6A9A3F9B5" box="[552,707,1530,1556]" pageId="1" pageNumber="56" value="1997-12-26">26 Dec 1997</date>
. WHT 7642, 2 ex.,
<quantity id="940C9B4A271CC0010EC23AB6AF7BF9B5" box="[942,1051,1530,1556]" metricMagnitude="-2" metricUnit="m" metricValue="5.16" pageId="1" pageNumber="56" unit="mm" value="51.6">51.6 mm</quantity>
,
<quantity id="940C9B4A271CC001094B3AB6AFF1F9B5" box="[1063,1169,1530,1556]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" pageId="1" pageNumber="56" unit="mm" value="51.8">51.8 mm</quantity>
SL,
<collectingCountry id="2BE3763F271CC00109A63AB6AE21F9B5" box="[1226,1345,1530,1556]" name="Sri Lanka" pageId="1" pageNumber="56">Sri Lanka</collectingCountry>
, Kelani River near Gampaha
<geoCoordinate id="36C05068271CC0010CE3396EAA88F99D" box="[399,488,1570,1596]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="7.05">7°03’N</geoCoordinate>
,
<geoCoordinate id="36C05068271CC0010C98396EA939F99D" box="[500,601,1570,1596]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="80.01667">80°01’E</geoCoordinate>
, elevation
<quantity id="940C9B4A271CC0010FBB396EA875F99D" box="[727,789,1570,1596]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="56" unit="m" value="10.0">10 m</quantity>
; collector, date unknown. WHT 7643,
<quantity id="940C9B4A271CC00109B6396EAE23F99D" box="[1242,1347,1570,1596]" metricMagnitude="-2" metricUnit="m" metricValue="7.27" pageId="1" pageNumber="56" unit="mm" value="72.7">72.7 mm</quantity>
SL,
<collectingCountry id="2BE3763F271CC0010816396EAB83F9C5" name="Sri Lanka" pageId="1" pageNumber="56">Sri Lanka</collectingCountry>
, Attidiya near Colombo (
<geoCoordinate id="36C05068271CC0010F613906A907F9C5" box="[525,615,1610,1636]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="6.8333335">6°50’N</geoCoordinate>
,
<geoCoordinate id="36C05068271CC0010F1F3906A9B8F9C5" box="[627,728,1610,1636]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="79.88333">79°53’E</geoCoordinate>
), elevation
<quantity id="940C9B4A271CC0010E0C3906A8F0F9C5" box="[864,912,1610,1636]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="56" unit="m" value="8.0">8 m</quantity>
; coll. D. Gabadage,
<date id="274A106F271CC00109133906AE70F9C5" box="[1151,1296,1610,1636]" pageId="1" pageNumber="56" value="1993-06-11">11 Jun 1993</date>
.
</paragraph>
</subSubSection>
<subSubSection id="1BEE6524271CC0010DA9393EA9D9F855" pageId="1" pageNumber="56" type="diagnosis">
<paragraph id="534B36AF271CC0010DA9393EA9D9F855" blockId="1.[151,1437,1330,2036]" pageId="1" pageNumber="56">
<emphasis id="6180EABD271CC0010DA9393EAA25F92D" bold="true" box="[197,325,1650,1676]" pageId="1" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="94F44D2C271CC0010C3C393FA941F92D" box="[336,545,1651,1676]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD271CC0010C3C393FA941F92D" box="[336,545,1651,1676]" italics="true" pageId="1" pageNumber="56">Puntius kamalika</emphasis>
</taxonomicName>
is distinguished from all Sri Lankan and peninsular-Indian congeners by having ½4/1/2½ scales in transverse line between mid-dorsal scale row and pelvic-fin origin and lacking any prominent markings on fins and body in both living and preserved examples. Additionally, it is distinguished from the Sri Lankan and peninsular Indian species of
<taxonomicName id="94F44D2C271CC0010E6639A7A804F8A5" box="[778,868,1771,1796]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="6180EABD271CC0010E6639A7A804F8A5" box="[778,868,1771,1796]" italics="true" pageId="1" pageNumber="56">Puntius</emphasis>
</taxonomicName>
that most closely resemble it as follows: from
<taxonomicName id="94F44D2C271CC00108EA39A7AA71F88D" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" sensu="stricto" species="amphibius">
<emphasis id="6180EABD271CC00108EA39A7AA71F88D" italics="true" pageId="1" pageNumber="56">P. amphibius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AB357C6271CC0010C75385EAACBF88D" box="[281,427,1810,1836]" pageId="1" pageNumber="56" sensu="stricto">sensu stricto</taxonomicNameLabel>
by having the head length 28.8–31.2% (
<emphasis id="6180EABD271CC0010EE0385FA8C4F88D" box="[908,932,1811,1836]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
26.4–28.3%) SL; eye diameter 7.7–10.2% (
<emphasis id="6180EABD271CC0010DCD3877ABD9F8F5" box="[161,185,1851,1876]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
6.6–7.3%) SL; maxillary barbel 3.7–5.3% (
<emphasis id="6180EABD271CC0010FD03877A9B4F8F5" box="[700,724,1851,1876]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
3.1%) SL; and 16+14 vertebrae (
<emphasis id="6180EABD271CC00109313877AF16F8F5" box="[1117,1142,1851,1876]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
17+14); from
<taxonomicName id="94F44D2C271CC001084D3877AEFBF8F5" box="[1313,1435,1851,1876]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="6180EABD271CC001084D3877AEFBF8F5" box="[1313,1435,1851,1876]" italics="true" pageId="1" pageNumber="56">P. dorsalis</emphasis>
</taxonomicName>
by its smaller size (maximum standard length
<quantity id="940C9B4A271CC0010FD3382EA876F8DD" box="[703,790,1890,1916]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="56" unit="mm" value="72.0">72 mm</quantity>
,
<emphasis id="6180EABD271CC0010E4F382FA85BF8DD" box="[803,827,1891,1916]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
<quantity id="940C9B4A271CC0010E28382EA8CCF8DD" box="[836,940,1890,1916]" metricMagnitude="-1" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="56" unit="mm" value="133.0">133 mm</quantity>
); and lacking any prominent markings on fins and body (
<emphasis id="6180EABD271CC0010C2438C7AA01F805" box="[328,353,1931,1956]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
black blotches on base of dorsal and caudal fins); and from
<taxonomicName id="94F44D2C271CC001094838C7AFD3F805" box="[1060,1203,1931,1956]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="mahecola">
<emphasis id="6180EABD271CC001094838C7AFD3F805" box="[1060,1203,1931,1956]" italics="true" pageId="1" pageNumber="56">P. mahecola</emphasis>
</taxonomicName>
by its smaller maximum size (up to
<quantity id="940C9B4A271CC0010C0738FEAAA0F86D" box="[363,448,1970,1996]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="56" unit="mm" value="73.0">73 mm</quantity>
SL,
<emphasis id="6180EABD271CC0010C9038FFA975F86D" box="[508,533,1971,1996]" italics="true" pageId="1" pageNumber="56">vs</emphasis>
<quantity id="940C9B4A271CC0010F7138FEA912F86D" box="[541,626,1970,1996]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="1" pageNumber="56" unit="mm" value="89.0">89 mm</quantity>
SL in
<taxonomicName id="94F44D2C271CC0010FAA38FFA837F86D" box="[710,855,1971,1996]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="mahecola">
<emphasis id="6180EABD271CC0010FAA38FFA837F86D" box="[710,855,1971,1996]" italics="true" pageId="1" pageNumber="56">P. mahecola</emphasis>
</taxonomicName>
), and by lacking a black blotch, larger than eye, across about 3½ scales of the caudal peduncle.
</paragraph>
</subSubSection>
<subSubSection id="1BEE6524271FC0050DA93FDBAE44FE8B" lastPageId="5" lastPageNumber="60" pageId="2" pageNumber="57" type="description">
<paragraph id="534B36AF271FC0020DA93FDBA831FEF3" blockId="2.[151,1436,151,338]" pageId="2" pageNumber="57">
<emphasis id="6180EABD271FC0020DA93FDBAA3DFF10" bold="true" box="[197,349,151,177]" pageId="2" pageNumber="57">Description.</emphasis>
General body shape and appearance are shown in
<figureCitation id="CBCF2A2A271FC0020EDB3FD4AF7CFF13" box="[951,1052,152,178]" captionStart="FIGURE 1" captionStartId="3.[151,255,1516,1540]" captionTargetBox="[280,1307,198,1478]" captionTargetId="figure@3.[280,1307,194,1012]" captionTargetPageId="3" captionText="FIGURE 1. Puntius kamalika. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo." httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="2" pageNumber="57">Figure 1</figureCitation>
. Morphometric data of
<typeStatus id="8C4F880D271FC002085A3FD4AEFCFF13" box="[1334,1436,152,178]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
and 17
<typeStatus id="8C4F880D271FC0020D9F3F8CAA03FF7B" box="[243,355,192,218]" pageId="2" pageNumber="57" type="paratype">paratypes</typeStatus>
are given in
<tableCitation id="1E760314271FC0020F653F8CA904FF7B" box="[521,612,192,218]" captionStart="TABLE 1" captionStartId="2.[151,239,426,450]" captionTargetBox="[159,1272,517,1607]" captionTargetPageId="2" captionText="TABLE 1. Measurements (as a percentage of standard length) of the holotype (WHT 7639, 50.2 mm SL) and 16 paratypes (WHT 7640, WHT 7641, WHT 7642, WHT 7643, 39.3 – 72.7 mm SL) of Puntius kamalika." httpUri="http://table.plazi.org/id/078B6627271FC0020DFB3EE6AFB2FE45" pageId="2" pageNumber="57" tableUuid="078B6627271FC0020DFB3EE6AFB2FE45">Table 1</tableCitation>
. Body elongate, laterally compressed; pre-dorsal contour ascending, convex, with a slight depression above first pre-dorsal scale. Ventral contour convex anterior to origin of caudal peduncle. Dorsal and ventral profiles of caudal peduncle concave to terminal scales on tail base. Caudal peduncle length 1.1–1.4 (mean 1.2, SD 0.1) times its depth.
</paragraph>
<caption id="078B6627271FC0020DFB3EE6AFB2FE45" ID-Table-UUID="078B6627271FC0020DFB3EE6AFB2FE45" httpUri="http://table.plazi.org/id/078B6627271FC0020DFB3EE6AFB2FE45" pageId="2" pageNumber="57" targetBox="[159,1272,517,1607]" targetIsTable="true" targetPageId="2">
<paragraph id="534B36AF271FC0020DFB3EE6AFB2FE45" blockId="2.[151,1436,426,484]" pageId="2" pageNumber="57">
<emphasis id="6180EABD271FC0020DFB3EE6AA6DFE63" bold="true" box="[151,269,426,450]" pageId="2" pageNumber="57">TABLE 1.</emphasis>
Measurements (as a percentage of standard length) of the holotype (WHT 7639, 50.2 mm SL) and 16 paratypes (WHT 7640, WHT 7641, WHT 7642, WHT 7643, 39.3–72.7 mm SL) of
<taxonomicName id="94F44D2C271FC002097D3E81AFABFE45" box="[1041,1227,461,484]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="57" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD271FC002097D3E81AFABFE45" box="[1041,1227,461,484]" italics="true" pageId="2" pageNumber="57">Puntius kamalika</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="534B36AF271FC0020FF53D49AF95F9E6" pageId="2" pageNumber="57">
<table id="21F4C40F271F3FFF0DF33D49AF98F9E6" box="[159,1272,517,1607]" gridcols="4" gridrows="26" pageId="2" pageNumber="57">
<tr id="EDC434ED271F3FFF0DF33D49AF98FDBD" box="[159,1272,517,540]" gridrow="0" pageId="2" pageNumber="57" rowspan-0="1">
<th id="AE155D91271F3FFF0FF43D49A87EFDBD" box="[664,798,517,540]" gridcol="1" gridrow="0" pageId="2" pageNumber="57">range</th>
<th id="AE155D91271F3FFF0EB03D49AF78FDBD" box="[988,1048,517,540]" gridcol="2" gridrow="0" pageId="2" pageNumber="57">mean</th>
<th id="AE155D91271F3FFF09BF3D49AF98FDBD" box="[1235,1272,517,540]" gridcol="3" gridrow="0" pageId="2" pageNumber="57">s.d.</th>
</tr>
<tr id="EDC434ED271F3FFF0DF33D7CAF98FDE6" box="[159,1272,560,583]" gridrow="1" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33D7CAAACFDE6" box="[159,460,560,583]" gridcol="0" gridrow="1" pageId="2" pageNumber="57">Total length</th>
<td id="AE155D91271F3FFF0FF43D7CA87EFDE6" box="[664,798,560,583]" gridcol="1" gridrow="1" pageId="2" pageNumber="57">125.8–133.9</td>
<td id="AE155D91271F3FFF0EB03D7CAF78FDE6" box="[988,1048,560,583]" gridcol="2" gridrow="1" pageId="2" pageNumber="57">129.9</td>
<td id="AE155D91271F3FFF09BF3D7CAF98FDE6" box="[1235,1272,560,583]" gridcol="3" gridrow="1" pageId="2" pageNumber="57">2.0</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33D16AF98FDD0" box="[159,1272,602,625]" gridrow="2" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33D16AAACFDD0" box="[159,460,602,625]" gridcol="0" gridrow="2" pageId="2" pageNumber="57">Predorsal length</th>
<td id="AE155D91271F3FFF0FF43D16A87EFDD0" box="[664,798,602,625]" gridcol="1" gridrow="2" pageId="2" pageNumber="57">48.8–53.2</td>
<td id="AE155D91271F3FFF0EB03D16AF78FDD0" box="[988,1048,602,625]" gridcol="2" gridrow="2" pageId="2" pageNumber="57">51.5</td>
<td id="AE155D91271F3FFF09BF3D16AF98FDD0" box="[1235,1272,602,625]" gridcol="3" gridrow="2" pageId="2" pageNumber="57">1.3</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33DC9AF98FD3D" box="[159,1272,645,668]" gridrow="3" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33DC9AAACFD3D" box="[159,460,645,668]" gridcol="0" gridrow="3" pageId="2" pageNumber="57">Post-dorsal length</th>
<td id="AE155D91271F3FFF0FF43DC9A87EFD3D" box="[664,798,645,668]" gridcol="1" gridrow="3" pageId="2" pageNumber="57">48.9–57.5</td>
<td id="AE155D91271F3FFF0EB03DC9AF78FD3D" box="[988,1048,645,668]" gridcol="2" gridrow="3" pageId="2" pageNumber="57">52.6</td>
<td id="AE155D91271F3FFF09BF3DC9AF98FD3D" box="[1235,1272,645,668]" gridcol="3" gridrow="3" pageId="2" pageNumber="57">2.2</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33DFCAF98FD66" box="[159,1272,688,711]" gridrow="4" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33DFCAAACFD66" box="[159,460,688,711]" gridcol="0" gridrow="4" pageId="2" pageNumber="57">Preanal length</th>
<td id="AE155D91271F3FFF0FF43DFCA87EFD66" box="[664,798,688,711]" gridcol="1" gridrow="4" pageId="2" pageNumber="57">70.9–76.2</td>
<td id="AE155D91271F3FFF0EB03DFCAF78FD66" box="[988,1048,688,711]" gridcol="2" gridrow="4" pageId="2" pageNumber="57">74.0</td>
<td id="AE155D91271F3FFF09BF3DFCAF98FD66" box="[1235,1272,688,711]" gridcol="3" gridrow="4" pageId="2" pageNumber="57">1.3</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33D96AF98FD50" box="[159,1272,730,753]" gridrow="5" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33D96AAACFD50" box="[159,460,730,753]" gridcol="0" gridrow="5" pageId="2" pageNumber="57">Prepelvic length</th>
<td id="AE155D91271F3FFF0FF43D96A87EFD50" box="[664,798,730,753]" gridcol="1" gridrow="5" pageId="2" pageNumber="57">49.1–52.2</td>
<td id="AE155D91271F3FFF0EB03D96AF78FD50" box="[988,1048,730,753]" gridcol="2" gridrow="5" pageId="2" pageNumber="57">50.6</td>
<td id="AE155D91271F3FFF09BF3D96AF98FD50" box="[1235,1272,730,753]" gridcol="3" gridrow="5" pageId="2" pageNumber="57">1.0</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33C49AF98FCBD" box="[159,1272,773,796]" gridrow="6" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33C49AAACFCBD" box="[159,460,773,796]" gridcol="0" gridrow="6" pageId="2" pageNumber="57">Caudal peduncle length</th>
<td id="AE155D91271F3FFF0FF43C49A87EFCBD" box="[664,798,773,796]" gridcol="1" gridrow="6" pageId="2" pageNumber="57">15.9–18.7</td>
<td id="AE155D91271F3FFF0EB03C49AF78FCBD" box="[988,1048,773,796]" gridcol="2" gridrow="6" pageId="2" pageNumber="57">17.6</td>
<td id="AE155D91271F3FFF09BF3C49AF98FCBD" box="[1235,1272,773,796]" gridcol="3" gridrow="6" pageId="2" pageNumber="57">0.9</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33C7CAF98FCE6" box="[159,1272,816,839]" gridrow="7" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33C7CAAACFCE6" box="[159,460,816,839]" gridcol="0" gridrow="7" pageId="2" pageNumber="57">Caudal peduncle depth</th>
<td id="AE155D91271F3FFF0FF43C7CA87EFCE6" box="[664,798,816,839]" gridcol="1" gridrow="7" pageId="2" pageNumber="57">13.5–15.5</td>
<td id="AE155D91271F3FFF0EB03C7CAF78FCE6" box="[988,1048,816,839]" gridcol="2" gridrow="7" pageId="2" pageNumber="57">14.2</td>
<td id="AE155D91271F3FFF09BF3C7CAF98FCE6" box="[1235,1272,816,839]" gridcol="3" gridrow="7" pageId="2" pageNumber="57">0.6</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33C16AF98FCD0" box="[159,1272,858,881]" gridrow="8" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33C16AAACFCD0" box="[159,460,858,881]" gridcol="0" gridrow="8" pageId="2" pageNumber="57">Head length</th>
<td id="AE155D91271F3FFF0FF43C16A87EFCD0" box="[664,798,858,881]" gridcol="1" gridrow="8" pageId="2" pageNumber="57">28.8–31.2</td>
<td id="AE155D91271F3FFF0EB03C16AF78FCD0" box="[988,1048,858,881]" gridcol="2" gridrow="8" pageId="2" pageNumber="57">29.7</td>
<td id="AE155D91271F3FFF09BF3C16AF98FCD0" box="[1235,1272,858,881]" gridcol="3" gridrow="8" pageId="2" pageNumber="57">0.7</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33CC9AF98FC3D" box="[159,1272,901,924]" gridrow="9" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33CC9AAACFC3D" box="[159,460,901,924]" gridcol="0" gridrow="9" pageId="2" pageNumber="57">Head depth</th>
<td id="AE155D91271F3FFF0FF43CC9A87EFC3D" box="[664,798,901,924]" gridcol="1" gridrow="9" pageId="2" pageNumber="57">16.6–19.0</td>
<td id="AE155D91271F3FFF0EB03CC9AF78FC3D" box="[988,1048,901,924]" gridcol="2" gridrow="9" pageId="2" pageNumber="57">18.0</td>
<td id="AE155D91271F3FFF09BF3CC9AF98FC3D" box="[1235,1272,901,924]" gridcol="3" gridrow="9" pageId="2" pageNumber="57">0.7</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33CFCAF98FC66" box="[159,1272,944,967]" gridrow="10" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33CFCAAACFC66" box="[159,460,944,967]" gridcol="0" gridrow="10" pageId="2" pageNumber="57">Snout length</th>
<td id="AE155D91271F3FFF0FF43CFCA87EFC66" box="[664,798,944,967]" gridcol="1" gridrow="10" pageId="2" pageNumber="57">7.7–9.5</td>
<td id="AE155D91271F3FFF0EB03CFCAF78FC66" box="[988,1048,944,967]" gridcol="2" gridrow="10" pageId="2" pageNumber="57">8.3</td>
<td id="AE155D91271F3FFF09BF3CFCAF98FC66" box="[1235,1272,944,967]" gridcol="3" gridrow="10" pageId="2" pageNumber="57">0.6</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33C96AF98FC50" box="[159,1272,986,1009]" gridrow="11" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33C96AAACFC50" box="[159,460,986,1009]" gridcol="0" gridrow="11" pageId="2" pageNumber="57">Eye diameter</th>
<td id="AE155D91271F3FFF0FF43C96A87EFC50" box="[664,798,986,1009]" gridcol="1" gridrow="11" pageId="2" pageNumber="57">7.7–10.3</td>
<td id="AE155D91271F3FFF0EB03C96AF78FC50" box="[988,1048,986,1009]" gridcol="2" gridrow="11" pageId="2" pageNumber="57">9.1</td>
<td id="AE155D91271F3FFF09BF3C96AF98FC50" box="[1235,1272,986,1009]" gridcol="3" gridrow="11" pageId="2" pageNumber="57">0.7</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33B49AF98FBBD" box="[159,1272,1029,1052]" gridrow="12" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33B49AAACFBBD" box="[159,460,1029,1052]" gridcol="0" gridrow="12" pageId="2" pageNumber="57">Interorbital width</th>
<td id="AE155D91271F3FFF0FF43B49A87EFBBD" box="[664,798,1029,1052]" gridcol="1" gridrow="12" pageId="2" pageNumber="57">9.9–12.9</td>
<td id="AE155D91271F3FFF0EB03B49AF78FBBD" box="[988,1048,1029,1052]" gridcol="2" gridrow="12" pageId="2" pageNumber="57">11.2</td>
<td id="AE155D91271F3FFF09BF3B49AF98FBBD" box="[1235,1272,1029,1052]" gridcol="3" gridrow="12" pageId="2" pageNumber="57">0.7</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33B7CAF98FBE6" box="[159,1272,1072,1095]" gridrow="13" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33B7CAAACFBE6" box="[159,460,1072,1095]" gridcol="0" gridrow="13" pageId="2" pageNumber="57">Maxillary barbel length</th>
<td id="AE155D91271F3FFF0FF43B7CA87EFBE6" box="[664,798,1072,1095]" gridcol="1" gridrow="13" pageId="2" pageNumber="57">3.7–5.3</td>
<td id="AE155D91271F3FFF0EB03B7CAF78FBE6" box="[988,1048,1072,1095]" gridcol="2" gridrow="13" pageId="2" pageNumber="57">4.4</td>
<td id="AE155D91271F3FFF09BF3B7CAF98FBE6" box="[1235,1272,1072,1095]" gridcol="3" gridrow="13" pageId="2" pageNumber="57">0.5</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33B16AF98FBD0" box="[159,1272,1114,1137]" gridrow="14" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33B16AAACFBD0" box="[159,460,1114,1137]" gridcol="0" gridrow="14" pageId="2" pageNumber="57">Body depth</th>
<td id="AE155D91271F3FFF0FF43B16A87EFBD0" box="[664,798,1114,1137]" gridcol="1" gridrow="14" pageId="2" pageNumber="57">28.0–33.6</td>
<td id="AE155D91271F3FFF0EB03B16AF78FBD0" box="[988,1048,1114,1137]" gridcol="2" gridrow="14" pageId="2" pageNumber="57">30.9</td>
<td id="AE155D91271F3FFF09BF3B16AF98FBD0" box="[1235,1272,1114,1137]" gridcol="3" gridrow="14" pageId="2" pageNumber="57">1.6</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33BC9AF98FB3D" box="[159,1272,1157,1180]" gridrow="15" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33BC9AAACFB3D" box="[159,460,1157,1180]" gridcol="0" gridrow="15" pageId="2" pageNumber="57">Dorsal-fin height</th>
<td id="AE155D91271F3FFF0FF43BC9A87EFB3D" box="[664,798,1157,1180]" gridcol="1" gridrow="15" pageId="2" pageNumber="57">26.7–32.1</td>
<td id="AE155D91271F3FFF0EB03BC9AF78FB3D" box="[988,1048,1157,1180]" gridcol="2" gridrow="15" pageId="2" pageNumber="57">28.2</td>
<td id="AE155D91271F3FFF09BF3BC9AF98FB3D" box="[1235,1272,1157,1180]" gridcol="3" gridrow="15" pageId="2" pageNumber="57">1.4</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33BFCAF98FB66" box="[159,1272,1200,1223]" gridrow="16" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33BFCAAACFB66" box="[159,460,1200,1223]" gridcol="0" gridrow="16" pageId="2" pageNumber="57">Dorsal-fin breadth</th>
<td id="AE155D91271F3FFF0FF43BFCA87EFB66" box="[664,798,1200,1223]" gridcol="1" gridrow="16" pageId="2" pageNumber="57">14.6–16.1</td>
<td id="AE155D91271F3FFF0EB03BFCAF78FB66" box="[988,1048,1200,1223]" gridcol="2" gridrow="16" pageId="2" pageNumber="57">15.7</td>
<td id="AE155D91271F3FFF09BF3BFCAF98FB66" box="[1235,1272,1200,1223]" gridcol="3" gridrow="16" pageId="2" pageNumber="57">0.5</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33B96AF98FB50" box="[159,1272,1242,1265]" gridrow="17" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33B96AAACFB50" box="[159,460,1242,1265]" gridcol="0" gridrow="17" pageId="2" pageNumber="57">Anal-fin depth</th>
<td id="AE155D91271F3FFF0FF43B96A87EFB50" box="[664,798,1242,1265]" gridcol="1" gridrow="17" pageId="2" pageNumber="57">17.5–19.9</td>
<td id="AE155D91271F3FFF0EB03B96AF78FB50" box="[988,1048,1242,1265]" gridcol="2" gridrow="17" pageId="2" pageNumber="57">18.4</td>
<td id="AE155D91271F3FFF09BF3B96AF98FB50" box="[1235,1272,1242,1265]" gridcol="3" gridrow="17" pageId="2" pageNumber="57">0.8</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33A49AF98FABD" box="[159,1272,1285,1308]" gridrow="18" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33A49AAACFABD" box="[159,460,1285,1308]" gridcol="0" gridrow="18" pageId="2" pageNumber="57">Anal-fin breadth</th>
<td id="AE155D91271F3FFF0FF43A49A87EFABD" box="[664,798,1285,1308]" gridcol="1" gridrow="18" pageId="2" pageNumber="57">8.3–10.7</td>
<td id="AE155D91271F3FFF0EB03A49AF78FABD" box="[988,1048,1285,1308]" gridcol="2" gridrow="18" pageId="2" pageNumber="57">29.5</td>
<td id="AE155D91271F3FFF09BF3A49AF98FABD" box="[1235,1272,1285,1308]" gridcol="3" gridrow="18" pageId="2" pageNumber="57">0.6</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33A7CAF98FAE6" box="[159,1272,1328,1351]" gridrow="19" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33A7CAAACFAE6" box="[159,460,1328,1351]" gridcol="0" gridrow="19" pageId="2" pageNumber="57">Pectoral-fin length</th>
<td id="AE155D91271F3FFF0FF43A7CA87EFAE6" box="[664,798,1328,1351]" gridcol="1" gridrow="19" pageId="2" pageNumber="57">19.3–22.5</td>
<td id="AE155D91271F3FFF0EB03A7CAF78FAE6" box="[988,1048,1328,1351]" gridcol="2" gridrow="19" pageId="2" pageNumber="57">21.0</td>
<td id="AE155D91271F3FFF09BF3A7CAF98FAE6" box="[1235,1272,1328,1351]" gridcol="3" gridrow="19" pageId="2" pageNumber="57">0.9</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33A16AF98FAD0" box="[159,1272,1370,1393]" gridrow="20" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33A16AAACFAD0" box="[159,460,1370,1393]" gridcol="0" gridrow="20" pageId="2" pageNumber="57">Pelvic-fin length</th>
<td id="AE155D91271F3FFF0FF43A16A87EFAD0" box="[664,798,1370,1393]" gridcol="1" gridrow="20" pageId="2" pageNumber="57">18.2–20.6</td>
<td id="AE155D91271F3FFF0EB03A16AF78FAD0" box="[988,1048,1370,1393]" gridcol="2" gridrow="20" pageId="2" pageNumber="57">19.2</td>
<td id="AE155D91271F3FFF09BF3A16AF98FAD0" box="[1235,1272,1370,1393]" gridcol="3" gridrow="20" pageId="2" pageNumber="57">0.7</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33AC9AF98FA3D" box="[159,1272,1413,1436]" gridrow="21" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33AC9AAACFA3D" box="[159,460,1413,1436]" gridcol="0" gridrow="21" pageId="2" pageNumber="57">Length of upper caudal lobe</th>
<td id="AE155D91271F3FFF0FF43AC9A87EFA3D" box="[664,798,1413,1436]" gridcol="1" gridrow="21" pageId="2" pageNumber="57">29.8–34.2</td>
<td id="AE155D91271F3FFF0EB03AC9AF78FA3D" box="[988,1048,1413,1436]" gridcol="2" gridrow="21" pageId="2" pageNumber="57">32.2</td>
<td id="AE155D91271F3FFF09BF3AC9AF98FA3D" box="[1235,1272,1413,1436]" gridcol="3" gridrow="21" pageId="2" pageNumber="57">1.5</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33AFCAF98FA66" box="[159,1272,1456,1479]" gridrow="22" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33AFCAAACFA66" box="[159,460,1456,1479]" gridcol="0" gridrow="22" pageId="2" pageNumber="57">Length of lower caudal lobe</th>
<td id="AE155D91271F3FFF0FF43AFCA87EFA66" box="[664,798,1456,1479]" gridcol="1" gridrow="22" pageId="2" pageNumber="57">28.5–34.5</td>
<td id="AE155D91271F3FFF0EB03AFCAF78FA66" box="[988,1048,1456,1479]" gridcol="2" gridrow="22" pageId="2" pageNumber="57">32.5</td>
<td id="AE155D91271F3FFF09BF3AFCAF98FA66" box="[1235,1272,1456,1479]" gridcol="3" gridrow="22" pageId="2" pageNumber="57">1.6</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33A96AF98FA50" box="[159,1272,1498,1521]" gridrow="23" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33A96AAACFA50" box="[159,460,1498,1521]" gridcol="0" gridrow="23" pageId="2" pageNumber="57">Medial length of caudal fin</th>
<td id="AE155D91271F3FFF0FF43A96A87EFA50" box="[664,798,1498,1521]" gridcol="1" gridrow="23" pageId="2" pageNumber="57">12.7–17.8</td>
<td id="AE155D91271F3FFF0EB03A96AF78FA50" box="[988,1048,1498,1521]" gridcol="2" gridrow="23" pageId="2" pageNumber="57">15.2</td>
<td id="AE155D91271F3FFF09BF3A96AF98FA50" box="[1235,1272,1498,1521]" gridcol="3" gridrow="23" pageId="2" pageNumber="57">1.5</td>
</tr>
<tr id="EDC434ED271F3FFF0DF33949AF98F9BD" box="[159,1272,1541,1564]" gridrow="24" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF33949AAACF9BD" box="[159,460,1541,1564]" gridcol="0" gridrow="24" pageId="2" pageNumber="57">Internarial width</th>
<td id="AE155D91271F3FFF0FF43949A87EF9BD" box="[664,798,1541,1564]" gridcol="1" gridrow="24" pageId="2" pageNumber="57">6.1–7.7</td>
<td id="AE155D91271F3FFF0EB03949AF78F9BD" box="[988,1048,1541,1564]" gridcol="2" gridrow="24" pageId="2" pageNumber="57">6.9</td>
<td id="AE155D91271F3FFF09BF3949AF98F9BD" box="[1235,1272,1541,1564]" gridcol="3" gridrow="24" pageId="2" pageNumber="57">0.5</td>
</tr>
<tr id="EDC434ED271F3FFF0DF3397CAF98F9E6" box="[159,1272,1584,1607]" gridrow="25" pageId="2" pageNumber="57">
<th id="AE155D91271F3FFF0DF3397CAAACF9E6" box="[159,460,1584,1607]" gridcol="0" gridrow="25" pageId="2" pageNumber="57">Post-orbital head length</th>
<td id="AE155D91271F3FFF0FF4397CA87EF9E6" box="[664,798,1584,1607]" gridcol="1" gridrow="25" pageId="2" pageNumber="57">12.5–13.8</td>
<td id="AE155D91271F3FFF0EB0397CAF78F9E6" box="[988,1048,1584,1607]" gridcol="2" gridrow="25" pageId="2" pageNumber="57">13.0</td>
<td id="AE155D91271F3FFF09BF397CAF98F9E6" box="[1235,1272,1584,1607]" gridcol="3" gridrow="25" pageId="2" pageNumber="57">0.4</td>
</tr>
</table>
</paragraph>
<paragraph id="534B36AF271FC0020DA9398CABA7F87E" blockId="2.[151,1437,1728,2015]" pageId="2" pageNumber="57">
Head slightly less than one-third of standard length, somewhat compressed laterally. Eye located slightly forward of centre, superiorly on head. Eyes just visible in ventral and dorsal views. Nares dorsolaterally orientated, a shallow transverse depression on snout immediately anterior to them, dividing dorsal profile of snout into two convex halves. Mouth U-shaped in ventral aspect, subterminal, small, angle of gape not reaching below anterior nares in lateral aspect. Lips fleshy, entire, adnate to jaws. Upper lip overhanging lower one; posterior margin of lower lip interrupted medially by isthmus. A single pair of maxillary barbels, their length slightly less than one-third eye diameter; no mandibular barbels. Fifth ceratobranchial with 5+3+2 teeth (
<figureCitation id="CBCF2A2A271FC002080138ECABC7F87E" captionStart="FIGURE 2" captionStartId="4.[151,255,1441,1465]" captionTargetBox="[277,1312,310,1394]" captionTargetId="figure@4.[266,1322,295,1416]" captionTargetPageId="4" captionText="FIGURE 2. Osteology of Puntius kamalika, 65.9 mm SL, WHT 7682 (left lateral view, except A, which is in left-medial view; and D and E, which are in dorsal view); A, maxilla; B, dentary and anguloarticular; C, fifth ceretobranchial; D, arrangement of premaxilla, maxilla and kinethmoid; E, cranium; F, caudal skeleton; G, circumorbital series; H, opercle (AA = anguloarticular; C 5 = fifth ceratobranchial; DN = dentary; DS = dermosphenotic; FO = fontanel; FR = frontal; HA = hypurapophysis; IO = interorbital; KE = kinethmoid; MX = maxillary; OP = operculum; PA = parietal; PH = parahypural; PM = premaxillary). Scale bars = 1 mm." httpUri="https://zenodo.org/record/183060/files/figure.png" pageId="2" pageNumber="57">Fig. 2</figureCitation>
F).
</paragraph>
<caption id="078B6627271EC0030DFB3AA0AA81F9E5" httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="3" pageNumber="58" targetBox="[280,1307,198,1478]" targetPageId="3">
<paragraph id="534B36AF271EC0030DFB3AA0AA81F9E5" blockId="3.[151,1436,1516,1604]" pageId="3" pageNumber="58">
<emphasis id="6180EABD271EC0030DFB3AA0AA7AF9A5" bold="true" box="[151,282,1516,1540]" pageId="3" pageNumber="58">FIGURE 1.</emphasis>
<taxonomicName id="94F44D2C271EC0030C4D3AA1AABCF9A5" box="[289,476,1517,1540]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="58" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD271EC0030C4D3AA1AABCF9A5" box="[289,476,1517,1540]" italics="true" pageId="3" pageNumber="58">Puntius kamalika</emphasis>
</taxonomicName>
. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo.
</paragraph>
</caption>
<paragraph id="534B36AF271EC0030DA9393CA877F803" blockId="3.[151,1437,1648,2034]" pageId="3" pageNumber="58">Pelvic-fin origin beneath dorsal-fin origin. Distal margin of dorsal fin slightly concave, that of anal fin straight. Distal margins of both pelvic and pectoral fins convex, rounded. Dorsal-fin origin above eighth scale of lateral line, its height 27.6–32.1% SL, with 4 unbranched rays (the last one smooth, strong) and 8 branched rays. Anal fin with 3 unbranched and 5 branched rays, its depth 17.5–19.9% SL. Posterior margins of dorsal and anal fins straight, their last rays, respectively, branched to base. Pelvic fin with one unbranched and eight branched rays, not reaching anus. Pectoral fin with one unbranched and 13 branched rays, not reaching pelvicfin origin. Caudal fin deeply forked, its lobes pointed, with 8 branched rays on upper lobe, 9 on lower lobe; 4 procurrent rays visible at top and bottom of caudal fin.</paragraph>
<paragraph id="534B36AF271EC0040DA938FCAFC4FEA3" blockId="3.[151,1437,1648,2034]" lastBlockId="4.[151,1436,152,258]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">Pored scales on lateral line 22 (1), 23 (12), 24 (5) on body, plus up to two on caudal-fin base; 4½ scales in transverse line between lateral line and origin of dorsal fin, 2½ between lateral line and origins of both pelvic and anal fins; ½2/1/2½ scales in transverse line on caudal peduncle; eight predorsal scales (excluding notched scale at dorsal-fin origin). An axillary scale approximately half length of pelvic fin extends backwards from pelvic-fin origin. Four scales on base of caudal fin. Dorsal-fin base sheathed by 7 scales.</paragraph>
<caption id="078B66272719C0040DFB3AEDA959F9C4" httpUri="https://zenodo.org/record/183060/files/figure.png" pageId="4" pageNumber="59" targetBox="[277,1312,310,1394]" targetPageId="4">
<paragraph id="534B36AF2719C0040DFB3AEDA959F9C4" blockId="4.[151,1436,1441,1637]" pageId="4" pageNumber="59">
<emphasis id="6180EABD2719C0040DFB3AEDAA78FA18" bold="true" box="[151,280,1441,1465]" pageId="4" pageNumber="59">FIGURE 2.</emphasis>
Osteology of
<taxonomicName id="94F44D2C2719C0040CC33AEDA908FA19" box="[431,616,1441,1464]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD2719C0040CC33AEDA908FA19" box="[431,616,1441,1464]" italics="true" pageId="4" pageNumber="59">Puntius kamalika</emphasis>
</taxonomicName>
, 65.9 mm SL, WHT 7682 (left lateral view, except A, which is in left-medial view; and D and E, which are in dorsal view); A, maxilla; B, dentary and anguloarticular; C, fifth ceretobranchial; D, arrangement of premaxilla, maxilla and kinethmoid; E, cranium; F, caudal skeleton; G, circumorbital series; H, opercle (AA= anguloarticular; C5= fifth ceratobranchial; DN= dentary; DS= dermosphenotic; FO=fontanel; FR=frontal; HA=hypurapophysis; IO=interorbital; KE=kinethmoid; MX= maxillary; OP=operculum; PA=parietal; PH=parahypural; PM=premaxillary). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="534B36AF2719C0040DA939D5AF12F8DA" blockId="4.[151,1436,1689,2038]" pageId="4" pageNumber="59">
Coloration. In life (
<figureCitation id="CBCF2A2A2719C0040CC639D5AA8EF912" box="[426,494,1689,1715]" captionStart="FIGURE 1" captionStartId="3.[151,255,1516,1540]" captionTargetBox="[280,1307,198,1478]" captionTargetId="figure@3.[280,1307,194,1012]" captionTargetPageId="3" captionText="FIGURE 1. Puntius kamalika. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo." httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
B), adults greenish-golden brown dorsally, with a faint, diffuse brownish lateral band about two scale-heights wide from top of operculum to base of caudal fin. Tail-base slightly darker than rest of body, but lacking melanophores. Body silvery-white ventrally. Fins faintly pinkish, except for caudal, which is hyaline. In preservative (
<figureCitation id="CBCF2A2A2719C0040F4F385DA904F88A" box="[547,612,1809,1835]" captionStart="FIGURE 1" captionStartId="3.[151,255,1516,1540]" captionTargetBox="[280,1307,198,1478]" captionTargetId="figure@3.[280,1307,194,1012]" captionTargetPageId="3" captionText="FIGURE 1. Puntius kamalika. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo." httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
A), head and body dorsally brownish olive, laterally yellowish brown; lower body, belly and chest pale cream-yellow. Fins hyaline. A faint darkening visible at caudal-fin base on some specimens (
<figureCitation id="CBCF2A2A2719C0040C0A382DAAC8F8DA" box="[358,424,1889,1915]" captionStart="FIGURE 1" captionStartId="3.[151,255,1516,1540]" captionTargetBox="[280,1307,198,1478]" captionTargetId="figure@3.[280,1307,194,1012]" captionTargetPageId="3" captionText="FIGURE 1. Puntius kamalika. A, holotype, WHT 7639, 50.2 mm SL, Sri Lanka, tributary of Kalu River at Walandure; B, living specimen, not preserved, same locality as holotype; C, paratype, WHT 7643, 72.7 mm SL, Sri Lanka, Attidiya- Bellanwila Wetland, Colombo." httpUri="https://zenodo.org/record/183059/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
A). Faint dark-grey speckles present on anal and caudal fins.
</paragraph>
<paragraph id="534B36AF2719C0040DA938C5AF2AF857" blockId="4.[151,1436,1689,2038]" pageId="4" pageNumber="59">
Distribution.
<taxonomicName id="94F44D2C2719C0040C0938C5A956F803" box="[357,566,1929,1954]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD2719C0040C0938C5A956F803" box="[357,566,1929,1954]" italics="true" pageId="4" pageNumber="59">Puntius kamalika</emphasis>
</taxonomicName>
is widely distributed in the wet-zone lowlands of
<collectingCountry id="2BE3763F2719C00409CD38C5AE77F802" box="[1185,1303,1929,1955]" name="Sri Lanka" pageId="4" pageNumber="59">Sri Lanka</collectingCountry>
(rainfall&gt;
<quantity id="940C9B4A2719C0040DFB38F8AA6EF86F" box="[151,270,1972,1998]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="59" unit="mm" value="2500.0">2,500 mm</quantity>
yr -1), where it occurs in streams, rivers and marshes between the Kelani-River basin draining to the island’s west, and the Gin River basin to the south (
<figureCitation id="CBCF2A2A2719C0040F9E3890A859F857" box="[754,825,2012,2038]" captionStart="FIGURE 3" captionStartId="5.[151,255,1050,1074]" captionTargetBox="[416,1178,345,1025]" captionTargetId="figure@5.[416,1179,345,1026]" captionTargetPageId="5" captionText="FIGURE 3. Distribution in Sri Lanka of Puntius kamalika (type locality indicated by a circle)." httpUri="https://zenodo.org/record/183061/files/figure.png" pageId="4" pageNumber="59">Fig. 3</figureCitation>
). It is locally common.
</paragraph>
<paragraph id="534B36AF2718C0050DA93FD4AE44FE8B" blockId="5.[151,1436,152,298]" pageId="5" pageNumber="60">
Etymology. The species-name is an eponym honouring
<taxonomicName id="94F44D2C2718C0050E363FD4A8ACFF13" box="[858,972,152,178]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="63" phylum="Chordata" rank="species" species="kamalika">Kamalika</taxonomicName>
Abeyaratne (
<date id="274A106F2718C00509003FD4AE6DFF13" box="[1132,1293,152,178]" pageId="5" pageNumber="60" value="1934-06-22">22 June 1934</date>
–
<date id="274A106F2718C00508713FD4AA63FF7B" pageId="5" pageNumber="60" value="2004-12-11">11 December 2004</date>
), formed as a noun in apposition.
<taxonomicName id="94F44D2C2718C0050FFB3F8CA804FF78" box="[663,868,192,217]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="60" phylum="Chordata" rank="species" species="kamalika">
<emphasis id="6180EABD2718C0050FFB3F8CA804FF78" box="[663,868,192,217]" italics="true" pageId="5" pageNumber="60">Puntius kamalika</emphasis>
</taxonomicName>
has previously been known in
<collectingCountry id="2BE3763F2718C00509BA3F8CAE2AFF7B" box="[1238,1354,192,218]" name="Sri Lanka" pageId="5" pageNumber="60">Sri Lanka</collectingCountry>
by the vernacular names Ipilli kadeya and Mada ipilla (
<bibRefCitation id="37654B5E2718C0050FB93FA4A8DBFEA3" author="Deraniyagala" box="[725,955,232,258]" pageId="5" pageNumber="64" refString="Deraniyagala, P. E. P. (1930) The Eventognathi of Ceylon. Spolia Zeylanica, 16, 1 - 41." type="journal article" year="1930">Deraniyagala 1930</bibRefCitation>
). We propose that in English it be called Kami’s barb, an allusion to the diminutive by which Dr Abeyaratne was known throughout her life.
</paragraph>
</subSubSection>
</treatment>
</document>
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