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<document id="5B69D3F012F1716B49ABB1FD83BE38DA" ID-DOI="10.5281/zenodo.184776" ID-GBIF-Dataset="45522745-642c-4528-a7da-0f641ff3164b" ID-ISSN="1175-5326" ID-Zenodo-Dep="184776" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460017201318" checkinUser="plazi" docAuthor="Mikhaljova, Elena V. &amp; Lim, Kil-Young" docDate="2008" docId="8C058934C42DFF96FF41FF44FEE3944A" docLanguage="en" docName="zt01925p061.pdf" docOrigin="Zootaxa 1925" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tokyosoma bellum Mikhaljova &amp; Lim, 2008, sp. n." docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="703CF14CC429FF93FFD6FFD3FFEB906D" masterDocTitle="The millipede family Diplomaragnidae Attems 1907 in the Korean Peninsula, with the descriptions of a new genus and new species (Diplopoda, Chordeumatida)" masterLastPageNumber="61" masterPageNumber="51" pageNumber="55" updateTime="1698233680532" updateUser="plazi">
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<mods:title id="CCED4EEEDFC105BADFEC04095078F401">The millipede family Diplomaragnidae Attems 1907 in the Korean Peninsula, with the descriptions of a new genus and new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:name id="85359B10382713AD39994181FB348197" type="personal">
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<mods:namePart id="1E804578E9C6CBE0B60D7A3E29370498">Mikhaljova, Elena V.</mods:namePart>
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<mods:namePart id="BE2AC2670C3BD5792A3BA0D28D38220D">Lim, Kil-Young</mods:namePart>
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<mods:typeOfResource id="E793B2A1D0DAED552F7CE1CD8556C89B">text</mods:typeOfResource>
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<mods:title id="2E95599E8282E95EBEB02FD093AE0F18">Zootaxa</mods:title>
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<mods:date id="F84A684571F6F4BFC8DA3FE0987F4BB1">2008</mods:date>
<mods:detail id="C2E52CD98E45587260A48BEF766FF6B1" type="volume">
<mods:number id="CB405E7E44F8A605AEB0ECE91B1E30B1">1925</mods:number>
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<mods:identifier id="667BA97DA34D74187345D43AB882BB82" type="DOI">10.5281/zenodo.184776</mods:identifier>
<mods:identifier id="5BDC63652678180057151FECAF8D3B4F" type="GBIF-Dataset">45522745-642c-4528-a7da-0f641ff3164b</mods:identifier>
<mods:identifier id="B78DE9C88F03FF12C524823404112830" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="639A600C142F3D53C170C3281ACB3558" type="Zenodo-Dep">184776</mods:identifier>
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<treatment id="8C058934C42DFF96FF41FF44FEE3944A" ID-DOI="http://doi.org/10.5281/zenodo.6229643" ID-GBIF-Taxon="119381638" ID-Zenodo-Dep="6229643" LSID="urn:lsid:plazi:treatment:8C058934C42DFF96FF41FF44FEE3944A" httpUri="http://treatment.plazi.org/id/8C058934C42DFF96FF41FF44FEE3944A" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
<subSubSection id="4CB66BA9C42DFF97FF41FF44FF1690B9" pageId="4" pageNumber="55" type="nomenclature">
<paragraph id="04133822C42DFF97FF41FF44FE2990DC" blockId="4.[151,450,151,212]" box="[151,450,151,177]" pageId="4" pageNumber="55">
<heading id="5F5B8F4EC42DFF97FF41FF44FE2990DC" bold="true" box="[151,450,151,177]" fontSize="11" level="1" pageId="4" pageNumber="55" reason="1">
<emphasis id="36D8E430C42DFF97FF41FF44FE2990DC" bold="true" box="[151,450,151,177]" pageId="4" pageNumber="55">
<taxonomicName id="C3AC43A1C42DFF97FF41FF44FE9290DC" box="[151,377,151,177]" class="Diplopoda" family="Diplomaragnidae" genus="Tokyosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="bellum" status="sp. nov.">
<emphasis id="36D8E430C42DFF97FF41FF44FE9290DC" bold="true" box="[151,377,151,177]" italics="true" pageId="4" pageNumber="55">Tokyosoma bellum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DEB594BC42DFF97FE56FF44FE2990DC" box="[384,450,151,177]" pageId="4" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="04133822C42DFF97FF41FF69FF1690B9" blockId="4.[151,450,151,212]" box="[151,253,186,212]" pageId="4" pageNumber="55">
<figureCitation id="9C9724A7C42DFF97FF41FF69FF1690B9" box="[151,253,186,212]" captionStart="FIGURES 7 9" captionStartId="4.[151,269,1509,1533]" captionTargetBox="[252,1304,622,1508]" captionTargetId="figure@4.[252,1335,601,1529]" captionTargetPageId="4" captionText="FIGURES 7 9. Tokyosoma bellum sp. n., male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm." httpUri="https://zenodo.org/record/184778/files/figure.png" pageId="4" pageNumber="55">Figs 79</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CB66BA9C42DFF97FF41FED9FDDC91A9" pageId="4" pageNumber="55" type="materials_examined">
<paragraph id="04133822C42DFF97FF41FED9FDDC91A9" blockId="4.[151,1437,266,572]" pageId="4" pageNumber="55">
<emphasis id="36D8E430C42DFF97FF41FED9FE93914E" box="[151,376,266,291]" italics="true" pageId="4" pageNumber="55">Material examined</emphasis>
:
<emphasis id="36D8E430C42DFF97FE5AFED9FE1C914E" box="[396,503,266,291]" italics="true" pageId="4" pageNumber="55">
<typeStatus id="DB178680C42DFF97FE5AFED9FE1C914E" box="[396,503,266,291]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="12AAF3ABC42DFF97FDDCFED9FDB69149" box="[522,605,266,292]" pageId="4" pageNumber="55" type="male">1 male</specimenCount>
(ChNU), Yangsan, Gyeongsangnamdo,
<collectingCountry id="7CBB78B2C42DFF97FB84FED9FB079149" box="[1106,1260,266,292]" name="South Korea" pageId="4" pageNumber="55">South Korea</collectingCountry>
,
<date id="70121EE2C42DFF97FB2DFED9FF3F9121" pageId="4" pageNumber="55" value="1990-12-28">28 December 1990</date>
, leg. K.Y. Lim;
<emphasis id="36D8E430C42DFF97FE74FEE1FDF29126" box="[418,537,306,331]" italics="true" pageId="4" pageNumber="55">
<typeStatus id="DB178680C42DFF97FE74FEE1FDF29126" box="[418,537,306,331]" pageId="4" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="12AAF3ABC42DFF97FDFBFEE1FD689121" box="[557,643,306,332]" pageId="4" pageNumber="55" type="male">1 male</specimenCount>
,
<specimenCount id="12AAF3ABC42DFF97FD42FEE1FCE29121" box="[660,777,306,332]" pageId="4" pageNumber="55" type="female">2 females</specimenCount>
(ChNU),
<specimenCount id="12AAF3ABC42DFF97FC5FFEE1FC0B9121" box="[905,992,306,332]" pageId="4" pageNumber="55" type="male">1 male</specimenCount>
,
<specimenCount id="12AAF3ABC42DFF97FC26FEE1FBB19121" box="[1008,1114,306,332]" pageId="4" pageNumber="55" type="female">1 female</specimenCount>
(
<collectionCode id="62BDA0E7C42DFF97FBB9FEE1FB599121" box="[1135,1202,306,332]" pageId="4" pageNumber="55">IBSS</collectionCode>
),
<specimenCount id="12AAF3ABC42DFF97FB1DFEE1FACA9121" box="[1227,1313,306,332]" pageId="4" pageNumber="55" type="male">1 male</specimenCount>
,
<specimenCount id="12AAF3ABC42DFF97FAE4FEE1FA779121" box="[1330,1436,306,332]" pageId="4" pageNumber="55" type="female">1 female</specimenCount>
(
<collectionCode id="62BDA0E7C42DFF97FF49FE89FEEA9119" box="[159,257,346,372]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="55">ZMUM</collectionCode>
), same locality as for
<typeStatus id="DB178680C42DFF97FDD1FE89FD9A9119" box="[519,625,346,372]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
,
<date id="70121EE2C42DFF97FDABFE89FC8A9119" box="[637,865,346,372]" pageId="4" pageNumber="55" value="1990-12-28">28 December 1990</date>
, leg. K.Y. Lim. One male
<typeStatus id="DB178680C42DFF97FB77FE89FAED9119" box="[1185,1286,346,372]" pageId="4" pageNumber="55" type="paratype">paratype</typeStatus>
is devoid of head. Long pointed outgrowth of the fused mesal sheath processes of posterior gonopod colpocoxites of one from dissected gonopods is broken.
</paragraph>
</subSubSection>
<subSubSection id="4CB66BA9C42DFF97FF10FE01FCDA9251" pageId="4" pageNumber="55" type="diagnosis">
<paragraph id="04133822C42DFF97FF10FE01FCDA9251" blockId="4.[151,1437,266,572]" pageId="4" pageNumber="55">
<emphasis id="36D8E430C42DFF97FF10FE01FED69186" box="[198,317,466,491]" italics="true" pageId="4" pageNumber="55">Diagnosis</emphasis>
: Differs from congeners mainly by shape (like cup with long pointed outgrowth) of mesal sheath structure of posterior gonopods, very low, waveshaped lateral sheath processes of posterior gonopod colpocoxites and scarcely subbasally fused colpocoxites.
</paragraph>
</subSubSection>
<caption id="50D368AAC42DFF97FF41FA36FECF962F" httpUri="https://zenodo.org/record/184778/files/figure.png" pageId="4" pageNumber="55" targetBox="[252,1304,622,1508]" targetPageId="4">
<paragraph id="04133822C42DFF97FF41FA36FECF962F" blockId="4.[151,1437,1509,1602]" pageId="4" pageNumber="55">
<emphasis id="36D8E430C42DFF97FF41FA36FEAE9590" bold="true" box="[151,325,1509,1533]" pageId="4" pageNumber="55">FIGURES 79.</emphasis>
<taxonomicName id="C3AC43A1C42DFF97FE98FA36FDF39591" box="[334,536,1509,1532]" class="Diplopoda" family="Diplomaragnidae" genus="Tokyosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="bellum" status="sp. nov.">
<emphasis id="36D8E430C42DFF97FE98FA36FDF39591" box="[334,536,1509,1532]" italics="true" pageId="4" pageNumber="55">Tokyosoma bellum</emphasis>
</taxonomicName>
<emphasis id="36D8E430C42DFF97FDF7FA36FDB49590" bold="true" box="[545,607,1509,1533]" pageId="4" pageNumber="55">
<taxonomicNameLabel id="2DEB594BC42DFF97FDF7FA36FDB49590" box="[545,607,1509,1533]" pageId="4" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm.
</paragraph>
</caption>
<subSubSection id="4CB66BA9C42DFF96FF10F9A0FEFF93BB" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55" type="description">
<paragraph id="04133822C42DFF97FF10F9A0FDBB96B0" blockId="4.[151,1437,1651,2037]" pageId="4" pageNumber="55">
<emphasis id="36D8E430C42DFF97FF10F9A0FEBA96E1" box="[198,337,1651,1676]" italics="true" pageId="4" pageNumber="55">Description</emphasis>
: Male. Length 7.07.5 mm, width about 0.7 mm. Coloration in alcohol pale with feebly marked beautiful design of light brown spots and strips on metazona and head. Legs with marbled brown distal part. Eyes black. Antennae brown.
</paragraph>
<paragraph id="04133822C42DFF97FF10F938FDAC97A0" blockId="4.[151,1437,1651,2037]" pageId="4" pageNumber="55">Body with 29 segments. Head covered with long and short setae, vertigial suture visible. Eye patches subtriangular, each composed of 1416 ocelli. Collum semicircular. Body width gradually increasing until somite 7, body parallelsided on somites 821(22), gradually tapering posteriorly. Beginning from somite 2(3) somites with normally developed paraterga which gradually grow less distinct toward hind part of body. Metazonital macrochaetae in transverse row on somites 2628, like elongate triangle on preceding somites. Macrochaetae long, pointed apically.</paragraph>
<paragraph id="04133822C42DFF96FF10F808FAEA919F" blockId="4.[151,1437,1651,2037]" lastBlockId="5.[151,1437,152,1063]" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
Legs long and slender. Each claw of legs 12 at base with two small additional claws dorsally and long setiform outgrowth ventrally. Leg pairs 37 not larger than other walking legs. Leg pairs 37 with small group of funnelshaped tarsal papillae apically near claw. Each claw of leg pairs 37 with long setiform outgrowth ventrally only i.e. without dorsal additional small claws. Leg pair 10 with several tarsal papillae subapically. Leg pair 11 with a few or without tarsal papillae. Midbody legs without tarsal papillae. Each claw of postgonopodal legs (including leg pairs 10 and 11) at base with long setiform outgrowth ventrally and two small additional claws dorsally; this additional claws gradually missing toward very hind legs. Each claw of very hind legs with long setiform outgrowth ventrally only. Legs 10 and 11 with coxal glands. Coxa 10 with ventral fovea of which caudomesal edge extended into thin process (
<figureCitation id="9C9724A7C42CFF96FCB0FE63FC4491A7" box="[870,943,432,458]" captionStart="FIGURES 7 9" captionStartId="4.[151,269,1509,1533]" captionTargetBox="[252,1304,622,1508]" captionTargetId="figure@4.[252,1335,601,1529]" captionTargetPageId="4" captionText="FIGURES 7 9. Tokyosoma bellum sp. n., male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm." httpUri="https://zenodo.org/record/184778/files/figure.png" pageId="5" pageNumber="56">Fig. 7</figureCitation>
). Coxa 11 without modifications (excluding opening of coxal gland ventrally). Trochanter 11 with small conical process directed caudad.
</paragraph>
<paragraph id="04133822C42CFF96FF10FDD3FDA093EF" blockId="5.[151,1437,152,1063]" pageId="5" pageNumber="56">
Anterior gonopod telopodite 1segmented, flagelliform, its distal part positioned inside elevated sheaths and somewhat protruding outside, apex curved, pointed, with tiny tooth subapically (
<figureCitation id="9C9724A7C42CFF96FB53FDFBFB25922F" box="[1157,1230,552,578]" captionStart="FIGURES 7 9" captionStartId="4.[151,269,1509,1533]" captionTargetBox="[252,1304,622,1508]" captionTargetId="figure@4.[252,1335,601,1529]" captionTargetPageId="4" captionText="FIGURES 7 9. Tokyosoma bellum sp. n., male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm." httpUri="https://zenodo.org/record/184778/files/figure.png" pageId="5" pageNumber="56">Fig. 8</figureCitation>
). Posterior gonopod colpocoxites scarcely fused subbasally. Each colpocoxite divided into broad median and narrow lateral branches; last one some flattened on sides. Mesal sheath processes of posterior gonopod colpocoxites fused medially into single cupshaped structure (ms) with long pointed outgrowth directed caudoventrally. Lateral sheath processes of colpocoxites (lp) very low, virtually undeveloped, waveshaped. Apex of colpocoxite curved posterad. Posterior gonopod angiocoxite with pearshaped globule in posterior view, depressed centrally in anterior view. Posterior angiocoxal process absent. Angiocoxite in anterior view with one process; anterior angiocoxal process long, its distal portion sheathed by colpocoxite on anterior side (
<figureCitation id="9C9724A7C42CFF96FB02FC93FAF79337" box="[1236,1308,832,858]" captionStart="FIGURES 7 9" captionStartId="4.[151,269,1509,1533]" captionTargetBox="[252,1304,622,1508]" captionTargetId="figure@4.[252,1335,601,1529]" captionTargetPageId="4" captionText="FIGURES 7 9. Tokyosoma bellum sp. n., male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm." httpUri="https://zenodo.org/record/184778/files/figure.png" pageId="5" pageNumber="56">Fig. 9</figureCitation>
). Posterior gonopod telopodite with short femur.
</paragraph>
<paragraph id="04133822C42CFF96FF10FC43FEFF93BB" blockId="5.[151,1437,152,1063]" pageId="5" pageNumber="56">Female. Length 88.5mm, width 0.80.9 mm. Body with 29 segments. Ocelli 1618. Vulvae not dissected.</paragraph>
</subSubSection>
<subSubSection id="4CB66BA9C42CFF96FF10FC36FEE3944A" pageId="5" pageNumber="56" type="etymology">
<paragraph id="04133822C42CFF96FF10FC36FEE3944A" blockId="5.[151,1437,152,1063]" pageId="5" pageNumber="56">
<emphasis id="36D8E430C42CFF96FF10FC36FEA89393" box="[198,323,997,1022]" italics="true" pageId="5" pageNumber="56">Etymology</emphasis>
: The specific epithet refers to the beautiful design of light brown spots and strips on metazona and head.
</paragraph>
</subSubSection>
</treatment>
</document>