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<document id="0AA3CC65E6854FB9C352D989C4043FC6" ID-DOI="10.3853/j.0067-1975.58.2006.1469" ID-ISSN="2201-4349" ID-Zenodo-Dep="4685692" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618342738327" checkinUser="carolina" docAuthor="Keable, Stephen J." docDate="2006" docId="8A0EDF188C076032FF3D8B2EFAC5F85D" docLanguage="en" docName="RecAustMus.58.2.133-244.pdf" docOrigin="Records of the Australian Museum 58 (2)" docSource="https://journals.australian.museum/keable-2006-rec-aust-mus-582-133244/" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Natatolana pilula" docType="treatment" docVersion="3" lastPageNumber="214" masterDocId="7637A7608C566063FFD48F2BFFB5FF90" masterDocTitle="Taxonomic Revision of Natatolana (Crustacea: Isopoda: Cirolanidae)" masterLastPageNumber="244" masterPageNumber="133" pageNumber="214" updateTime="1698905545581" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="7246CB64F0E936C195F39613A94AE5A2">Taxonomic Revision of Natatolana (Crustacea: Isopoda: Cirolanidae)</mods:title>
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<mods:namePart id="07A1FDDD1133BC52EFFAF756B11E596A">Keable, Stephen J.</mods:namePart>
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<mods:date id="FA42C312A3EF59A33A33A85C848FC666">2006</mods:date>
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<treatment id="8A0EDF188C076032FF3D8B2EFAC5F85D" LSID="urn:lsid:plazi:treatment:8A0EDF188C076032FF3D8B2EFAC5F85D" httpUri="http://treatment.plazi.org/id/8A0EDF188C076032FF3D8B2EFAC5F85D" lastPageNumber="214" pageId="81" pageNumber="214">
<subSubSection id="4ABD3D858C076032FF3D8B2EFDC8FB8D" box="[233,637,1029,1053]" pageId="81" pageNumber="214" type="nomenclature">
<paragraph id="02186E0E8C076032FF3D8B2EFDC8FB8D" blockId="81.[233,637,1029,1053]" box="[233,637,1029,1053]" pageId="81" pageNumber="214">
<heading id="5950D9628C076032FF3D8B2EFDC8FB8D" box="[233,637,1029,1053]" centered="true" fontSize="11" level="2" pageId="81" pageNumber="214" reason="2">
<taxonomicName id="C5A7158D8C076032FF3D8B2EFDC8FB8D" ID-CoL="45TD8" authority="(Barnard, 1955)" baseAuthorityName="Barnard" baseAuthorityYear="1955" box="[233,637,1029,1053]" class="Malacostraca" family="Cirolanidae" genus="Natatolana" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="214" phylum="Arthropoda" rank="species" species="pilula">
<emphasis id="30D3B21C8C076032FF3D8B2EFDC8FB8D" bold="true" box="[233,637,1029,1053]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FF3D8B2EFE00FB8D" bold="true" box="[233,437,1029,1053]" italics="true" pageId="81" pageNumber="214">Natatolana pilula</emphasis>
(
<bibRefCitation id="663613FF8C076032FE118B2EFDC0FB8D" author="Barnard, K" box="[453,629,1029,1053]" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard, 1955</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FEA48B1AFE40FBD8" box="[368,501,1073,1096]" pageId="81" pageNumber="214" type="description">
<paragraph id="02186E0E8C076032FEA48B1AFE40FBD8" blockId="81.[368,501,1073,1096]" box="[368,501,1073,1096]" pageId="81" pageNumber="214">
<figureCitation id="9A9C728B8C076032FEA48B1AFE40FBD8" box="[368,501,1073,1096]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="82.[186,221,1790,1811]" captionStartId-1="83.[446,481,1835,1856]" captionStartId-2="84.[491,526,1835,1856]" captionTargetBox-0="[189,1444,152,1773]" captionTargetBox-1="[129,1375,154,1762]" captionTargetBox-2="[210,1465,154,1805]" captionTargetId-0="figure-8@82.[188,1444,151,1811]" captionTargetId-1="figure-10@83.[121,1375,151,1762]" captionTargetId-2="figure-9@84.[207,1467,151,1809]" captionTargetPageId-0="82" captionTargetPageId-1="83" captionTargetPageId-2="84" captionText-0="Fig. 38. Natatolana pilula, male SAM A-14532. Scales for MD, MP = 0.5 mm, for MX1, MX2 = 0.2 mm, for A1, A2 = 0.5 mm." captionText-1="Fig. 39. Natatolana pilula, male SAM A-14532. Scales = 0.5 mm." captionText-2="Fig. 40. Natatolana pilula, male SAM A-14532. Scales = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4685822" figureDoi-1="http://doi.org/10.5281/zenodo.4685826" figureDoi-2="http://doi.org/10.5281/zenodo.4685828" httpUri-0="https://zenodo.org/record/4685822/files/figure.png" httpUri-1="https://zenodo.org/record/4685826/files/figure.png" httpUri-2="https://zenodo.org/record/4685828/files/figure.png" pageId="81" pageNumber="214">Figs. 3840</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FFAD8B75FE59FB53" pageId="81" pageNumber="214" type="reference_group">
<paragraph id="02186E0E8C076032FFAD8B75FECAFB1D" blockId="81.[121,749,1118,1219]" pageId="81" pageNumber="214">
<treatmentCitationGroup id="22B749208C076032FFAD8B75FECAFB1D" pageId="81" pageNumber="214">
<treatmentCitation id="8306481F8C076032FFAD8B75FE61FBE3" author="Barnard, K" box="[121,468,1118,1139]" page="54" pageId="81" pageNumber="214" year="1955">
<taxonomicName id="C5A7158D8C076032FFAD8B75FE61FBE3" ID-CoL="VGFC" authority="Barnard, 1955: 54" authorityName="Barnard" authorityPageNumber="54" authorityYear="1955" box="[121,468,1118,1139]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="214" phylum="Arthropoda" rank="species" species="pilula">
<emphasis id="30D3B21C8C076032FFAD8B75FEA7FBE3" box="[121,274,1118,1139]" italics="true" pageId="81" pageNumber="214">Cirolana pilula</emphasis>
<bibRefCitation id="663613FF8C076032FECF8B75FE61FBE3" author="Barnard, K" box="[283,468,1118,1139]" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard, 1955: 54</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 26ac.
<treatmentCitation id="8306481F8C076032FD828B75FF04FB1D" author="Kensley, B" page="39" pageId="81" pageNumber="214" year="1975">
<bibRefCitation id="663613FF8C076032FD828B75FF04FB1D" author="Kensley, B" pageId="81" pageNumber="214" pagination="35 - 89" refId="ref85881" refString="Kensley, B., 1975. Marine Isopoda from the continental shelf of South Africa. Annals of the South African Museum 67: 35 - 89." type="journal article" year="1975">Kensley, 1975: 39</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="8306481F8C076032FF168B53FEAEFB1D" author="Kensley, B" box="[194,283,1144,1165]" page="69" pageId="81" pageNumber="214" year="1978">
<bibRefCitation id="663613FF8C076032FF168B53FEAEFB1D" author="Kensley, B" box="[194,283,1144,1165]" pageId="81" pageNumber="214" refId="ref85910" refString="Kensley, B., 1978. Guide to the Marine Isopods of Southern Africa. Cape Town: Trustees of the South African Museum, pp. 173." type="book" year="1978">1978: 69</bibRefCitation>
</treatmentCitation>
, fig. 28H.
</treatmentCitationGroup>
</paragraph>
<paragraph id="02186E0E8C076032FFAD8BB8FE59FB53" blockId="81.[121,749,1118,1219]" pageId="81" pageNumber="214">
<treatmentCitationGroup id="22B749208C076032FFAD8BB8FE59FB53" pageId="81" pageNumber="214">
<taxonomicName id="C5A7158D8C076032FFAD8BB8FE9CFB38" ID-CoL="45TD8" baseAuthorityName="Barnard" baseAuthorityYear="1955" box="[121,297,1171,1192]" class="Malacostraca" family="Cirolanidae" genus="Natatolana" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="214" phylum="Arthropoda" rank="species" species="pilula">
<emphasis id="30D3B21C8C076032FFAD8BB8FE9CFB38" box="[121,297,1171,1192]" italics="true" pageId="81" pageNumber="214">Natatolana pilula</emphasis>
</taxonomicName>
.
<treatmentCitation id="8306481F8C076032FEEE8BB8FE5CFB38" author="Bruce, N" box="[314,489,1171,1192]" page="958" pageId="81" pageNumber="214" year="1981">
<bibRefCitation id="663613FF8C076032FEEE8BB8FE5CFB38" author="Bruce, N" box="[314,489,1171,1192]" pageId="81" pageNumber="214" pagination="945 - 966" refId="ref82900" refString="Bruce, N. L., 1981. Cirolanidae (Crustacea: Isopoda) of Australia: Diagnoses of Cirolana Leach, Metacirolana Nierstrasz, Neocirolana Hale, Anopsilana Paulian &amp; Debouteville, and Three New Genera - Natatolana, Politolana and Cartetolana. Australian Journal of Marine and Freshwater Research 32: 945 - 966." type="journal article" year="1981">Bruce 1981: 958</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="8306481F8C076032FE2E8BB8FDF2FB38" author="Bruce, N" box="[506,583,1171,1192]" page="8" pageId="81" pageNumber="214" year="1986">
<bibRefCitation id="663613FF8C076032FE2E8BB8FDF2FB38" author="Bruce, N" box="[506,583,1171,1192]" pageId="81" pageNumber="214" pagination="1 - 239" refId="ref82958" refString="Bruce, N. L., 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239." type="journal article" year="1986">1986: 8</bibRefCitation>
</treatmentCitation>
, 222.
<treatmentCitation id="8306481F8C076032FD588BB8FEA1FB53" author="Brusca, R" page="82" pageId="81" pageNumber="214" year="1995">
<bibRefCitation id="663613FF8C076032FD588BB8FEA1FB53" author="Brusca, R" pageId="81" pageNumber="214" pagination="1 - 96" refId="ref83454" refString="Brusca, R. C., R. Wetzer &amp; S. C. France, 1995. Cirolanidae (Crustacea: Isopoda: Flabellifera) of the Tropical Eastern Pacific. Proceedings of the San Diego Society of Natural History 30: 1 - 96." type="journal article" year="1995">
Brusca
<emphasis id="30D3B21C8C076032FD0F8BB8FF1FFB53" italics="true" pageId="81" pageNumber="214">et al</emphasis>
., 1995: 82
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="8306481F8C076032FEF08B85FE5DFB53" author="Kensley, B" box="[292,488,1198,1219]" page="230" pageId="81" pageNumber="214" year="2001">
<bibRefCitation id="663613FF8C076032FEF08B85FE5DFB53" author="Kensley, B" box="[292,488,1198,1219]" pageId="81" pageNumber="214" pagination="205 - 264" refId="ref86099" refString="Kensley, B., 2001. Biogeography of the marine Isopoda of the Indian Ocean, with a check-list of species and records. In Isopod Systematics and Evolution, Crustacean Issues, ed. B. Kensley &amp; R. C. Brusca, vol. 13, pp. 205 - 264. Rotterdam: A. A. Balkema." type="journal article" year="2001">Kensley, 2001: 230</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FFAD8BFCFD5BF9D1" pageId="81" pageNumber="214" type="materials_examined">
<paragraph id="02186E0E8C076032FFAD8BFCFEE0FA6A" blockId="81.[121,750,1239,1530]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FFAD8BFCFE9EFB7F" bold="true" box="[121,299,1239,1263]" pageId="81" pageNumber="214">Type material</emphasis>
.
<typeStatus id="DD1CD0AC8C076032FEE98BF2FE17FB7E" box="[317,418,1241,1262]" pageId="81" pageNumber="214" type="syntype">Syntypes</typeStatus>
:
<specimenCount id="14A1A5878C076032FE658BF2FD8CFB7E" box="[433,569,1241,1262]" pageId="81" pageNumber="214" type="generic">3 specimens</specimenCount>
,
<collectionCode id="64B6F6CB8C076032FD9C8BF2FDCBFB7E" box="[584,638,1241,1262]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="81" pageNumber="214">SAM</collectionCode>
A-43144. According to
<bibRefCitation id="663613FF8C076032FED68BDEFE2CFA9A" author="Barnard, K" box="[258,409,1269,1290]" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard (1955)</bibRefCitation>
“including male but no ovigerous female”. All examined. The material registered as the types has been largely dismembered. One specimen (including pereonites 27, pleon and pleotelson), however, has ooestegites and developing embryos. The remaining specimens cannot be sexed. The pleopod 2 with the appendix masculina figured by
<bibRefCitation id="663613FF8C076032FD4A8A51FF0FFA3A" author="Barnard, K" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard (1955)</bibRefCitation>
is not present. None of the type material is in good enough condition to use for a full redescription. Type locality: False Bay,
<collectingCountry id="7AB02E9E8C076032FF658AE1FE8EFA4F" box="[177,315,1482,1503]" name="South Africa" pageId="81" pageNumber="214">South Africa</collectingCountry>
,
<geoCoordinate id="679308C98C076032FE9D8AE1FE0BFA4F" box="[329,446,1482,1503]" degrees="34" direction="south" minutes="17" orientation="latitude" pageId="81" pageNumber="214" precision="15" seconds="06" value="-34.285">34°17'06&quot;S</geoCoordinate>
<geoCoordinate id="679308C98C076032FE1C8AE1FD8BFA4F" box="[456,574,1482,1503]" degrees="18" direction="east" minutes="39" orientation="longitude" pageId="81" pageNumber="214" precision="15" seconds="02" value="18.650555">18°39'02&quot;E</geoCoordinate>
and
<geoCoordinate id="679308C98C076032FDAC8AE1FD58FA4F" box="[632,749,1482,1503]" degrees="34" direction="south" minutes="12" orientation="latitude" pageId="81" pageNumber="214" precision="15" seconds="08" value="-34.202225">34°12'08&quot;S</geoCoordinate>
<geoCoordinate id="679308C98C076032FFAD8ACEFF5EFA6A" box="[121,235,1509,1530]" degrees="18" direction="east" minutes="36" orientation="longitude" pageId="81" pageNumber="214" precision="15" seconds="05" value="18.601389">18°36'05&quot;E</geoCoordinate>
,
<quantity id="C55FC3EB8C076032FF238ACEFEE4FA6A" box="[247,337,1509,1530]" metricMagnitude="1" metricUnit="m" metricValue="5.4" metricValueMax="6.2" metricValueMin="4.6" pageId="81" pageNumber="214" unit="m" value="54.0" valueMax="62.0" valueMin="46.0">
46
<specimenCount id="14A1A5878C076032FEC88ACEFEE4FA6A" box="[284,337,1509,1530]" pageId="81" pageNumber="214" type="generic">62 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="02186E0E8C076032FFAD8925FD5BF9D1" blockId="81.[121,750,1550,1601]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FFAD8925FEE7F9B6" bold="true" box="[121,338,1550,1574]" pageId="81" pageNumber="214">Material examined</emphasis>
.
<materialsCitation id="B2CF64538C076032FE89893BFD5CF9D1" collectingDate="1961-05-15" country="South Africa" latitude="-34.208332" location="False Bay" longLatPrecision="771" longitude="18.616667" pageId="81" pageNumber="214" specimenCount="1">
<emphasis id="30D3B21C8C076032FE89893BFE55F9B5" bold="true" box="[349,480,1552,1573]" pageId="81" pageNumber="214">
<collectingCountry id="7AB02E9E8C076032FE89893BFE55F9B5" box="[349,480,1552,1573]" name="South Africa" pageId="81" pageNumber="214">South Africa</collectingCountry>
</emphasis>
: 3, 10 mm, SAM-A14532,
<location id="077838D58C076032FFAD8907FF6FF9D1" LSID="urn:lsid:plazi:treatment:8A0EDF188C076032FF3D8B2EFAC5F85D:077838D58C076032FFAD8907FF6FF9D1" box="[121,218,1580,1601]" country="South Africa" latitude="-34.208332" longLatPrecision="771" longitude="18.616667" name="False Bay" pageId="81" pageNumber="214">False Bay</location>
,
<geoCoordinate id="679308C98C076032FF308907FEF6F9D1" box="[228,323,1580,1601]" degrees="34" direction="south" minutes="12.5" orientation="latitude" pageId="81" pageNumber="214" precision="92" value="-34.208332">34°12.5'S</geoCoordinate>
<geoCoordinate id="679308C98C076032FE9D8907FE2CF9D1" box="[329,409,1580,1601]" degrees="18" direction="east" minutes="37" orientation="longitude" pageId="81" pageNumber="214" precision="925" value="18.616667">18°37'E</geoCoordinate>
,
<date id="761948CE8C076032FE778907FD92F9D1" box="[419,551,1580,1601]" pageId="81" pageNumber="214" value="1961-05-15">
<collectingDate id="665DB1268C076032FE778907FD92F9D1" box="[419,551,1580,1601]" pageId="81" pageNumber="214" value="1961-05-15">15 May 1961</collectingDate>
</date>
, Sta No: FAL 414V
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FFAD897EFA3BF9FC" pageId="81" pageNumber="214" type="diagnosis">
<paragraph id="02186E0E8C076032FFAD897EFAD4FEF0" blockId="81.[121,749,1621,1997]" lastBlockId="81.[794,1422,153,352]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FFAD897EFF5DF9FD" bold="true" box="[121,232,1621,1645]" pageId="81" pageNumber="214">Diagnosis</emphasis>
.
<emphasis id="30D3B21C8C076032FF22897DFE75F9FD" box="[246,448,1622,1645]" italics="true" pageId="81" pageNumber="214">Interocular furrow</emphasis>
: well developed, extending across the cephalon; smoothly convex.
<emphasis id="30D3B21C8C076032FDE88958FD51F91A" box="[572,740,1651,1674]" italics="true" pageId="81" pageNumber="214">Frontal lamina</emphasis>
: lateral margins straight, parallel.
<emphasis id="30D3B21C8C076032FE3689BAFD89F938" box="[482,572,1681,1704]" italics="true" pageId="81" pageNumber="214">Antenna</emphasis>
: c. 0.2× as long as body, reaching to just posterior of pereonite 1.
<emphasis id="30D3B21C8C076032FD788985FF0FF972" italics="true" pageId="81" pageNumber="214">Coxal plates</emphasis>
: furrows variously developed, strong but incomplete on coxae 24, complete on coxae 57.
<emphasis id="30D3B21C8C076032FDE289C2FD1DF890" box="[566,680,1769,1792]" italics="true" pageId="81" pageNumber="214">Pleonite 4</emphasis>
: apex broadly rounded.
<emphasis id="30D3B21C8C076032FE97882DFE02F88D" box="[323,439,1798,1821]" italics="true" pageId="81" pageNumber="214">Pleotelson</emphasis>
: broad, length 0.67× basal width; anterodorsal depression absent; anterolateral margins convex; posterolateral margins broadly rounded; apex truncate; RS absent.
<emphasis id="30D3B21C8C076032FE8D8875FE4DF8E5" box="[345,504,1886,1909]" italics="true" pageId="81" pageNumber="214">Pereopods 23</emphasis>
: propodus without RS on palm.
<emphasis id="30D3B21C8C076032FF0F8850FEC2F802" box="[219,375,1915,1938]" italics="true" pageId="81" pageNumber="214">Pereopods 57</emphasis>
: propodus short and robust on each pereopod.
<emphasis id="30D3B21C8C076032FF3B88B2FED8F820" box="[239,365,1945,1968]" italics="true" pageId="81" pageNumber="214">Pereopod 7</emphasis>
: basis broad, width 0.58× length; distance between anterior margin and medial carina less than between posterior margin and medial carina; posterior margin with setae along entire length.
<emphasis id="30D3B21C8C076032FB3C8F9CFA9CFF5E" box="[1256,1321,183,206]" italics="true" pageId="81" pageNumber="214">Penes</emphasis>
: absent.
<emphasis id="30D3B21C8C076032FCCE8FFFFBDAFF7B" box="[794,1135,212,235]" italics="true" pageId="81" pageNumber="214">Pleopod 2 appendix masculina</emphasis>
: extending subequal with tip of endopod, 0.99× length of endopod; margins straight or sinuate; slender; apex bent laterally slightly, serrulate with a minute digitiform process.
<emphasis id="30D3B21C8C076032FBAB8E07FB69FED3" box="[1151,1244,300,323]" italics="true" pageId="81" pageNumber="214">Uropods</emphasis>
: exopod slightly shorter than endopod, 0.8× the length of the endopod.
</paragraph>
<paragraph id="02186E0E8C076032FCCE8E5EFC13F9AF" blockId="81.[794,1422,373,1599]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FCCE8E5EFBCFFE1D" bold="true" box="[794,1146,373,397]" pageId="81" pageNumber="214">Additional descriptive characters</emphasis>
.
<emphasis id="30D3B21C8C076032FB508E5DFB0DFE1D" box="[1156,1208,374,397]" italics="true" pageId="81" pageNumber="214">Body</emphasis>
: length c. 2.5× width. Colour cream to white in alcohol. Chromatophores absent.
<emphasis id="30D3B21C8C076032FCCE8E9BFCC8FE57" box="[794,893,432,455]" italics="true" pageId="81" pageNumber="214">Cephalon</emphasis>
: submarginal cephalic furrow moderately developed, does not run entire length of anterior margin.
<emphasis id="30D3B21C8C076032FAD38EE5FA8DFE75" box="[1287,1336,462,485]" italics="true" pageId="81" pageNumber="214">Eyes</emphasis>
: with 7 ommatidia in horizontal diameter; with 9 ommatidia in vertical diameter; ovate; colour deep red in alcohol.
<emphasis id="30D3B21C8C076032FB3E8D23FA33FD8F" box="[1258,1414,520,543]" italics="true" pageId="81" pageNumber="214">Frontal lamina</emphasis>
: length c. 3× basal width; apex expanded, anterior margin angled.
<emphasis id="30D3B21C8C076032FCBD8D68FC64FDCA" box="[873,977,579,602]" italics="true" pageId="81" pageNumber="214">Antennule</emphasis>
: short, just reaching pereonite 1. Peduncular article 1 longer than article 2; article 2 with 1 large pappose seta; article 3 very long, subequal to or longer than the combined lengths of articles 1 and 2. Flagellum 10-articulate.
<emphasis id="30D3B21C8C076032FCCE8D93FCC5FD5F" box="[794,880,696,719]" italics="true" pageId="81" pageNumber="214">Antenna</emphasis>
: peduncular article 4 with long SS projecting from posterolateral margin; article 5 with 2 conspicuous pappose setae at posterodistal angle. Flagellum 15-articulate.
<emphasis id="30D3B21C8C076032FAF08DD8FA33FC9A" box="[1316,1414,755,778]" italics="true" pageId="81" pageNumber="214">Mandible</emphasis>
: setal row with 13 RS.
<emphasis id="30D3B21C8C076032FC248C3BFBE4FCB7" box="[1008,1105,784,807]" italics="true" pageId="81" pageNumber="214">Maxillule</emphasis>
: medial lobe with 3 large robust pappose setae; lateral lobe with 13 RS on distal surface.
<emphasis id="30D3B21C8C076032FAEC8C05FA33FCD5" box="[1336,1414,814,837]" italics="true" pageId="81" pageNumber="214">Maxilla</emphasis>
: lateral lobe with 6 SS; medial lobe with 5 SS and 6 PS; middle lobe with 15 SS.
<emphasis id="30D3B21C8C076032FC078C43FBF6FCEF" box="[979,1091,872,895]" italics="true" pageId="81" pageNumber="214">Maxilliped</emphasis>
: endite with 2 coupling hooks, and 5 PS.
<emphasis id="30D3B21C8C076032FC478CADFC6BFC0D" box="[915,990,902,925]" italics="true" pageId="81" pageNumber="214">Pereon</emphasis>
: ornamentation consists of 1 strongly developed furrow on the lateral margins of pereonite 1 and 1 weakly developed furrow on lateral margins of pereonites 2 7; pereonite 1 longest, 25 subequal, 67 progressively shorter.
<emphasis id="30D3B21C8C076032FCCE8CD0FCE9FB82" box="[794,860,1019,1042]" italics="true" pageId="81" pageNumber="214">Coxae</emphasis>
: pereonite 1, coxae 24 with rounded posteroventral corners, coxa 5 with a short square corner, coxae 6 and 7 with broad, acute, posteroventral corners.
<emphasis id="30D3B21C8C076032FB748B1DFAB5FBDD" box="[1184,1280,1078,1101]" italics="true" pageId="81" pageNumber="214">Pleonites</emphasis>
: 1 concealed by pereonite 7, 25 visible.
<emphasis id="30D3B21C8C076032FBE68B78FB2FFBFA" box="[1074,1178,1107,1130]" italics="true" pageId="81" pageNumber="214">Pleonite 2</emphasis>
: not produced.
<emphasis id="30D3B21C8C076032FAE38B78FC92FB17" italics="true" pageId="81" pageNumber="214">Pleonite 3</emphasis>
: subacute.
<emphasis id="30D3B21C8C076032FC418B5BFC4AFB17" box="[917,1023,1136,1159]" italics="true" pageId="81" pageNumber="214">Pleotelson</emphasis>
: PS restricted to apex.
<emphasis id="30D3B21C8C076032FB0A8B5BFAE5FB17" box="[1246,1360,1136,1159]" italics="true" pageId="81" pageNumber="214">Pereopod 7</emphasis>
: basis anterior margin straight; medial carina with PS along entire length; posterior margin convex, posterior margin SS present. Ischium anterior margin with non-RS absent; posterior margin with 2 RS (submarginal), PS present. Merus posterior margin with 8 RS, SS present. Carpus posterior margin with 1 RS, SS absent. Propodus subequal to carpus; posterior margin with 3 RS, SS present.
<emphasis id="30D3B21C8C076032FC6C8A15FB58FAC5" box="[952,1261,1342,1365]" italics="true" pageId="81" pageNumber="214">Pleopod 2 appendix masculina</emphasis>
: arising basally.
<emphasis id="30D3B21C8C076032FCCE8A70FC16FAE2" box="[794,931,1371,1394]" italics="true" pageId="81" pageNumber="214">Pleopods 15</emphasis>
: exopod suture absent on pleopods 35; endopod PS on most of margins on pleopods 14, absent on pleopod 5.
<emphasis id="30D3B21C8C076032FCCE8ABDFCC6FA3D" box="[794,883,1430,1453]" italics="true" pageId="81" pageNumber="214">Uropods</emphasis>
: endopod subcircular; medial margin rounded, with 3 RS, PS along entire length; apex with 3 RS; lateral margin straight, with 1 RS, PS on distal three-quarters. Exopod medial margin convex, with 1 RS, PS present on distal half; apex rounded, with 3 RS; lateral margin convex, with 2 RS, PS on distal quarter.
</paragraph>
<paragraph id="02186E0E8C076032FCCE897FFA3BF9FC" blockId="81.[794,1422,1620,1644]" box="[794,1422,1620,1644]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FCCE897FFCFDF9FC" bold="true" box="[794,840,1620,1644]" pageId="81" pageNumber="214">Size</emphasis>
.
<quantity id="C55FC3EB8C076032FC87897EFC70F9FC" box="[851,965,1621,1644]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="81" pageNumber="214" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
(
<bibRefCitation id="663613FF8C076032FC05897EFBD9F9FC" author="Barnard, K" box="[977,1132,1621,1644]" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard, 1955</bibRefCitation>
;
<bibRefCitation id="663613FF8C076032FBA2897EFABAF9FC" author="Kensley, B" box="[1142,1295,1621,1644]" pageId="81" pageNumber="214" refId="ref85910" refString="Kensley, B., 1978. Guide to the Marine Isopods of Southern Africa. Cape Town: Trustees of the South African Museum, pp. 173." type="book" year="1978">Kensley, 1978</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FCCE89AAFA3BF813" pageId="81" pageNumber="214" type="discussion">
<paragraph id="02186E0E8C076032FCCE89AAFA3EF960" blockId="81.[794,1422,1665,1923]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FCCE89AAFC31F909" bold="true" box="[794,900,1665,1689]" pageId="81" pageNumber="214">Remarks</emphasis>
. The material examined matches the original description and illustrations of
<taxonomicName id="C5A7158D8C076032FB8F89B5FAA0F925" baseAuthorityName="Barnard" baseAuthorityYear="1955" box="[1115,1301,1694,1717]" class="Malacostraca" family="Cirolanidae" genus="Natatolana" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="214" phylum="Arthropoda" rank="species" species="pilula">
<emphasis id="30D3B21C8C076032FB8F89B5FAA0F925" box="[1115,1301,1694,1717]" italics="true" pageId="81" pageNumber="214">Natatolana pilula</emphasis>
</taxonomicName>
by
<bibRefCitation id="663613FF8C076032FAEC89B5FCD7F943" author="Barnard, K" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard (1955)</bibRefCitation>
except that chromatophores are absent, whereas the original description mentions pinkish or greyish speckles.
</paragraph>
<paragraph id="02186E0E8C076032FCEC89DDFA3BF813" blockId="81.[794,1422,1665,1923]" pageId="81" pageNumber="214">
The following characters are most useful in distinguishing
<taxonomicName id="C5A7158D8C076032FCCE883FFC61F8BB" baseAuthorityName="Barnard" baseAuthorityYear="1955" box="[794,980,1812,1835]" class="Malacostraca" family="Cirolanidae" genus="Natatolana" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="214" phylum="Arthropoda" rank="species" species="pilula">
<emphasis id="30D3B21C8C076032FCCE883FFC61F8BB" box="[794,980,1812,1835]" italics="true" pageId="81" pageNumber="214">Natatolana pilula</emphasis>
</taxonomicName>
from other species within the genus; the short antenna, the shape of the posterolateral margins of pleonites 3 and 4, the shape and setation of the pleotelson and uropods; and the morphology of the appendix masculina apex.
</paragraph>
</subSubSection>
<subSubSection id="4ABD3D858C076032FCCE88B3FAC5F85D" pageId="81" pageNumber="214" type="distribution">
<paragraph id="02186E0E8C076032FCCE88B3FAC5F85D" blockId="81.[794,1421,1944,1997]" pageId="81" pageNumber="214">
<emphasis id="30D3B21C8C076032FCCE88B3FC15F820" bold="true" box="[794,928,1944,1968]" pageId="81" pageNumber="214">Distribution</emphasis>
.
<collectingCountry id="7AB02E9E8C076032FC7E88B3FB85F83F" box="[938,1072,1944,1967]" name="South Africa" pageId="81" pageNumber="214">South Africa</collectingCountry>
: Lamberts Bay to Natal.At depths of
<quantity id="C55FC3EB8C076032FCE2889DFC27F85D" box="[822,914,1974,1997]" metricMagnitude="1" metricUnit="m" metricValue="4.2" metricValueMax="6.6" metricValueMin="1.8" pageId="81" pageNumber="214" unit="m" value="42.0" valueMax="66.0" valueMin="18.0">1866 m</quantity>
(
<bibRefCitation id="663613FF8C076032FC4B889DFB8DF85D" author="Barnard, K" box="[927,1080,1974,1997]" pageId="81" pageNumber="214" pagination="1 - 107" refId="ref82536" refString="Barnard, K. H., 1955. Additions to the Fauna-list of South African Crustacea and Pycnogonida. Annals of the South African Museum 43: 1 - 107." type="journal article" year="1955">Barnard, 1955</bibRefCitation>
;
<bibRefCitation id="663613FF8C076032FB96889DFB6DF85D" author="Kensley, B" box="[1090,1240,1974,1997]" pageId="81" pageNumber="214" refId="ref85910" refString="Kensley, B., 1978. Guide to the Marine Isopods of Southern Africa. Cape Town: Trustees of the South African Museum, pp. 173." type="book" year="1978">Kensley, 1978</bibRefCitation>
;
<bibRefCitation id="663613FF8C076032FB36889DFAD1F85D" author="Bruce, N" box="[1250,1380,1974,1997]" pageId="81" pageNumber="214" pagination="1 - 239" refId="ref82958" refString="Bruce, N. L., 1986. Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement 6: 1 - 239." type="journal article" year="1986">Bruce, 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>