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<document id="F0342FF770DEEE69B0647DB459995784" ID-DOI="10.5281/zenodo.13244981" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244981" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970368039" checkinUser="felipe" docAuthor="Stuart Ko Sok Thy Neang, Bryan L." docDate="2006" docId="CB2487A5FFF969012CEAC789FF02F62F" docLanguage="en" docName="RafflesBZool.54.1.129-155.pdf" docOrigin="Raffles Bulletin of Zoology 54 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Limnonectes dabanus" docType="treatment" docVersion="2" lastPageNumber="138" masterDocId="371DFFDDFFF169082F30C000FB36F417" masterDocTitle="A Collection Of Amphibians And Reptiles From Hilly Eastern Cambodia" masterLastPageNumber="155" masterPageNumber="129" pageNumber="137" updateTime="1722975809971" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="35F7D90169A84545A4F4A6AD11CC2877" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FD656651B54AE7269B79872350AFC4FC">
<mods:title id="30579245E09B01554A512D2D7E47FCDC">A Collection Of Amphibians And Reptiles From Hilly Eastern Cambodia</mods:title>
</mods:titleInfo>
<mods:name id="581B2EB3D3F5999531C93A5121C6A054" type="personal">
<mods:role id="F30321453D1AAC27F135B0BD5851E726">
<mods:roleTerm id="F622D1BAB089931587AEA24D99224B28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D13CDB9C83AD597AE5ADC39C3C8EAAAE">Stuart Ko Sok Thy Neang, Bryan L.</mods:namePart>
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<mods:typeOfResource id="3B4AA0776B8B7ACFE8FA24BDA9AFF85D">text</mods:typeOfResource>
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<mods:title id="F1C2A788F54B246C6A177522296890EB">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="BADD3F5D2DE3CD7B27FA857CAFC31844">
<mods:date id="1EF3F4907BC909A37C9CBFAB8A326BD4">2006</mods:date>
<mods:detail id="310C612B1177BEB0A6006E6BBBE5C292" type="pubDate">
<mods:number id="E9BFD805FEDE79E77953F5C06CA71C1B">2006-02-28</mods:number>
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<mods:number id="60979E4AAA42AF31EF29686FC3A573C7">54</mods:number>
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<mods:number id="389A07D3D8DB5EF23555CE440FC26D89">1</mods:number>
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<mods:identifier id="357B13B6E87056AB74AD764929804E40" type="DOI">10.5281/zenodo.13244981</mods:identifier>
<mods:identifier id="08A511BD12A2F9D7C9A0E30FD29421F7" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="FDE76DAE3EBCAA7727C83026787C0C8A" type="Zenodo-Dep">13244981</mods:identifier>
</mods:mods>
<treatment id="CB2487A5FFF969012CEAC789FF02F62F" LSID="urn:lsid:plazi:treatment:CB2487A5FFF969012CEAC789FF02F62F" httpUri="http://treatment.plazi.org/id/CB2487A5FFF969012CEAC789FF02F62F" lastPageId="9" lastPageNumber="138" pageId="8" pageNumber="137">
<subSubSection id="0B976538FFF969002CEAC789FE1CF3B6" box="[986,1322,1929,1953]" pageId="8" pageNumber="137" type="nomenclature">
<paragraph id="433236B3FFF969002CEAC789FE1CF3B6" blockId="8.[986,1322,1929,1985]" box="[986,1322,1929,1953]" pageId="8" pageNumber="137">
<heading id="187A81DFFFF969002CEAC789FE1CF3B6" bold="true" box="[986,1322,1929,1953]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="137" reason="2">
<taxonomicName id="848D4D30FFF969002CEAC789FE1CF3B6" ID-CoL="72DL9" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[986,1322,1929,1953]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="137" phylum="Chordata" rank="species" species="dabanus">
<emphasis id="71F9EAA1FFF969002CEAC789FE1CF3B6" bold="true" box="[986,1322,1929,1953]" pageId="8" pageNumber="137">
<emphasis id="71F9EAA1FFF969002CEAC789FFFAF3B6" bold="true" box="[986,1228,1929,1953]" italics="true" pageId="8" pageNumber="137">Limnonectes dabanus</emphasis>
(Smith)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B976538FFF969002B6AC7A9FF9DF3D6" box="[1114,1195,1961,1985]" pageId="8" pageNumber="137" type="description">
<paragraph id="433236B3FFF969002B6AC7A9FF9DF3D6" blockId="8.[986,1322,1929,1985]" box="[1114,1195,1961,1985]" pageId="8" pageNumber="137">
(
<figureCitation id="DBB62A36FFF969002B52C7A9FF95F3D6" box="[1122,1187,1961,1985]" captionStart="Fig" captionStartId="9.[113,148,2064,2086]" captionTargetBox="[112,752,1606,2047]" captionTargetId="figure-408@9.[112,752,1606,2047]" captionTargetPageId="9" captionText="Fig. 7. Male Limnonectes dabanus from Phnom Nam Lyr Wildlife Sanctuary, Pichrada District, Mondolkiri Province, Cambodia." figureDoi="http://doi.org/10.5281/zenodo.13244992" httpUri="https://zenodo.org/record/13244992/files/figure.png" pageId="8" pageNumber="137">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0B976538FFF969012C73C7E8F9AEF5E2" lastPageId="9" lastPageNumber="138" pageId="8" pageNumber="137" type="materials_examined">
<paragraph id="433236B3FFF969002C73C7E8F88AFC20" blockId="8.[835,1474,2024,2103]" pageId="8" pageNumber="137">
<emphasis id="71F9EAA1FFF969002C73C7E8FF29F3E9" bold="true" box="[835,1055,2024,2046]" italics="true" pageId="8" pageNumber="137">Material examined. </emphasis>
<materialsCitation id="F3E53CEEFFF969002B18C7E9F88EFC20" collectingDate="2003-11-03" collectionCode="FMNH" collectorName="O'Rang &amp; O Ronas Stream" elevation="450" latitude="12.291862" longLatPrecision="1" longitude="107.05167" pageId="8" pageNumber="137" specimenCode="FMNH 262747" specimenCount="1">
<collectorName id="EE785365FFF969002B18C7E9FF4EF3E9" box="[1064,1144,2025,2046]" pageId="8" pageNumber="137">ORang</collectorName>
:
<specimenCode id="132B9EC8FFF969002BB5C7E9FE29F3E8" box="[1157,1311,2025,2047]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="137" type="Museum">FMNH 262747</specimenCode>
, hilly evergreen forest,
<collectorName id="EE785365FFF969002CB5C806FF15FC0C" box="[901,1059,2054,2075]" pageId="8" pageNumber="137">O Ronas Stream</collectorName>
,
<geoCoordinate id="26B95074FFF969002B1CC806FF99FC0C" box="[1068,1199,2054,2075]" degrees="12" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="137" precision="1" seconds="30.7" value="12.291862">12°17'30.7&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFF969002B84C806FE75FC0C" box="[1204,1347,2053,2075]" degrees="107" direction="east" minutes="03" orientation="longitude" pageId="8" pageNumber="137" precision="1" seconds="06.0" value="107.05167">107°03'06.0&quot;E</geoCoordinate>
,
<quantity id="84759B56FFF969002A7CC805FEBFFC0C" box="[1356,1417,2053,2075]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="137" unit="m" value="450.0">
<elevation id="C8A0D180FFF969002A7CC805FEBFFC0C" box="[1356,1417,2053,2075]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="137" unit="m" value="450.0">450 m</elevation>
</quantity>
elev.,
<date id="37331073FFF969002C73C822F88EFC20" box="[835,952,2082,2103]" pageId="8" pageNumber="137" value="2003-11-03">
<collectingDate id="2777E99BFFF969002C73C822F88EFC20" box="[835,952,2082,2103]" pageId="8" pageNumber="137" value="2003-11-03">3 Nov.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="433236B3FFF869012F41C0C0F98DF572" blockId="9.[113,752,192,358]" pageId="9" pageNumber="138">
<materialsCitation id="F3E53CEEFFF869012F41C0C0F9A9F518" collectingDate="2000-06-16" collectionCode="FMNH" collectorName="O Nam Lyr Stream" elevation="600" latitude="12.537778" location="Phnom Nam Lyr Mountain" longLatPrecision="21" longitude="107.53333" pageId="9" pageNumber="138" specimenCode="FMNH 261924-27" specimenCount="1">
Pichrada:
<specimenCode id="132B9EC8FFF869012FFFC0C1FAB4F4C1" box="[207,386,192,214]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">FMNH 261924-27</specimenCode>
,
<location id="46526068FFF869012EBAC0C0F9BCF4C1" LSID="urn:lsid:plazi:treatment:CB2487A5FFF969012CEAC789FF02F62F:46526068FFF869012EBAC0C0F9BCF4C1" box="[394,650,192,214]" latitude="12.537778" longLatPrecision="21" longitude="107.53333" name="Phnom Nam Lyr Mountain" pageId="9" pageNumber="138">Phnom Nam Lyr Mountain</location>
, evergreen mixed with deciduous and bamboo forest along
<collectorName id="EE785365FFF869012D05C0DDF9DAF4E5" box="[565,748,221,242]" pageId="9" pageNumber="138">O Nam Lyr Stream</collectorName>
, near
<geoCoordinate id="26B95074FFF869012F92C0FAFA22F507" box="[162,276,250,272]" degrees="12" direction="north" minutes="32" orientation="latitude" pageId="9" pageNumber="138" precision="15" seconds="16" value="12.537778">12°32'16&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFF869012E2BC0FAFAA1F518" box="[283,407,250,271]" degrees="107" direction="east" minutes="32" orientation="longitude" pageId="9" pageNumber="138" precision="15" seconds="00" value="107.53333">107°32'00&quot;E</geoCoordinate>
,
<quantity id="84759B56FFF869012E93C0FAFAD4F518" box="[419,482,250,272]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="138" unit="m" value="600.0">
<elevation id="C8A0D180FFF869012E93C0FAFAD4F518" box="[419,482,250,272]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="138" unit="m" value="600.0">600 m</elevation>
</quantity>
elev.,
<date id="37331073FFF869012D15C0FAF9A9F518" box="[549,671,250,272]" pageId="9" pageNumber="138" value="2000-06-16">
<collectingDate id="2777E99BFFF869012D15C0FAF9A9F518" box="[549,671,250,272]" pageId="9" pageNumber="138" value="2000-06-16">16 Jun.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="F3E53CEEFFF869012D9BC0FAF981F572" collectingDate="2000-06-17" collectingDateMax="2000-06-21" collectingDateMin="2000-06-17" collectionCode="FMNH" elevation="650" latitude="12.537778" location="Phnom Nam Lyr Mountain" longLatPrecision="21" longitude="107.53333" pageId="9" pageNumber="138" specimenCode="FMNH 261929-32, 261935, 261937, 261942, 261944-45" specimenCount="1">
<specimenCode id="132B9EC8FFF869012D9BC0FAFBEDF53B" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">FMNH 261929-32</specimenCode>
,
<specimenCode id="132B9EC8FFF869012FD4C116FA18F53B" box="[228,302,278,300]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">261935</specimenCode>
,
<specimenCode id="132B9EC8FFF869012E07C116FAB7F53B" box="[311,385,278,300]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">261937</specimenCode>
,
<specimenCode id="132B9EC8FFF869012EBAC116FAE2F53B" box="[394,468,278,300]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">261942</specimenCode>
,
<specimenCode id="132B9EC8FFF869012EEDC116F97EF53B" box="[477,584,278,300]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">261944-45</specimenCode>
,
<location id="46526068FFF869012D60C116FBE6F55E" LSID="urn:lsid:plazi:treatment:CB2487A5FFF969012CEAC789FF02F62F:46526068FFF869012D60C116FBE6F55E" latitude="12.537778" longLatPrecision="21" longitude="107.53333" name="Phnom Nam Lyr Mountain" pageId="9" pageNumber="138">Phnom Nam Lyr Mountain</location>
, evergreen mixed with deciduous and bamboo forest, near
<geoCoordinate id="26B95074FFF869012F41C150FBD4F571" box="[113,226,336,358]" degrees="12" direction="north" minutes="32" orientation="latitude" pageId="9" pageNumber="138" precision="15" seconds="16" value="12.537778">12°32'16&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFF869012FD9C150FA53F572" box="[233,357,336,357]" degrees="107" direction="east" minutes="32" orientation="longitude" pageId="9" pageNumber="138" precision="15" seconds="00" value="107.53333">107°32'00&quot;E</geoCoordinate>
,
<quantity id="84759B56FFF869012E41C150FAEDF571" box="[369,475,336,358]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="138" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<elevation id="C8A0D180FFF869012E41C150FAEDF571" box="[369,475,336,358]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="138" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
</quantity>
elev.,
<date id="37331073FFF869012D2DC150F981F572" box="[541,695,336,357]" pageId="9" pageNumber="138" value="2000-06-17" valueMax="2000-06-21" valueMin="2000-06-17">
<collectingDate id="2777E99BFFF869012D2DC150F981F572" box="[541,695,336,357]" pageId="9" pageNumber="138" value="2000-06-17" valueMax="2000-06-21" valueMin="2000-06-17">17-21 Jun.2000</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="433236B3FFF869012F41C18AF9AEF5E2" blockId="9.[113,752,394,502]" pageId="9" pageNumber="138">
<materialsCitation id="F3E53CEEFFF869012F41C18AF995F5AB" collectingDate="2003-12-01" collectionCode="FMNH" collectorName="Siem Pang" elevation="600" latitude="14.267694" longLatPrecision="1" longitude="106.62906" pageId="9" pageNumber="138" specimenCode="FMNH 262744-45" specimenCount="1">
<collectorName id="EE785365FFF869012F41C18AFBEEF588" box="[113,216,394,416]" pageId="9" pageNumber="138">Siem Pang</collectorName>
:
<specimenCode id="132B9EC8FFF869012FEFC18AFAB9F5B7" box="[223,399,394,416]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">FMNH 262744-45</specimenCode>
, bamboo mixed with evergreen forest, near
<geoCoordinate id="26B95074FFF869012FAEC1A7FA17F5AB" box="[158,289,422,444]" degrees="14" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="138" precision="1" seconds="03.7" value="14.267694">14°16'03.7&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFF869012E15C1A6FA85F5AB" box="[293,435,422,444]" degrees="106" direction="east" minutes="37" orientation="longitude" pageId="9" pageNumber="138" precision="1" seconds="44.6" value="106.62906">106°37'44.6&quot;E</geoCoordinate>
,
<quantity id="84759B56FFF869012E8BC1A6FAC0F5AB" box="[443,502,422,444]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="138" unit="m" value="600.0">
<elevation id="C8A0D180FFF869012E8BC1A6FAC0F5AB" box="[443,502,422,444]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="138" unit="m" value="600.0">600 m</elevation>
</quantity>
elev.,
<date id="37331073FFF869012D03C1A7F995F5AB" box="[563,675,423,444]" pageId="9" pageNumber="138" value="2003-12-01">
<collectingDate id="2777E99BFFF869012D03C1A7F995F5AB" box="[563,675,423,444]" pageId="9" pageNumber="138" value="2003-12-01">1 Dec.2003</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="F3E53CEEFFF869012D9BC1A7F9A2F5E2" collectingDate="2003-12-02" collectionCode="FMNH" collectorName="O Chay Stream" elevation="370" latitude="14.294084" longLatPrecision="1" longitude="106.61244" pageId="9" pageNumber="138" specimenCode="FMNH 262920" specimenCount="1">
<specimenCode id="132B9EC8FFF869012D9BC1A7FB8BF5CE" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="138" type="Museum">FMNH 262920</specimenCode>
, bamboo mixed with evergreen forest,
<collectorName id="EE785365FFF869012D7DC1C3F9DAF5CF" box="[589,748,451,473]" pageId="9" pageNumber="138">O Chay Stream</collectorName>
,
<geoCoordinate id="26B95074FFF869012F41C1E0FBC2F5E2" box="[113,244,480,501]" degrees="14" direction="north" minutes="17" orientation="latitude" pageId="9" pageNumber="138" precision="1" seconds="38.7" value="14.294084">14°17'38.7&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFF869012FCBC1E0FABCF5E1" box="[251,394,480,502]" degrees="106" direction="east" minutes="36" orientation="longitude" pageId="9" pageNumber="138" precision="1" seconds="44.8" value="106.61244">106°36'44.8&quot;E</geoCoordinate>
,
<quantity id="84759B56FFF869012EA5C1E0FAE5F5E1" box="[405,467,480,502]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="138" unit="m" value="370.0">
<elevation id="C8A0D180FFF869012EA5C1E0FAE5F5E1" box="[405,467,480,502]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="138" unit="m" value="370.0">370 m</elevation>
</quantity>
elev.,
<date id="37331073FFF869012D25C1E0F9A2F5E2" box="[533,660,480,501]" pageId="9" pageNumber="138" value="2003-12-02">
<collectingDate id="2777E99BFFF869012D25C1E0F9A2F5E2" box="[533,660,480,501]" pageId="9" pageNumber="138" value="2003-12-02">02 Dec.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0B976538FFF869012F41C21DFF02F62F" pageId="9" pageNumber="138" type="discussion">
<paragraph id="433236B3FFF869012F41C21DFADFF063" blockId="9.[113,752,541,1141]" pageId="9" pageNumber="138">
<emphasis id="71F9EAA1FFF869012F41C21DFBDBF623" bold="true" box="[113,237,541,565]" italics="true" pageId="9" pageNumber="138">Remarks. </emphasis>
<specimenCount id="558BFD3AFFF869012FC5C21DFAA9F622" box="[245,415,541,565]" count="17" pageId="9" pageNumber="138" type="adult">Seventeen adult</specimenCount>
males fully agree with
<bibRefCitation id="271C4B42FFF869012DADC21DFB8CF642" author="Smith, M" pageId="9" pageNumber="138" pagination="215 - 229" refId="ref27116" refString="Smith, M., 1922. The frogs allied to Rana doriae. Journal of the Natural History Society of Siam, 4 (4): 215 - 229." type="journal article" year="1922">Smiths (1922)</bibRefCitation>
brief original description of
<taxonomicName id="848D4D30FFF869012D3DC23EFBF7F662" authorityName="Smith" authorityYear="1922" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="subSpecies" species="macrognathus" subSpecies="dabana">
<emphasis id="71F9EAA1FFF869012D3DC23EFBF7F662" italics="true" pageId="9" pageNumber="138">Rana macrognathus dabana</emphasis>
</taxonomicName>
from the Langbian Plateau of southern
<collectingCountry id="3B9A7623FFF869012D41C25DF9FBF662" box="[625,717,605,629]" name="Vietnam" pageId="9" pageNumber="138">Vietnam</collectingCountry>
by having distinctly enlarged heads, encompassing nearly half the SVL; a swelling consisting of dense fibrous tissue on top of the head, commencing at the interorbital region and extending to or near to the level of the posterior border of the tympanum; the heels not or only slightly touching when the hindlimbs are folded at right angles to the body; and in measurements (Table 4). The specimens also fully agree with the
<typeStatus id="9C368811FFF869012FAFC35DFA35F762" box="[159,259,861,885]" pageId="9" pageNumber="138" type="holotype">holotype</typeStatus>
(MNHN 1948.0126) of
<taxonomicName id="848D4D30FFF869012D1CC35EFA24F782" authority="Bourret, 1941 b" authorityName="Bourret" authorityYear="1941" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="toumanoffi">
<emphasis id="71F9EAA1FFF869012D1CC35EF9D9F762" box="[556,751,861,885]" italics="true" pageId="9" pageNumber="138">Rana toumanoffi</emphasis>
<bibRefCitation id="271C4B42FFF869012F41C37DFA24F782" author="Bourret, R" box="[113,274,893,917]" pageId="9" pageNumber="138" pagination="1 - 29" refId="ref25096" refString="Bourret, R., 1941 b. Notes herpetologiques sur l'Indochine francaise. XXII. Reptiles et batraciens recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1941. Description d'une espece et d'une variete nouvelles. Direction de l'Instruction Publique: 1 - 29." type="book chapter" year="1941">Bourret, 1941b</bibRefCitation>
</taxonomicName>
, a single male from Mimot,
<collectingCountry id="3B9A7623FFF869012D06C37DF990F782" box="[566,678,893,917]" name="Cambodia" pageId="9" pageNumber="138">Cambodia</collectingCountry>
, which we have examined. There are no apparent differences in body measurements among Smiths
<typeStatus id="9C368811FFF869012EFFC3BDF904F7C2" box="[463,562,957,981]" pageId="9" pageNumber="138" type="syntype">syntypes</typeStatus>
, our Cambodian specimens, and Bourrets
<typeStatus id="9C368811FFF869012EBBC3DDFADEF7E2" box="[395,488,989,1013]" pageId="9" pageNumber="138" type="holotype">holotype</typeStatus>
of
<taxonomicName id="848D4D30FFF869012D3FC3DEF995F7E2" authorityName="Bourret" authorityYear="1941" box="[527,675,989,1013]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="toumanoffi">
<emphasis id="71F9EAA1FFF869012D3FC3DEF995F7E2" box="[527,675,989,1013]" italics="true" pageId="9" pageNumber="138">R. toumanoffi</emphasis>
</taxonomicName>
(Table 1), although the degree of enlargement of the head and head swelling varies considerably among individual males, even from the same locality. We treat
<taxonomicName id="848D4D30FFF869012EFAC43EF94BF042" authorityName="Bourret" authorityYear="1941" box="[458,637,1085,1109]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="toumanoffi">
<emphasis id="71F9EAA1FFF869012EFAC43EF94BF042" box="[458,637,1085,1109]" italics="true" pageId="9" pageNumber="138">Rana toumanoffi</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="848D4D30FFF869012FC9C45EFAD4F062" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[249,482,1117,1141]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="dabanus">
<emphasis id="71F9EAA1FFF869012FC9C45EFAD4F062" box="[249,482,1117,1141]" italics="true" pageId="9" pageNumber="138">Limnonectes dabanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="433236B3FFF869012F41C49DF940F203" blockId="9.[113,752,1181,1557]" pageId="9" pageNumber="138">
<bibRefCitation id="271C4B42FFF869012F41C49DFBC8F0A2" author="Smith, M" box="[113,254,1181,1205]" pageId="9" pageNumber="138" pagination="215 - 229" refId="ref27116" refString="Smith, M., 1922. The frogs allied to Rana doriae. Journal of the Natural History Society of Siam, 4 (4): 215 - 229." type="journal article" year="1922">Smith (1922)</bibRefCitation>
stated that juveniles and females of the species belonging to the
<taxonomicName id="848D4D30FFF869012E13C4BEFA91F0C2" authorityName="Boulenger" authorityYear="1887" box="[291,423,1213,1237]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="doriae">
<emphasis id="71F9EAA1FFF869012E13C4BEFA91F0C2" box="[291,423,1213,1237]" italics="true" pageId="9" pageNumber="138">Rana doriae</emphasis>
</taxonomicName>
group, in which he included
<taxonomicName id="848D4D30FFF869012DEAC4BEFBFDF0E2" baseAuthorityName="Smith" baseAuthorityYear="1922" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="dabanus">
<emphasis id="71F9EAA1FFF869012DEAC4BEFBFDF0E2" italics="true" pageId="9" pageNumber="138">L. dabanus</emphasis>
</taxonomicName>
, resemble each other so closely that “it is practically impossible to distinguish between them.” A large series (FMNH 261928, 261933-34, 261936, 261938-41, 261943, 262665, 262669, 262743, 262746, 262902-19, 262921-31, 262933-38, 262940-51, N = 60) of females, juveniles, and undeveloped males are tentatively assigned here to
<taxonomicName id="848D4D30FFF869012D4DC57EF9C6F182" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[637,752,1405,1429]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="dabanus">
<emphasis id="71F9EAA1FFF869012D4DC57EF9C6F182" box="[637,752,1405,1429]" italics="true" pageId="9" pageNumber="138">L. dabanus</emphasis>
</taxonomicName>
because they generally agree in skin texture and size (except head measurements) and were collected with the 17 developed males. Nine of the 77 (11.7%) specimens have a light vertebral stripe from the tip of the snout to near the vent.
</paragraph>
<paragraph id="433236B3FFF869012C1CC0C0FF47F5AF" blockId="9.[812,1452,192,440]" pageId="9" pageNumber="138">
Specimens were collected in and along small streams. This was the most frequently encountered, forest-dwelling frog species during our fieldwork. A developed male (FMNH 261937) was collected at night (2015 hrs.) in June sitting in a depression about twice its body size in a sandy vertical bank with exposed tree roots,
<quantity id="84759B56FFF869012B70C160FFB0F56F" box="[1088,1158,352,376]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="138" unit="cm" value="15.0">15 cm</quantity>
above a stream pool. The depression appeared to have been constructed by the frog, possibly for breeding activity.
</paragraph>
<paragraph id="433236B3FFF869012C1CC1E0FF02F62F" blockId="9.[812,1451,480,568]" pageId="9" pageNumber="138">
This is the first report of the species in
<collectingCountry id="3B9A7623FFF869012BC9C1E0FE5FF5EF" box="[1273,1385,480,504]" name="Cambodia" pageId="9" pageNumber="138">Cambodia</collectingCountry>
since
<bibRefCitation id="271C4B42FFF869012C1CC200F8CFF60F" author="Bourret, R" box="[812,1017,512,536]" pageId="9" pageNumber="138" pagination="1 - 29" refId="ref25096" refString="Bourret, R., 1941 b. Notes herpetologiques sur l'Indochine francaise. XXII. Reptiles et batraciens recus au Laboratoire des Sciences Naturelles de l'Universite au cours de l'annee 1941. Description d'une espece et d'une variete nouvelles. Direction de l'Instruction Publique: 1 - 29." type="book chapter" year="1941">Bourrets (1941b)</bibRefCitation>
description of its junior synonym
<taxonomicName id="848D4D30FFF869012AA3C201F896F62F" authorityName="Bourret" authorityYear="1941" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="toumanoffi">
<emphasis id="71F9EAA1FFF869012AA3C201F896F62F" italics="true" pageId="9" pageNumber="138">R. toumanoffi</emphasis>
</taxonomicName>
from Mimot.
</paragraph>
</subSubSection>
</treatment>
</document>